treatments-xml/data/DA/84/2B/DA842B5F321A57828FF213CA540661B9.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.136.38558" ID-PMC="PMC6917624" ID-Pensoft-Pub="1314-2003-136-1" ID-Pensoft-UUID="B5F4606C22E55D648519EBC4D21D52AF" ID-PubMed="31866735" ModsDocID="1314-2003-136-1" checkinTime="1576162279418" checkinUser="pensoft" docAuthor="Wurdack, Kenneth J. &amp; Michelangeli, Fabian A." docDate="2019" docId="DA842B5F321A57828FF213CA540661B9" docLanguage="en" docName="PhytoKeys 136: 1-21" docOrigin="PhytoKeys 136" docSource="http://dx.doi.org/10.3897/phytokeys.136.38558" docTitle="Tryssophyton merumense Wurdack, Mem. New York Bot. Gard. 10: 155. 1964" docType="treatment" docVersion="5" id="B5F4606C22E55D648519EBC4D21D52AF" lastPageNumber="1" masterDocId="B5F4606C22E55D648519EBC4D21D52AF" masterDocTitle="Systematics and relationships of Tryssophyton (Melastomataceae), with a second species from the Pakaraima Mountains of Guyana" masterLastPageNumber="21" masterPageNumber="1" pageNumber="1" updateTime="1668139578533" updateUser="ExternalLinkService">
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<mods:title>Systematics and relationships of Tryssophyton (Melastomataceae), with a second species from the Pakaraima Mountains of Guyana</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wurdack, Kenneth J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Michelangeli, Fabian A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>136</mods:number>
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<mods:start>1</mods:start>
<mods:end>21</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.136.38558</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.136.38558</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-136-1</mods:identifier>
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<treatment ID-GBIF-Taxon="160702308" LSID="urn:lsid:plazi:treatment:DA842B5F321A57828FF213CA540661B9" httpUri="http://treatment.plazi.org/id/DA842B5F321A57828FF213CA540661B9" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="DA842B5F-321A-5782-8FF2-13CA540661B9" authority="Wurdack, Mem. New York Bot. Gard. 10: 155. 1964." class="Magnoliopsida" family="Melastomataceae" genus="Tryssophyton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tryssophyton merumense" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="merumense">Tryssophyton merumense Wurdack, Mem. New York Bot. Gard. 10: 155. 1964.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Type.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guyana. Cuyuni-Mazaruni Region, Partang River, Merume Mtns., Merume Mt.; common on mossy logs in forest, 1140 m elev., 4 Jul 1960 (fl., fr.), 
<emphasis italics="true" pageId="0" pageNumber="1">S.S. Tillett</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">C.L. Tillett</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">&amp; R. Boyan 43988</emphasis>
(holotype: US-2343844!; isotypes: K-000329332!, NY-00245868!).
</paragraph>
<paragraph pageId="0" pageNumber="1">
It should be noted that both the US and NY sheets are each marked as the holotype. However, the protologue clearly states that the US specimen is the holotype, even citing the sheet number and has that designation in J. 
<normalizedToken originalValue="Wurdack’s">Wurdack's</normalizedToken>
handwriting. The NY sheet has 
<normalizedToken originalValue="“holotype”">&quot;holotype&quot;</normalizedToken>
merely typed on the label. Thus, there is no need to lectotypify this name and the NY specimen should be considered as an isotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The genus is combined from 
<emphasis italics="true" pageId="0" pageNumber="1">tryssos</emphasis>
(Greek, dainty or delicate) and 
<emphasis italics="true" pageId="0" pageNumber="1">phyton</emphasis>
(Greek, plant) and refers to the plant habit. The specific epithet refers to Merume Mountain where the type was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="additional collections examined">
<paragraph pageId="0" pageNumber="1">Additional collections examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guyana. Cuyuni-Mazaruni Region: Pakaraima Mountains, upper Karowrieng River at Maipuri Falls; mixed bryophyte, pteridophyte, herb community; sandstone boulders, white sand, large cave behind falls, 
<geoCoordinate degrees="5" direction="north" minutes="41" orientation="latitude" precision="925" value="5.6833334">5°41'N</geoCoordinate>
, 
<geoCoordinate degrees="60" direction="west" minutes="13" orientation="longitude" precision="925" value="-60.216667">60°13'W</geoCoordinate>
, 575-600 m elev., 13 Oct 1992 (fl.), 
<emphasis italics="true" pageId="0" pageNumber="1">B. Hoffmann 2939</emphasis>
(NY!, US!). 2nd and 3rd escarpments (of four) of Kamakusa Mt., upper west-facing slopes below summit, rich forest with 
<taxonomicName authorityName="Aublet" authorityYear="1775" class="Monocotyledoneae" family="Arecaceae" genus="Licania" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Licania" order="Principes" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Licania</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Aublet" authorityYear="1775" class="Dicotyledoneae" family="Ebenaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Ebenales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Ebenaceae</taxonomicName>
, tree ferns, 
<taxonomicName authorityName="Aublet" authorityYear="1775" class="Monocotyledoneae" family="Arecaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Principes" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Arecaceae</taxonomicName>
, 
<geoCoordinate degrees="5" direction="north" minutes="52" orientation="latitude" precision="1" seconds="55.2" value="5.882">5°52'55.2&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="60" direction="west" minutes="6" orientation="longitude" precision="1" seconds="34.5" value="-60.10958">60°6'34.5&quot;W</geoCoordinate>
, 1330 m elev., 8 June 2012 (fl., fr.), 
<emphasis italics="true" pageId="0" pageNumber="1">K. Wurdack 5870</emphasis>
(US!). Potaro-Siparuni Region: Mt. Wokomung, easternmost pinnacle of massif, scrub forest on sandstone and peat, with 
<taxonomicName authorityName="Kunth" authorityYear="1822" class="Liliopsida" family="Poaceae" genus="Guadua" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guadua" order="Poales" pageId="0" pageNumber="1" phylum="Liliopsida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Guadua</emphasis>
</taxonomicName>
, 
<taxonomicName class="Liliopsida" family="Arecaceae" genus="Euterpe" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euterpe" order="Arecales" pageId="0" pageNumber="1" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Euterpe</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum</emphasis>
</taxonomicName>
, 
<geoCoordinate degrees="5" direction="north" minutes="5" orientation="latitude" precision="1" seconds="34.4" value="5.092889">5°5'34.4&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="59" direction="west" minutes="50" orientation="longitude" precision="1" seconds="13.3" value="-59.837025">59°50'13.3&quot;W</geoCoordinate>
, 1524 m elev., 13 Jul 2003 (fl.), 
<emphasis italics="true" pageId="0" pageNumber="1">H.D. Clarke 10822</emphasis>
(NY!, US!). Mt. Ayanganna, east slope, plateau above second escarpment, growing on mossy tree trunks and roots, 
<geoCoordinate degrees="5" direction="north" minutes="22" orientation="latitude" precision="15" seconds="28" value="5.3744445">5°22'28&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="59" direction="west" minutes="58" orientation="longitude" precision="15" seconds="06" value="-59.968334">59°58'06&quot;W</geoCoordinate>
, 1340 m elev., 16 Mar 2014 (fl.), 
<emphasis italics="true" pageId="0" pageNumber="1">A. Radosavljevic 165</emphasis>
(US!).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The five collections of 
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="T. merumense" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="merumense">
<emphasis italics="true" pageId="0" pageNumber="1">T. merumense</emphasis>
</taxonomicName>
span a 90 km section of the central Pakaraima Mountains, but further exploration is likely to expand its range into similar habitats. The species was only recently discovered (
<emphasis italics="true" pageId="0" pageNumber="1">Radosavljevic 165</emphasis>
) on the slopes of relatively well-explored Mount Ayanganna, the highest mountain (2041 m) wholly within Guyana. At mid-elevations on the western slopes of Kamakusa Mountain, it occurred (
<emphasis italics="true" pageId="0" pageNumber="1">Wurdack 5870</emphasis>
) as scattered, rarely-reproductive individuals on rotting logs and peaty-humus zones around the bases of trees.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The mountainous area north of the village of Imbaimadai and including Kamakusa Mountain has been variously mapped as the Merume Mountains. However, exactly what corresponds to the peak &quot;Merume Mountain&quot; within the region and indicated as the type locality of 
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="T. merumense" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="merumense">
<emphasis italics="true" pageId="0" pageNumber="1">T. merumense</emphasis>
</taxonomicName>
, is unclear. Field notes (Bassett Maguire Field Collections, vol. 19, Archives of The New York Botanical Garden) reveal that during 11 
<normalizedToken originalValue="Jun–">Jun-</normalizedToken>
16 Jul 1960, after leaving Imbaimadai, the collecting team, led by Stephen Tillett entered the Kamakusa Mountain area from along the Partang River. After reaching Partang Falls, they ascended into the uplands following existing trails, which were probably made by gold-miners or 
<normalizedToken originalValue="“pork-knockers.”">&quot;pork-knockers.&quot;</normalizedToken>
It is likely that &quot;Merume Mountain&quot; of Tillett et al. is equivalent to Kamakusa Mountain, but details referring to lower elevations, southeast ridge, southeast side and cliffs do not indicate they reached summit where 
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="T. quadrifolius" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="quadrifolius">
<emphasis italics="true" pageId="0" pageNumber="1">T. quadrifolius</emphasis>
</taxonomicName>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>