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316 lines
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<document id="F3E267A5BE4466D8A6DCBDA424BF430E" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago & Vaz-De, Fernando Z." docDate="2021" docId="03850669FFB9FFE5FE2A274D39D4CFB0" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius (Luederwaldtinia) simplicicornis" docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="42" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="53167433F1D45693FCB85CC2F33DE7FC">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="241F766149717DF5965E8F10A65F37F9">Montoya-Molina, Santiago</mods:namePart>
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<mods:affiliation id="34254DD424F3C9B212C7ED8599BEDC75">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="C3CA750E7921892EF2B9B8AD9533849D" type="email">horomo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="176307B12E01092A3D13A22423B25EDD">Vaz-De, Fernando Z.</mods:namePart>
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<mods:title id="811A380342F5438467283DC3786C62BB">European Journal of Taxonomy</mods:title>
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<mods:part id="14D6A0E97C3F68B15B79E39053EB0787">
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<mods:date id="58028089EE448EF244AA7298C114F4CB">2021</mods:date>
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<mods:number id="D948CF5CC493212DE94F3B700CEEDC32">2021-02-11</mods:number>
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<mods:number id="8AE9301CE73CD96AD0C485B9CE71E3BB">734</mods:number>
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<treatment id="03850669FFB9FFE5FE2A274D39D4CFB0" ID-DOI="http://doi.org/10.5281/zenodo.4545859" ID-GBIF-Taxon="177420803" ID-Zenodo-Dep="4545859" LSID="urn:lsid:plazi:treatment:03850669FFB9FFE5FE2A274D39D4CFB0" httpUri="http://treatment.plazi.org/id/03850669FFB9FFE5FE2A274D39D4CFB0" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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<subSubSection id="C336E4F4FFB9FFE7FE2A274D382ECA1E" pageId="41" pageNumber="42" type="nomenclature">
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<paragraph id="8B93B77FFFB9FFE7FE2A274D382ECA1E" blockId="41.[391,1196,264,329]" pageId="41" pageNumber="42">
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<taxonomicName id="4C2CCCFCFFB9FFE7FE2A274D3FF5CA75" ID-CoL="35ND2" authority="(Luederwaldt, 1935)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1935" box="[391,1196,264,291]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="simplicicornis" subGenus="Luederwaldtinia">
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<emphasis id="B9586B6DFFB9FFE7FE2A274D397BCA75" bold="true" box="[391,546,264,291]" italics="true" pageId="41" pageNumber="42">Dichotomius</emphasis>
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(
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<emphasis id="B9586B6DFFB9FFE7FD9E274D39A3CA75" bold="true" box="[563,762,264,291]" italics="true" pageId="41" pageNumber="42">Luederwaldtinia</emphasis>
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)
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<emphasis id="B9586B6DFFB9FFE7FCA6274D38ECCA75" bold="true" box="[779,949,264,291]" italics="true" pageId="41" pageNumber="42">simplicicornis</emphasis>
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(
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<bibRefCitation id="EFBDCA8EFFB9FFE7FC6E274D3FFDCA75" author="Luederwaldt H." box="[963,1188,264,291]" pageId="41" pageNumber="42" pagination="334 - 342" refId="ref30199" refString="Luederwaldt H. 1935. Zur Kenntnis der Gattung Pinotus (Coleoptera: Scarabaeidae). Revista Entomologica 5: 334 - 342." type="journal article" year="1935">Luederwaldt, 1935</bibRefCitation>
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)
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</taxonomicName>
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<figureCitation id="1317ABFAFFB9FFE7FD11276B3848CA1E" box="[700,785,302,328]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="41" pageNumber="42">Fig. 14</figureCitation>
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, Map 12
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</paragraph>
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</subSubSection>
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<subSubSection id="C336E4F4FFB9FFE7FF1027363E75CAA3" pageId="41" pageNumber="42" type="reference_group">
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<paragraph id="8B93B77FFFB9FFE7FF10273638B5CADB" blockId="41.[189,1004,370,397]" box="[189,1004,370,397]" pageId="41" pageNumber="42">
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<treatmentCitationGroup id="AB3C9051FFB9FFE7FF10273638B5CADB" box="[189,1004,370,397]" pageId="41" pageNumber="42">
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<treatmentCitation id="0A8D916EFFB9FFE7FF10273639BCCADA" author="Luederwaldt H." box="[189,741,370,397]" page="340" pageId="41" pageNumber="42" year="1935">
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<taxonomicName id="4C2CCCFCFFB9FFE7FF10273639BCCADA" ID-CoL="4HZST" authority="Luederwaldt, 1935: 340" authorityName="Luederwaldt" authorityPageNumber="340" authorityYear="1935" box="[189,741,370,397]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="simplicicornis">
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<emphasis id="B9586B6DFFB9FFE7FF1027363A9DCADA" box="[189,452,370,396]" italics="true" pageId="41" pageNumber="42">Pinotus simplicicornis</emphasis>
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<bibRefCitation id="EFBDCA8EFFB9FFE7FE66273739BCCADA" author="Luederwaldt H." box="[459,741,370,397]" pageId="41" pageNumber="42" pagination="334 - 342" refId="ref30199" refString="Luederwaldt H. 1935. Zur Kenntnis der Gattung Pinotus (Coleoptera: Scarabaeidae). Revista Entomologica 5: 334 - 342." type="journal article" year="1935">Luederwaldt, 1935: 340</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B93B77FFFB9FFE7FF1027F339A2CA86" blockId="41.[189,1324,438,502]" box="[189,763,438,464]" pageId="41" pageNumber="42">
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<treatmentCitationGroup id="AB3C9051FFB9FFE7FF1027F339A2CA86" box="[189,763,438,464]" pageId="41" pageNumber="42">
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<taxonomicName id="4C2CCCFCFFB9FFE7FF1027F33A9DCA86" ID-CoL="4HZST" authorityName="Luederwaldt" authorityYear="1935" box="[189,452,438,464]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="simplicicornis">
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<emphasis id="B9586B6DFFB9FFE7FF1027F33A9DCA86" box="[189,452,438,464]" italics="true" pageId="41" pageNumber="42">Pinotus simplicicornis</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A8D916EFFB9FFE7FE4C27F339AECA86" author="Blackwelder R. E." box="[481,759,438,464]" page="208" pageId="41" pageNumber="42" year="1944">
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<bibRefCitation id="EFBDCA8EFFB9FFE7FE4C27F339AECA86" author="Blackwelder R. E." box="[481,759,438,464]" pageId="41" pageNumber="42" pagination="189 - 341" refId="ref29526" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 208</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B93B77FFFB9FFE7FF10279E3E75CAA3" blockId="41.[189,1324,438,502]" box="[189,1324,475,502]" pageId="41" pageNumber="42">
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<treatmentCitationGroup id="AB3C9051FFB9FFE7FF10279E3E75CAA3" box="[189,1324,475,502]" pageId="41" pageNumber="42">
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<taxonomicName id="4C2CCCFCFFB9FFE7FF10279E3959CAA3" ID-CoL="35ND2" baseAuthorityName="Luederwaldt" baseAuthorityYear="1935" box="[189,512,475,501]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="simplicicornis">
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<emphasis id="B9586B6DFFB9FFE7FF10279E3959CAA3" box="[189,512,475,501]" italics="true" pageId="41" pageNumber="42">Dichotomius simplicicornis</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A8D916EFFB9FFE7FDB3279E381FCAA3" author="Ratcliffe B. C. & Jameson M. L. & Figueroa L. & Cave R. V. & Paulsen M. L." box="[542,838,475,502]" page="197" pageId="41" pageNumber="42" year="2015">
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<bibRefCitation id="EFBDCA8EFFB9FFE7FDB3279E381FCAA3" author="Ratcliffe B. C. & Jameson M. L. & Figueroa L. & Cave R. V. & Paulsen M. L." box="[542,838,475,502]" pageId="41" pageNumber="42" pagination="186 - 207" refId="ref30717" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. V. & Paulsen M. L. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Scarabaeoidea. Journal of the Kansas Entomological Society 88 (2): 186 - 207. https: // doi. org / 10.2317 / kent- 88 - 02 - 186 - 207.1" type="journal article" year="2015">
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Ratcliffe
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<emphasis id="B9586B6DFFB9FFE7FD26279939E6CAA3" box="[651,703,475,501]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
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. 2015: 197
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</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0A8D916EFFB9FFE7FCDA279E3FFBCAA0" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[887,1186,475,502]" page="95" pageId="41" pageNumber="42" year="2018">
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<bibRefCitation id="EFBDCA8EFFB9FFE7FCDA279E3FFBCAA0" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[887,1186,475,502]" pageId="41" pageNumber="42" pagination="72 - 100" refId="ref29681" refString="Chamorro W., Marin-Armijos D., Granda V. & Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
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Chamorro
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<emphasis id="B9586B6DFFB9FFE7FC5827993F69CAA3" box="[1013,1072,475,501]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
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2018: 95
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A8D916EFFB9FFE7FB03279E3E7ECAA3" author="Chamorro W. & Marin-Armijos D. & Asenjo A. & Vaz-De-Mello F. Z." box="[1198,1319,475,501]" httpUri="http://treatment.plazi.org/id/ED33B49961AB966659162A5D2CF1F4BB" page="132" pageId="41" pageNumber="42" year="2019">
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<bibRefCitation id="EFBDCA8EFFB9FFE7FB03279E3E7ECAA3" author="Chamorro W. & Marin-Armijos D. & Asenjo A. & Vaz-De-Mello F. Z." box="[1198,1319,475,501]" pageId="41" pageNumber="42" pagination="1 - 343" refId="ref29753" refString="Chamorro W., Marin-Armijos D., Asenjo A. & Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 132</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C336E4F4FFB9FFE7FF10245A3FE8C856" pageId="41" pageNumber="42" type="diagnosis">
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<paragraph id="8B93B77FFFB9FFE7FF10245A3A6FC96F" blockId="41.[189,310,543,569]" box="[189,310,543,569]" pageId="41" pageNumber="42">
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<heading id="D0DB0013FFB9FFE7FF10245A3A6FC96F" bold="true" box="[189,310,543,569]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
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<emphasis id="B9586B6DFFB9FFE7FF10245A3A6FC96F" bold="true" box="[189,310,543,569]" pageId="41" pageNumber="42">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B93B77FFFB9FFE7FF1024153FE8C856" blockId="41.[188,1400,592,769]" pageId="41" pageNumber="42">
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<taxonomicName id="4C2CCCFCFFB9FFE7FF1024153958C93C" baseAuthorityName="Luederwaldt" baseAuthorityYear="1935" box="[189,513,592,618]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="simplicicornis">
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<emphasis id="B9586B6DFFB9FFE7FF1024153958C93C" box="[189,513,592,618]" italics="true" pageId="41" pageNumber="42">Dichotomius simplicicornis</emphasis>
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</taxonomicName>
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|
|
is separated from other species in the group by the following combination of characters: cephalic carina median tubercle higher than lateral ones, apically rounded and emarginate. Lateral tubercles short with acute on apex. Ventral clypeal process conical apically acute. Pronotum medial edge of the anterior disc declivity deeper than lateral sides, resembling two underdeveloped lobes. Elytral striae well marked, punctuation same width of striae; interstriae smooth.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C336E4F4FFB9FFE7FF10256F3F89CF7C" pageId="41" pageNumber="42" type="materials_examined">
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF10256F3AF3C812" blockId="41.[189,426,810,836]" box="[189,426,810,836]" pageId="41" pageNumber="42">
|
|||
|
|
<heading id="D0DB0013FFB9FFE7FF10256F3AF3C812" bold="true" box="[189,426,810,836]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FF10256F3AF3C812" bold="true" box="[189,426,810,836]" pageId="41" pageNumber="42">Material examined</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF47251E3873C820" blockId="41.[188,1399,859,961]" box="[234,810,859,886]" pageId="41" pageNumber="42">
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FF47251E3A3FC823" bold="true" box="[234,358,859,885]" pageId="41" pageNumber="42">
|
|||
|
|
<typeStatus id="549709DDFFB9FFE7FF47251E3A3FC823" box="[234,358,859,885]" pageId="41" pageNumber="42" type="lectotype">Lectotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
(head and pronotum, here designated)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF1025C43E74C897" blockId="41.[188,1399,859,961]" pageId="41" pageNumber="42">
|
|||
|
|
<materialsCitation id="3B44BD22FFB9FFE7FF1025C43E74C897" ID-GBIF-Occurrence="3037418328" collectionCode="BMNH" country="Peru" location="PERU" pageId="41" pageNumber="42" specimenCount="1" typeStatus="holotype">
|
|||
|
|
<collectingCountry id="F33BF7EFFFB9FFE7FF1025C43A51C8CD" box="[189,264,897,923]" name="Peru" pageId="41" pageNumber="42">PERU</collectingCountry>
|
|||
|
|
•
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FEB325C43A6DC8CD" bold="true" box="[286,308,897,923]" pageId="41" pageNumber="42">
|
|||
|
|
<specimenCount id="9D2A7CF6FFB9FFE7FEB325C43A6DC8CD" box="[286,308,897,923]" pageId="41" pageNumber="42" type="male">♂</specimenCount>
|
|||
|
|
</emphasis>
|
|||
|
|
; “Peru” [white, hw]/ “
|
|||
|
|
<taxonomicName id="4C2CCCFCFFB9FFE7FD9425C43866C8CC" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[569,831,896,922]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="simplicicornis" status="sp. nov.">
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FD9425C43866C8CC" box="[569,831,896,922]" italics="true" pageId="41" pageNumber="42">Pinotus simplicicornis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26BD616FFB9FFE7FCE825C438DAC8CD" box="[837,899,897,923]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
Lueder. Det. 1934” [white, hw]/ “Fry Coll. 1905-100” [white, p]/ “34600” [white, hw]/ “
|
|||
|
|
<typeStatus id="549709DDFFB9FFE7FD6025E33837C896" box="[717,878,934,960]" pageId="41" pageNumber="42" type="holotype">HOLOTYPE</typeStatus>
|
|||
|
|
” [red with black margins, p];
|
|||
|
|
<collectionCode id="ED3D2FBAFFB9FFE7FB6025E23E70C897" box="[1229,1321,935,961]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="41" pageNumber="42">BMNH</collectionCode>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF4725AF3AB8CF52" blockId="41.[189,1232,1002,1066]" box="[234,481,1002,1028]" pageId="41" pageNumber="42">
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FF4725AF3AB8CF52" bold="true" box="[234,481,1002,1028]" pageId="41" pageNumber="42">Additional material</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF1022553F89CF7C" blockId="41.[189,1232,1002,1066]" box="[189,1232,1037,1066]" pageId="41" pageNumber="42">
|
|||
|
|
<materialsCitation id="3B44BD22FFB9FFE7FF1022553F92CF7C" ID-GBIF-Occurrence="3037418503" box="[189,1227,1037,1066]" collectingDate="2006-04-02" collectionCode="CEMT" collectorName="A. Santibanez" country="Peru" location="Yaroni" municipality="Pichinaqui" pageId="41" pageNumber="42" specimenCount="12" specimenCount-female="4" specimenCount-male="8" stateProvince="Junin">
|
|||
|
|
<collectingCountry id="F33BF7EFFFB9FFE7FF1022553A51CF7C" box="[189,264,1040,1066]" name="Peru" pageId="41" pageNumber="42">PERU</collectingCountry>
|
|||
|
|
•
|
|||
|
|
<specimenCount id="9D2A7CF6FFB9FFE7FE8C22553A3DCF7F" box="[289,356,1037,1066]" pageId="41" pageNumber="42" type="male">8 ♂♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D2A7CF6FFB9FFE7FEDF22553AECCF7C" box="[370,437,1040,1066]" pageId="41" pageNumber="42" type="female">4 ♀♀</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectingRegion id="49E8799DFFB9FFE7FE6E2255395FCF7C" box="[451,518,1040,1066]" country="Peru" name="Junin" pageId="41" pageNumber="42">Junín</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="6BF72D05FFB9FFE7FDBC224A39CBCF7F" box="[529,658,1039,1065]" pageId="41" pageNumber="42">Pichinaqui</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<location id="8EF3E1A4FFB9FFE7FD31224A39B7CF7F" LSID="urn:lsid:plazi:treatment:03850669FFB9FFE5FE2A274D39D4CFB0:8EF3E1A4FFB9FFE7FD31224A39B7CF7F" box="[668,750,1039,1065]" country="Peru" municipality="Pichinaqui" name="Yaroni" pageId="41" pageNumber="42" stateProvince="Junin">Yaroni</location>
|
|||
|
|
;
|
|||
|
|
<date id="FF9291BFFFB9FFE7FD57225538DECF7F" box="[762,903,1039,1066]" pageId="41" pageNumber="42" value="2006-04-02">
|
|||
|
|
<collectingDate id="EFD66857FFB9FFE7FD57225538DECF7F" box="[762,903,1039,1066]" pageId="41" pageNumber="42" value="2006-04-02">2 Apr. 2006</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
;
|
|||
|
|
<collectorName id="26D9D2A9FFB9FFE7FC3F22553F6ECF7F" box="[914,1079,1039,1066]" pageId="41" pageNumber="42">A. Santibáñez</collectorName>
|
|||
|
|
leg.;
|
|||
|
|
<collectionCode id="ED3D2FBAFFB9FFE7FBD522553F92CF7C" box="[1144,1227,1040,1066]" collectionName="CEMT" pageId="41" pageNumber="42">CEMT</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C336E4F4FFB9FFE5FF10221638B8C8C4" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF1022163A32CF3B" blockId="41.[189,363,1107,1133]" box="[189,363,1107,1133]" pageId="41" pageNumber="42">
|
|||
|
|
<heading id="D0DB0013FFB9FFE7FF1022163A32CF3B" bold="true" box="[189,363,1107,1133]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FF1022163A32CF3B" bold="true" box="[189,363,1107,1133]" pageId="41" pageNumber="42">Redescription</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF4722C13A73CFC8" blockId="41.[189,604,1156,1220]" box="[234,298,1156,1182]" pageId="41" pageNumber="42">
|
|||
|
|
<emphasis id="B9586B6DFFB9FFE7FF4722C13A73CFC8" bold="true" box="[234,298,1156,1182]" pageId="41" pageNumber="42">Male</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF1022EF3905CF92" blockId="41.[189,604,1156,1220]" box="[189,604,1194,1220]" pageId="41" pageNumber="42">
|
|||
|
|
MEASUREMENTS. Length:
|
|||
|
|
<quantity id="4CD41A9AFFB9FFE7FE7322EF3901CF92" box="[478,600,1194,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.017" pageId="41" pageNumber="42" unit="mm" value="20.17">20.17 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF1022AB39E4CE92" blockId="41.[188,1399,1261,1476]" pageId="41" pageNumber="42">
|
|||
|
|
HEAD. Anterior third surface transversely rugose, extending posteriorly not reaching base of cephalic carina. Interocular space smooth, with scattered deep punctures (
|
|||
|
|
<figureCitation id="1317ABFAFFB9FFE7FC7E23563F73CE7B" box="[979,1066,1299,1325]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="41" pageNumber="42">Fig. 14</figureCitation>
|
|||
|
|
C–D). Clypeal margin with large medial emargination separating two rounded clypeal teeth feebly angled upwards. Cephalic carina, wider than high and conical in shape (
|
|||
|
|
<figureCitation id="1317ABFAFFB9FFE7FDDF231B3986CE2E" box="[626,735,1374,1400]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="41" pageNumber="42">Fig. 14D</figureCitation>
|
|||
|
|
). Median tubercle three times as high as lateral tubercles, apically rounded and emarginate. Lateral tubercles short and acute on apex (
|
|||
|
|
<figureCitation id="1317ABFAFFB9FFE7FBEE23C13FF6CEC8" box="[1091,1199,1412,1438]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="41" pageNumber="42">Fig. 14C</figureCitation>
|
|||
|
|
). Ventral clypeal process conical in shape and apically acute.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF1023AB3F7CCDC8" blockId="41.[186,1399,1517,1694]" pageId="41" pageNumber="42">
|
|||
|
|
PRONOTUM. Twice as wide as it is long, wider than elytra. Pronotal disc shiny and with well-impressed simple punctures, larger punctures on anterolateral edges and near posterior basal central margin.Anterior disc with sharp declivity, medial portion deeper than lateral sides, resembling two underdeveloped lobes (
|
|||
|
|
<figureCitation id="1317ABFAFFB9FFE7FF68201B3A0CCD2E" box="[197,341,1630,1656]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="41" pageNumber="42">Fig. 14A, D</figureCitation>
|
|||
|
|
). Hypomeron medial disc smooth and shiny, external margin bearing rows of disorganized setigerous punctures, setae all approximately of same length (see
|
|||
|
|
<figureCitation id="1317ABFAFFB9FFE7FC1320C63F40CDC8" box="[958,1049,1667,1694]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="41" pageNumber="42">Fig. 2E</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFB9FFE7FF10208D3A28CCBF" blockId="41.[186,1401,1735,2025]" pageId="41" pageNumber="42">
|
|||
|
|
ABDOMEN. Prosternum surface shagreened, with scattered ocellate setigerous punctures. Mesosternum and metepisternum surface shagreened, with large ocellate setigerous punctures, separated by less than twice diameter of a puncture. Metasternum anterior surface and margins of mesocoxa microsculpture shagreened, covered by dense long setae that surrounds the margins of mesocoxa, joining the anteromedial with lateral portions of metasternum (see
|
|||
|
|
<figureCitation id="1317ABFAFFB9FFE7FD02211B384BCC2E" box="[687,786,1886,1912]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="41" pageNumber="42">Fig. 2G</figureCitation>
|
|||
|
|
). Longitudinal sulcus distinctly sculpted, ending as shallow mid-fossa. Sternites with ocellate punctures along anterior margins. Pygidium delimited by incomplete margin, faded apically, large punctures restricted to basal margin, medial and apical surfaces finely punctate.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF53E7F7FFBAFFE4FF1021043A1FCCBE" ID-DOI="http://doi.org/10.5281/zenodo.4542135" ID-Zenodo-Dep="4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="42" pageNumber="43" startId="42.[189,232,1857,1883]" targetBox="[231,1353,264,1817]" targetPageId="42">
|
|||
|
|
<paragraph id="8B93B77FFFBAFFE4FF1021043A1FCCBE" blockId="42.[188,1399,1855,2024]" pageId="42" pageNumber="43">
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FF1021043A43CC0D" bold="true" box="[189,282,1857,1883]" pageId="42" pageNumber="43">Fig. 14.</emphasis>
|
|||
|
|
<taxonomicName id="4C2CCCFCFFBAFFE4FE8F2104383ACC0A" authority="(Luederwaldt, 1935)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1935" box="[290,867,1857,1884]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="simplicicornis">
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FE8F2104393CCC0D" box="[290,613,1857,1883]" italics="true" pageId="42" pageNumber="43">Dichotomius simplicicornis</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFBDCA8EFFBAFFE4FDD821043800CC0A" author="Luederwaldt H." box="[629,857,1857,1884]" pageId="42" pageNumber="43" pagination="334 - 342" refId="ref30199" refString="Luederwaldt H. 1935. Zur Kenntnis der Gattung Pinotus (Coleoptera: Scarabaeidae). Revista Entomologica 5: 334 - 342." type="journal article" year="1935">Luederwaldt, 1935</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FCC3210438FCCC0D" bold="true" box="[878,933,1857,1883]" pageId="42" pageNumber="43">A–B</emphasis>
|
|||
|
|
. Dorsal habitus.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FBC321043FDACC0D" bold="true" box="[1134,1155,1857,1883]" pageId="42" pageNumber="43">A</emphasis>
|
|||
|
|
. ♂.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FB1A21043F92CC0D" bold="true" box="[1207,1227,1857,1883]" pageId="42" pageNumber="43">B</emphasis>
|
|||
|
|
. ♀.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FB5221043E4DCC0D" bold="true" box="[1279,1300,1857,1883]" pageId="42" pageNumber="43">C</emphasis>
|
|||
|
|
. Lateral view of the aedeagus.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FE6A21213A82CC28" bold="true" box="[455,475,1892,1918]" pageId="42" pageNumber="43">D</emphasis>
|
|||
|
|
. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FD6721C23987CCF7" bold="true" box="[714,734,1927,1953]" pageId="42" pageNumber="43">E</emphasis>
|
|||
|
|
. Ventral view of the aedeagus.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FBE121C23FDACCF7" bold="true" box="[1100,1155,1927,1953]" pageId="42" pageNumber="43">F–G</emphasis>
|
|||
|
|
. Lamella copulatrix.
|
|||
|
|
<emphasis id="B9586B6DFFBAFFE4FF1021EE3B8DCC93" bold="true" box="[189,212,1963,1989]" pageId="42" pageNumber="43">H</emphasis>
|
|||
|
|
. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF10274C3A6ECA30" blockId="43.[188,1399,265,358]" pageId="43" pageNumber="44">ELYTRA. Shiny, wider than long. Striae distinctly punctate separated by two times diameter of a puncture on elytral disc; punctures same width of striae. Interstriae smooth with conspicuous randomly arranged punctures.</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF1027CE3FCECAB1" blockId="43.[187,1398,394,487]" pageId="43" pageNumber="44">LEGS. Metacoxa microsculpture shagreened, posterior edge covered by row of dense setigerous punctures separated by less than diameter of a puncture, setae dense. In ventral view, profemora surface with scattered well-imprinted punctation, setigerous punctation restricted to medial edge.</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF102448386BC970" blockId="43.[189,818,524,551]" box="[189,818,524,551]" pageId="43" pageNumber="44">
|
|||
|
|
PARAMERES AND LAMELLA COPULATRIX. See
|
|||
|
|
<figureCitation id="1317ABFAFFBBFFE5FD0F244939ADC970" box="[674,756,524,550]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="43" pageNumber="44">Fig. 14</figureCitation>
|
|||
|
|
E–G.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF10240E3AB7C933" blockId="43.[189,494,587,613]" box="[189,494,587,613]" pageId="43" pageNumber="44">
|
|||
|
|
<heading id="D0DB0013FFBBFFE5FF10240E3AB7C933" bold="true" box="[189,494,587,613]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
|
|||
|
|
<emphasis id="B9586B6DFFBBFFE5FF10240E3AB7C933" bold="true" box="[189,494,587,613]" pageId="43" pageNumber="44">Morphological variation</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF47243D3A8AC9C4" blockId="43.[189,722,632,691]" box="[234,467,632,658]" pageId="43" pageNumber="44">
|
|||
|
|
<emphasis id="B9586B6DFFBBFFE5FF47243D3A08C9C4" bold="true" box="[234,337,632,658]" pageId="43" pageNumber="44">Females</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="1317ABFAFFBBFFE5FECC243D3A93C9C4" box="[353,458,632,658]" captionStart="Fig" captionStartId="42.[189,232,1857,1883]" captionTargetBox="[231,1353,264,1817]" captionTargetId="figure-86@42.[228,1359,257,1825]" captionTargetPageId="42" captionText="Fig. 14. Dichotomius simplicicornis (Luederwaldt, 1935). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the male head, cephalic carina apically rounded, emarginate, lateral tubercles short and acute on the apex. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Pronotum arcuate viewed from above, anterior edge declivity resembling two under-developed lobes at the sides." figureDoi="http://doi.org/10.5281/zenodo.4542135" httpUri="https://zenodo.org/record/4542135/files/figure.png" pageId="43" pageNumber="44">Fig. 14B</figureCitation>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF1024DC398BC9E5" blockId="43.[189,722,632,691]" box="[189,722,665,691]" pageId="43" pageNumber="44">As males except for the following characters:</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF10249C38ACC845" blockId="43.[189,1399,728,787]" pageId="43" pageNumber="44">HEAD. Surface completely rugose. Space between clypeal teeth less separated than in males. Genae border protruding clypeal margin. Fronto-clypeal carina bituberculate.</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF10257C39F2C805" blockId="43.[189,683,825,851]" box="[189,683,825,851]" pageId="43" pageNumber="44">PRONOTUM. Narrower than elytra, gibbous.</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF10253D38B8C8C4" blockId="43.[189,993,888,914]" box="[189,993,888,914]" pageId="43" pageNumber="44">ELYTRA. Twice as long as pronotum with shagreened microsculpture.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C336E4F4FFBBFFE5FF1025F239D4CFB0" pageId="43" pageNumber="44" type="discussion">
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF1025F23A76C887" blockId="43.[189,303,951,977]" box="[189,303,951,977]" pageId="43" pageNumber="44">
|
|||
|
|
<heading id="D0DB0013FFBBFFE5FF1025F23A76C887" bold="true" box="[189,303,951,977]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
|
|||
|
|
<emphasis id="B9586B6DFFBBFFE5FF1025F23A76C887" bold="true" box="[189,303,951,977]" pageId="43" pageNumber="44">Remarks</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B93B77FFFBBFFE5FF1025A139D4CFB0" blockId="43.[188,1399,996,1254]" pageId="43" pageNumber="44">
|
|||
|
|
The
|
|||
|
|
<typeStatus id="549709DDFFBBFFE5FF5C25A13A01C8A8" box="[241,344,996,1022]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
|||
|
|
has the first half of the body (head and pronotum) of a male and the second half (abdomen) of a female. As the ICZN states in Article 73.1.5 “
|
|||
|
|
<emphasis id="B9586B6DFFBBFFE5FCB122403BA7CFF5" italics="true" pageId="43" pageNumber="44">
|
|||
|
|
If a subsequent author finds that a
|
|||
|
|
<typeStatus id="549709DDFFBBFFE5FB6F22403E7ECF49" box="[1218,1319,1029,1055]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
|||
|
|
which consists of a set of components (e.g. disarticulated body parts) is not derived from an individual animal, the extraneous components may, by appropriate citation, be excluded from the
|
|||
|
|
<typeStatus id="549709DDFFBBFFE5FB3A22023FA5CF37" box="[1175,1276,1095,1121]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
|||
|
|
(material may be excluded from a hapantotype if it is found to contain components representing more than one taxon
|
|||
|
|
</emphasis>
|
|||
|
|
”; therefore, we here designate the head and the pronotum as
|
|||
|
|
<typeStatus id="549709DDFFBBFFE5FC4322CC3F04CFF5" box="[1006,1117,1161,1187]" pageId="43" pageNumber="44" type="lectotype">lectotype</typeStatus>
|
|||
|
|
. Endemic species from
|
|||
|
|
<collectingCountry id="F33BF7EFFFBBFFE5FF1022EE3BAACF93" box="[189,243,1195,1221]" name="Peru" pageId="43" pageNumber="44">Peru</collectingCountry>
|
|||
|
|
and individuals cited by
|
|||
|
|
<bibRefCitation id="EFBDCA8EFFBBFFE5FDB522EF3810CF93" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[536,841,1194,1221]" pageId="43" pageNumber="44" pagination="72 - 100" refId="ref29681" refString="Chamorro W., Marin-Armijos D., Granda V. & Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
|
|||
|
|
Chamorro
|
|||
|
|
<emphasis id="B9586B6DFFBBFFE5FD3822EE3996CF92" box="[661,719,1194,1220]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
|
|||
|
|
(2018: 95
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFBDCA8EFFBBFFE5FCF822EF3892CF92" author="Chamorro W. & Marin-Armijos D. & Asenjo A. & Vaz-De-Mello F. Z." box="[853,971,1194,1220]" pageId="43" pageNumber="44" pagination="1 - 343" refId="ref29753" refString="Chamorro W., Marin-Armijos D., Asenjo A. & Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 132</bibRefCitation>
|
|||
|
|
) from
|
|||
|
|
<collectingCountry id="F33BF7EFFFBBFFE5FBB622EF3F24CF92" box="[1051,1149,1194,1220]" name="Ecuador" pageId="43" pageNumber="44">Ecuador</collectingCountry>
|
|||
|
|
were the rare species
|
|||
|
|
<taxonomicName id="4C2CCCFCFFBBFFE5FF10228E39D0CFB0" authority="(Kirsch, 1871)" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[189,649,1227,1254]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="foveicollis">
|
|||
|
|
<emphasis id="B9586B6DFFBBFFE5FF10228E3A8ACFB3" box="[189,467,1227,1253]" italics="true" pageId="43" pageNumber="44">Dichotomius foveicollis</emphasis>
|
|||
|
|
(Kirsch, 1871)
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|