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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="186" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="100" approvalRequired_for_treatments="80" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FFFF737D1B9FFF7CFD8D978A" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Canis hallstromi Troughton 1957" docType="treatment" docVersion="3" lastPageNumber="349" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="347" updateTime="1674153641407" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Parnaby, Harry E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingleby, Sandy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Divljan, Anja</mods:namePart>
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<mods:date>2017</mods:date>
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<mods:number>2017-10-06</mods:number>
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<mods:number>69</mods:number>
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<mods:number>5</mods:number>
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<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DD87C8FFFF737D1B9FFF7CFD8D978A" httpUri="http://treatment.plazi.org/id/03DD87C8FFFF737D1B9FFF7CFD8D978A" lastPageId="72" lastPageNumber="349" pageId="70" pageNumber="347">
<subSubSection pageId="70" pageNumber="347" type="nomenclature">
<paragraph blockId="70.[941,1365,151,178]" box="[941,1365,151,178]" pageId="70" pageNumber="347">
<heading box="[941,1365,151,178]" centered="true" fontSize="11" level="2" pageId="70" pageNumber="347" reason="2">
<taxonomicName authority="Troughton, 1957" authorityName="Troughton" authorityYear="1957" box="[941,1365,151,178]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="hallstromi">
<emphasis bold="true" box="[941,1365,151,178]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[941,1147,151,178]" italics="true" pageId="70" pageNumber="347">Canis hallstromi</emphasis>
Troughton, 1957
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="70.[869,1422,197,219]" box="[869,1422,197,219]" pageId="70" pageNumber="347">
<emphasis box="[869,1121,197,219]" italics="true" pageId="70" pageNumber="347">Proc. R. Zool. Soc. N.S.W.</emphasis>
19551956: 93. (
<date box="[1291,1411,197,219]" pageId="70" pageNumber="347" value="1957-05-08">8 May 1957</date>
).
</paragraph>
</subSubSection>
<subSubSection pageId="70" pageNumber="347" type="description">
<paragraph blockId="70.[839,1310,241,265]" box="[839,1310,241,265]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,1013,241,265]" pageId="70" pageNumber="347">Common name</emphasis>
. New
<collectingCountry box="[1083,1160,241,265]" name="Guinea-Bissau" pageId="70" pageNumber="347">Guinea</collectingCountry>
Singing Dog.
</paragraph>
<paragraph blockId="70.[837,1467,286,487]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,999,286,310]" pageId="70" pageNumber="347">Current name</emphasis>
.
<taxonomicName authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[1011,1353,286,310]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="familiaris">
<emphasis box="[1011,1182,286,310]" italics="true" pageId="70" pageNumber="347">Canis familiaris</emphasis>
<bibRefCitation author="Linnaeus, C" box="[1187,1353,286,310]" pageId="70" pageNumber="347" refId="ref137371" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tome 1. Holmiae: Laurentii Salvii. 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
; following
<bibRefCitation author="Jackson, S &amp; Groves, P &amp; Fleming, K &amp; Aplin, M &amp; Eldridge, A" box="[838,1061,316,340]" pageId="70" pageNumber="347" pagination="201 - 224" refId="ref133800" refString="Jackson, S. M., C. P. Groves, P. J. S. Fleming, K. P. Aplin, M. D. B. Eldridge, A. Gonzalez, and K. M. Helgen. 2017. The Wayward Dog: is the Australian native dog or Dingo a distinct species? Zootaxa: 4317 (2): 201 - 224. https: // doi. org / 10.11646 / zootaxa. 4317.2.1" type="journal article" year="2017">
Jackson
<emphasis box="[929,982,316,340]" italics="true" pageId="70" pageNumber="347">et al.</emphasis>
(2017)
</bibRefCitation>
. Taxonomic status has been contested; variously considered to be a feral lineage of the domesticated dog (e.g.,
<bibRefCitation author="Helgen, K" box="[946,1094,374,398]" pageId="70" pageNumber="347" pagination="689 - 749" refId="ref132662" refString="Helgen, K. M. 2007. A taxonomic and geographic overview of the mammals of Papua. In The Ecology of Papua, ed. A. J. Marshall and B. M. Beehler, Ecology of Indonesia Series, Vol. VI, Part 1 pp. 689 - 749. Singapore: Periplus Editions." type="book chapter" year="2007">Helgen, 2007</bibRefCitation>
;
<bibRefCitation author="Oskarsson, M &amp; Klutsch, U &amp; Boonyaprakob, A &amp; Wilton, Y" box="[1105,1343,374,398]" pageId="70" pageNumber="347" pagination="967 - 974" refId="ref140110" refString="Oskarsson, M. C. R., C. F. C. Klutsch, U. Boonyaprakob, A. Wilton, Y. Tanabe, and P. Savolainen. 2012. Mitochondrial DNA data indicate an introduction through Mainland Southeast Asia for Australian dingoes and Polynesian domestic dogs. Proceedings of the Royal Society of London B: Biological Sciences 279: 967 - 974. https: // doi. org / 10.1098 / rspb. 2011.1395" type="journal article" year="2012">
Oskarsson
<emphasis box="[1225,1280,374,398]" italics="true" pageId="70" pageNumber="347">et al.</emphasis>
2012
</bibRefCitation>
; Jackson &amp; Groves, 2015); a valid species (e.g.,
<bibRefCitation author="Crowther, M &amp; Fillios, N" box="[1231,1463,404,428]" pageId="70" pageNumber="347" pagination="192 - 203" refId="ref126659" refString="Crowther, M. S., M. Fillios, N. Colman, and M. Letnic. 2014. An updated description of the Australian dingo (Canis dingo Meyer, 1793). Journal of Zoology (London) 292 (3): 192 - 203. https: // doi. org / 10.1111 / jzo. 12134" type="journal article" year="2014">
Crowther
<emphasis box="[1338,1399,404,428]" italics="true" pageId="70" pageNumber="347">et al.,</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation author="Koler-Matznick, J &amp; Yates, S" box="[839,1130,433,457]" pageId="70" pageNumber="347" pagination="47 - 56" refId="ref134825" refString="Koler-Matznick, J., B. C. Yates, S. Bulmer, and I. L. Brisbin, Jr. 2007. The New Guinea singing dog: its status and scientific importance. Australian Mammalogy 29 (1): 47 - 56. https: // doi. org / 10.1071 / AM 07005" type="journal article" year="2007">
Koler-Matznick
<emphasis box="[1012,1055,433,457]" italics="true" pageId="70" pageNumber="347">et al</emphasis>
., 2007
</bibRefCitation>
;
<bibRefCitation author="Troughton, E" box="[1138,1314,433,457]" pageId="70" pageNumber="347" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton, 1971</bibRefCitation>
) or a synonym of
<taxonomicName box="[866,997,462,486]" class="Mammalia" family="Canidae" genus="Canus" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="lupus">
<emphasis box="[866,997,462,486]" italics="true" pageId="70" pageNumber="347">Canus lupus</emphasis>
</taxonomicName>
<taxonomicName authority="Meyer, 1793" authorityName="Meyer" authorityYear="1793" box="[1002,1199,462,487]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="dingo">
<emphasis box="[1002,1063,462,486]" italics="true" pageId="70" pageNumber="347">dingo</emphasis>
<bibRefCitation author="Meyer, F" box="[1066,1199,462,487]" pageId="70" pageNumber="347" refId="ref138969" refString="Meyer, F. A. A. 1793. Systematisch-summarische Uebersicht der neuesten zoologischen Entdeckungen in Neuholland und Afrika. Nebst zwey andern zoologischen Abhandlungen. Leipzig: Dykischen Buchhandlung. 178 pp. https: // doi. org / 10.5962 / bhl. title. 39685" type="book" year="1793">Meyer, 1793</bibRefCitation>
</taxonomicName>
(e.g.,
<bibRefCitation author="Wozencraft, W" box="[1264,1456,462,486]" pageId="70" pageNumber="347" pagination="532 - 628" refId="ref150291" refString="Wozencraft, W. C. 2005. Order Carnivora. In Mammal Species of the World: A Taxonomic and Geographic Reference, ed. D. E. Wilson and D. M. Reeder, pp. 532 - 628. Baltimore, USA: The Johns Hopkins University Press." type="book chapter" year="2005">Wozencraft, 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="70" pageNumber="347" type="materials_examined">
<paragraph blockId="70.[839,1467,507,649]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,943,508,532]" pageId="70" pageNumber="347">
<typeStatus box="[839,943,508,532]" pageId="70" pageNumber="347">Holotype</typeStatus>
</emphasis>
. M.8502, by subsequent designation (
<bibRefCitation author="Troughton, E" pageId="70" pageNumber="347" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton, 1971</bibRefCitation>
). Male, skull (
<figureCitation box="[1043,1121,537,561]" captionStart="Figure 29" captionStartId="71.[222,285,1888,1910]" captionTargetBox="[256,1285,154,1853]" captionTargetId="figure-9@71.[220,1323,151,1859]" captionTargetPageId="71" captionText="Figure 29. AM M.8502, holotype skull of Canis hallstromi Troughton, 1957. (Photography by Stuart Humphreys)." figureDoi="http://doi.org/10.5281/zenodo.5237864" httpUri="https://zenodo.org/record/5237864/files/figure.png" pageId="70" pageNumber="347">Fig. 29</figureCitation>
), flat skin and skeleton, collected by J. A. Sinclair in 1956. Registered
<date box="[1259,1463,566,590]" pageId="70" pageNumber="347" value="1964-11-06">6 November 1964</date>
. An entry against the specimen “KBO
<date box="[1257,1456,595,620]" pageId="70" pageNumber="347" value="1964-11-06">6 November 1964</date>
” possibly means “killed by order”.
</paragraph>
<paragraph blockId="70.[839,1468,670,812]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,953,670,694]" pageId="70" pageNumber="347">Condition</emphasis>
. Cranium and both dentaries: bones are porous on the labial side of all gum lines, exposing the tooth roots; cranium missing 1st upper right incisor and 1st upper right post-canine tooth. Whole skeleton is partly articulated. Flat skin: minor holes present throughout the skin.
</paragraph>
<paragraph blockId="70.[837,1467,833,1004]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,1161,833,857]" pageId="70" pageNumber="347">Cranial measurements (mm)</emphasis>
. M.8502: GL, 162.72; ConL, 152.85; BasL, 145.36; NasL, —; NasB, 14.60; UC1C1 (alv.), 26.33*; APV, 10.23; PAL, 79.02; UPM (alv.), 14.51*; UMR (alv.), 13.22*; ZB, 101.05; POC, 27.78; BUL, 22.76; MB, 61.65; DL (condyl.), 121.58; LPM (alv.), 9.50*; LMR (alv.), 29.77*. [* = estimate, bone eroded].
</paragraph>
<paragraph blockId="70.[837,1467,1025,1196]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,1004,1025,1049]" pageId="70" pageNumber="347">Type locality</emphasis>
.
<materialsCitation collectorName="Dwyer &amp; Minnegal" country="Papua New Guinea" county="Lavani Valley" location="Lavani Valley" municipality="The" pageId="70" pageNumber="347" specimenCount="1" stateProvince="Southern Highlands" typeStatus="holotype">
<location LSID="urn:lsid:plazi:treatment:03DD87C8FFFF737D1B9FFF7CFD8D978A:8EAB6005FFFF73731C33FBEAFB7E9325" box="[1025,1195,1025,1049]" country="Papua New Guinea" county="Lavani Valley" municipality="The" name="Lavani Valley" pageId="70" pageNumber="347" stateProvince="Southern Highlands">Lavani Valley</location>
,
<location LSID="urn:lsid:plazi:treatment:03DD87C8FFFF737D1B9FFF7CFD8D978A:8EAB6005FFFF73731C8EFBEAFA959324" box="[1212,1344,1025,1049]" country="Papua New Guinea" county="Lavani Valley" municipality="The" name="Huri Duna" pageId="70" pageNumber="347" stateProvince="Southern Highlands">Huri Duna</location>
,
<collectingRegion country="Papua New Guinea" name="Southern Highlands" pageId="70" pageNumber="347">Southern Highlands</collectingRegion>
,
<collectingCountry box="[971,1194,1054,1078]" name="Papua New Guinea" pageId="70" pageNumber="347">Papua New Guinea</collectingCountry>
.
<collectingMunicipality box="[1208,1250,1054,1078]" pageId="70" pageNumber="347">The</collectingMunicipality>
provenance of the
<typeStatus box="[840,933,1084,1108]" pageId="70" pageNumber="347">holotype</typeStatus>
remains unsubstantiated as it is not clear whether it came from the
<collectingCounty box="[1015,1161,1113,1137]" pageId="70" pageNumber="347">Lavani Valley</collectingCounty>
or an adjoining region, nor is it known if the type specimens were wild or village animals (
<collectorName box="[845,919,1172,1196]" pageId="70" pageNumber="347">Dwyer</collectorName>
&amp;
<collectorName box="[954,1059,1172,1196]" pageId="70" pageNumber="347">Minnegal</collectorName>
, 2016)
</materialsCitation>
.
</paragraph>
<paragraph blockId="70.[839,1468,1217,1329]" pageId="70" pageNumber="347">
<materialsCitation collectingDate="1967-07-05" collectorName="Dwyer &amp; Minnegal" country="Papua New Guinea" county="Lavani Valley" location="Lavani Valley" municipality="The" pageId="70" pageNumber="347" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<emphasis bold="true" box="[839,943,1217,1241]" pageId="70" pageNumber="347">
<typeStatus box="[839,943,1217,1241]" pageId="70" pageNumber="347">Paratype</typeStatus>
</emphasis>
. M.8917 by subsequent designation (
<bibRefCitation author="Troughton, E" pageId="70" pageNumber="347" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton, 1971</bibRefCitation>
). Female, skull, flat skin and skeleton, donated by Taronga Park Zoo, 1967. Same locality and collection data as holotype. Registered
<date box="[1095,1224,1305,1329]" pageId="70" pageNumber="347" value="1967-07-05">
<collectingDate box="[1095,1224,1305,1329]" pageId="70" pageNumber="347" value="1967-07-05">5 July 1967</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="72" lastPageNumber="349" pageId="70" pageNumber="347" type="discussion">
<paragraph blockId="70.[839,1469,1351,1609]" pageId="70" pageNumber="347">
<emphasis bold="true" box="[839,962,1350,1374]" pageId="70" pageNumber="347">Comments</emphasis>
. Troughtons description was based on external criteria taken from two live animals (adult male and young female), sent to Taronga Zoo, Sydney in 1956.
<bibRefCitation author="Troughton, E" pageId="70" pageNumber="347" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton (1971)</bibRefCitation>
cited the registration numbers of
<typeStatus box="[1319,1417,1438,1462]" pageId="70" pageNumber="347">holotype</typeStatus>
and
<typeStatus box="[839,933,1468,1492]" pageId="70" pageNumber="347">paratype</typeStatus>
.A re-diagnosis is provided by
<bibRefCitation author="Koler-Matznick, J &amp; Brisbin, Jr, M" pageId="70" pageNumber="347" pagination="109 - 118" refId="ref134761" refString="Koler-Matznick, J., I. L. Brisbin, Jr, M. Feinstein, and S. Bulmer. 2003. An updated description of the New Guinea singing dog (Canis hallstromi, Troughton 1957). Journal of Zoology (London) 261: 109 - 118. https: // doi. org / 10.1017 / S 0952836903004060" type="journal article" year="2003">
Koler-Matznick
<emphasis box="[1414,1465,1468,1492]" italics="true" pageId="70" pageNumber="347">et al.</emphasis>
(2003)
</bibRefCitation>
. Jackson &amp; Groves (2015: 287) consider
<taxonomicName authorityName="Troughton" authorityYear="1957" box="[1358,1467,1497,1521]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="hallstromi">
<emphasis box="[1358,1467,1497,1521]" italics="true" pageId="70" pageNumber="347">hallstromi</emphasis>
</taxonomicName>
to be an ancient breed of the domestic dog, and provide a useful discussion of the problems of applying subspecific nomenclature to introduced populations.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5237864" ID-Zenodo-Dep="5237864" box="[222,1321,1888,1910]" httpUri="https://zenodo.org/record/5237864/files/figure.png" pageId="71" pageNumber="348" startId="71.[222,285,1888,1910]" targetBox="[256,1285,154,1853]" targetPageId="71">
<paragraph blockId="71.[222,1321,1888,1910]" box="[222,1321,1888,1910]" pageId="71" pageNumber="348">
Figure 29. AM M.8502, holotype skull of
<taxonomicName authority="Troughton, 1957" authorityName="Troughton" authorityYear="1957" box="[626,955,1888,1910]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="71" pageNumber="348" phylum="Chordata" rank="species" species="hallstromi">
<emphasis box="[626,786,1888,1910]" italics="true" pageId="71" pageNumber="348">Canis hallstromi</emphasis>
Troughton, 1957
</taxonomicName>
. (Photography by Stuart Humphreys).
</paragraph>
</caption>
<paragraph blockId="72.[360,600,151,182]" box="[360,600,151,182]" pageId="72" pageNumber="349">
<emphasis bold="true" box="[360,600,151,182]" pageId="72" pageNumber="349">
Order
<taxonomicName authorityName="Bowdich" authorityYear="1821" box="[463,600,151,182]" class="Mammalia" kingdom="Animalia" order="Rodentia" pageId="72" pageNumber="349" phylum="Chordata" rank="order">Rodentia</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>