<documentid="CC79FD362C3BAD28588A51FFA1424574"ID-CLB-Dataset="40804"ID-DOI="10.5281/zenodo.188725"ID-GBIF-Dataset="97a0d3d1-80c3-450b-9404-255b5a20c7f5"ID-ISSN="1175-5326"ID-Zenodo-Dep="188725"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"checkinTime="1461096479258"checkinUser="plazi"docAuthor="Shear, William A., Taylor, Steven J., Wynne, Judson & Krejca, Jean K."docDate="2009"docId="03DD87D0AB1ECC24FF2AF9D4FEF3FC34"docLanguage="en"docName="zt02151p065.pdf"docOrigin="Zootaxa 2151"docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article"docStyleId="890A69B780ED73D6DB8551B71C8AC79E"docStyleName="Zootaxa.2009-2012.journal_article"docStyleVersion="4"docTitle="Pratherodesmus despaini Shear, new species"docType="treatment"docUuid="390D8938-B824-467D-88FC-6AB62969AB82"docUuidSource="ZooBank"docVersion="12"lastPageNumber="58"masterDocId="FFE4FFA8AB17CC2FFFBDFFF5FFAEFFA4"masterDocTitle="Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae)"masterLastPageNumber="65"masterPageNumber="47"pageNumber="56"updateTime="1732492908304"updateUser="ExternalLinkService">
<mods:titleid="4CCB8EA3A0D97B3E1027D65C588239F4">Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae)</mods:title>
<figureCitationid="134F2A43AB1ECC26FF2AF9B6FF49F9F9"box="[151,231,1603,1629]"captionStart="FIGURES 1, 2"captionStartId="4.[151,269,906,930]"captionTargetBox="[154,1436,197,890]"captionTargetId="figure@4.[154,1439,197,891]"captionTargetPageId="4"captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi."httpUri="https://zenodo.org/record/188726/files/figure.png"pageId="9"pageNumber="56">Figs. 1</figureCitation>
<figureCitationid="134F2A43AB1ECC26FC94F8DEFC2FF8E1"box="[809,897,1835,1861]"captionStart="FIGURE 39"captionStartId="17.[151,255,987,1011]"captionTargetBox="[151,1435,390,962]"captionTargetId="figure@17.[151,1436,390,963]"captionTargetPageId="17"captionText="FIGURE 39. Living example of Pratherodesmus despaini, photographed in Kaweah Cave (photo by J. Krejca)."httpUri="https://zenodo.org/record/188736/files/figure.png"pageId="9"pageNumber="56">Fig. 39</figureCitation>
). Head about 50% wider than collum (
<figureCitationid="134F2A43AB1ECC26FAF8F8DEFA24F8E1"box="[1349,1418,1835,1861]"captionStart="FIGURES 1, 2"captionStartId="4.[151,269,906,930]"captionTargetBox="[154,1436,197,890]"captionTargetId="figure@4.[154,1439,197,891]"captionTargetPageId="4"captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi."httpUri="https://zenodo.org/record/188726/files/figure.png"pageId="9"pageNumber="56">Fig. 1</figureCitation>
). Antennae long, extending back to posterior border of fifth segment. Collum with anterior margin arcuate, with 12 marginal setae, posterior margin straight to slightly sinuate, with eight setae, middle row with six setae; posteriolateral angles slightly produced (
<figureCitationid="134F2A43AB1ECC26FD3EF855FD64F81E"box="[643,714,1952,1978]"captionStart="FIGURES 1, 2"captionStartId="4.[151,269,906,930]"captionTargetBox="[154,1436,197,890]"captionTargetId="figure@4.[154,1439,197,891]"captionTargetPageId="4"captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi."httpUri="https://zenodo.org/record/188726/files/figure.png"pageId="9"pageNumber="56">Fig. 1</figureCitation>
, segment 10) with three lateral marginal teeth, each subtended by a seta; anterior row of six setae very strongly procurved, lateralmost seta of row widely separated, not appearing to be part of row; middle row of six setae slightly recurved; posterior marginal row with six setae. Posteriolateral metazonital corners strongly produced beyond ozopores, ozopores subtended by usual three setae. Pygidium as described for
<figureCitationid="134F2A43AB1DCC25FD09FEF9FD52FE82"box="[692,764,268,294]"captionStart="FIGURES 1, 2"captionStartId="4.[151,269,906,930]"captionTargetBox="[154,1436,197,890]"captionTargetId="figure@4.[154,1439,197,891]"captionTargetPageId="4"captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi."httpUri="https://zenodo.org/record/188726/files/figure.png"pageId="10"pageNumber="57">Fig. 1</figureCitation>
). Gonopods (Figs. 20–24, 32, 35) with gonocoxae and gonostome as described for
. Prefemora strongly transverse, articulating process narrow, pointed; prefemoral stem not marked. Exomere present, small, slightly curved, acute. Endomerite large, trullate, apically with two small lobes set with prominent, regular, small warts. Solenomere longer than in other species, with fewer cuticular scales; subtending process short, acute. tibiotarsus not flattened, as acute process similar in length to solenomere.
locality. Kaweah Cave is located close to the western boundary of Sequoia Kings Canyon National Park, and is unique among the caves in the park because it occurs at low elevation and has greater biodiversity than is found among the higher altitude caves (for locations of caves in the park, see the map in
<bibRefCitationid="EFE54B37AB1CCC24FD02FEF9FC63FE82"author="Shear"box="[703,973,268,294]"pageId="11"pageNumber="65"refString="Shear, W. A., & Shelley, R. M. (2008) Cave millipeds of the United States. VI. Sequoiadesmus krejcae, n. gen., n. sp., from Sequoia and Kings Canyon National Parks, California, USA (Diplopoda, Polyesmida, Trichopolydesmidea, Macrosternodesmidae). Zootaxa, 1693, 41 - 48."type="journal article"year="2008">Shear & Shelley 2008</bibRefCitation>
, and two other endemic troglophiles are to be found there. The other troglobites are an unidentified trichoniscid isopod, a cambalid millipede, and a pseudoscorpion, probably a species of
long, ending in a large room where the animals were collected. One male was collected from a root, and the other specimens were associated with unidentified guano (probably rodent or bat) on the floor. The temperature was not measured on the day of collections, but during an earlier visit to the same room of the cave the temperature was recorded at 11 degrees Celsius. The surrounding vegetation is California lower montane blue oak-foothill pine woodland and savanna (SWReGAP land cover
Named for Joel Despain, Cave Management Specialist at Sequoia and Kings Canyon National Parks, who organized and performed much of the sampling that resulted in the discovery of many new cave species.
