<documentid="36A0B7C7CDB79BFF70F98B5E5D4A37DF"ID-CLB-Dataset="54097"ID-DOI="10.11646/megataxa.4.1.1"ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb"ID-ISSN="2703-3090"ID-Zenodo-Dep="4591138"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="jonas"IM.taxonomicNames_approvedBy="jonas"IM.treatments_approvedBy="felipe"checkinTime="1614033145335"checkinUser="plazi"docAuthor="Kim, Il-Hoi & Boxshall, Geoff A."docDate="2020"docId="03C487CBED4D3828FCEFF94BFA6FFCC8"docLanguage="en"docName="megataxa.4.1.1.pdf.imf"docOrigin="Megataxa 4 (1)"docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01"docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article"docStyleId="50B6B96D96A55C21793B52C23E95B310"docStyleName="Megataxa.2020-.journal_article"docStyleVersion="5"docTitle="Ammonixys quadridens Kim & Boxshall 2020, gen. etsp. nov."docType="treatment"docVersion="11"lastPageNumber="542"masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1"masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)"masterLastPageNumber="6"masterPageNumber="1"pageNumber="541"updateTime="1738783121566"updateUser="jonas">
<mods:titleid="60DFECBE8427288F2B21C3DC20C90A3F">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
<taxonomicNameid="4C6D4D5EED4D3829FCEFF94BFB82F961"ID-CoL="7ZFY3"authority="Kim & Boxshall, 2020"authorityName="Kim & Boxshall"authorityYear="2020"box="[810,1095,1669,1696]"class="Copepoda"family="Notodelphyidae"genus="Ammonixys"kingdom="Animalia"order="Cyclopoida"pageId="540"pageNumber="541"phylum="Arthropoda"rank="species"species="quadridens"status="gen. et sp. nov.">
<figureCitationid="13562A58ED4D3829FCF7F966FC58F900"box="[818,925,1703,1730]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="540"pageNumber="541">Figs. 361</figureCitation>
,
<figureCitationid="13562A58ED4D3829FC6CF969FC10F903"box="[937,981,1703,1730]"captionStart="FIGURE 362"captionStartId="543.[136,245,1465,1491]"captionTargetBox="[144,1414,230,1423]"captionTargetId="figure-194@543.[136,1451,197,1433]"captionTargetPageId="543"captionText="FIGURE 362. Ammonixys quadridens gen. et sp. nov., female. A, maxillule; B, maxilla; C, maxilliped; D, leg 1; E, leg 2; F, leg 3; G, leg 4. Scale bars: A–C, 0.01 mm; D–G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592038"httpUri="https://zenodo.org/record/4592038/files/figure.png"pageId="540"pageNumber="541">362</figureCitation>
<collectionCodeid="ED7CAE18ED4D3829FA93F8BAFA69F84F"box="[1366,1452,1908,1934]"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="540"pageNumber="541">MNHN</collectionCode>
<collectingDateid="EF97E9F5ED4D3829FCACF858FBC7F871"box="[873,1026,1942,1968]"pageId="540"pageNumber="541"value="2010-05-01">01 May 2010</collectingDate>
<figureCitationid="13562A58ED4C3828FDCEFF63FD5CFF06"box="[523,665,173,199]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361A, B</figureCitation>
)globular, greatly swollen: body length 1.12 mm from frontal margin of cephalosome to posterior margin of fourth pedigerous somite. Prosome with smooth surface; lateral margins of dorsal cephalic shield expanded ventrally, completely concealing mouthparts in lateral view. Metasome 4- segmented; first and second articulations complete, third articulation incomplete; first to third pedigerous somites much wider than long, Fourth pedigerous somite greatly expanded, spherical, 785×774 μm. Freeurosome (
<figureCitationid="13562A58ED4C3828FD19FE11FF03FDDD"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361C</figureCitation>
) small, stout, 5-segmented, inserted into ventral surface of brood pouch (
<figureCitationid="13562A58ED4C3828FE5CFDEDFDCBFDFF"box="[409,526,547,574]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361B</figureCitation>
). All urosomites much widerthan long; 4 abdominal somitessparsely ornamented with minute spinules. Caudal ramus (
<figureCitationid="13562A58ED4C3828FDF8FDA9FD76FD43"box="[573,691,615,642]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361C</figureCitation>
) small, lobate, slightly longerthan wide, not articulated from anal somite, bearing minute setules; caudal setae apparently absent.
<figureCitationid="13562A58ED4C3828FF55FCFDFF3BFC8F"box="[144,254,819,846]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361F</figureCitation>
) archedposteriorly, 117 μmlong, 6-segmented; third and fourth segments articulated from each other only on one surface; first segment with 2 setae; setation of other segments obscure due to similarity with setules; third, fifth and terminal segments each bearing 1 small aesthetasc. Antenna (
<figureCitationid="13562A58ED4C3828FE4FFC13FDCCFC39"box="[394,521,989,1016]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361G</figureCitation>
) stout, 4-segmented; coxa, basis, and first endopodal segment unarmed; basis shorter than first endopodal segment; narrower second endopodal segment about 2.3 times longer than wide (41×18 μm) and as long as first segment: armed with 3 small setae plus small terminal claw, about one-third as long as second endopodal segment.
<figureCitationid="13562A58ED4C3828FEEBFB05FE6CFB27"box="[302,425,1227,1254]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig. 361H</figureCitation>
) simple, unornamented, with short,broadposteromedian lobe.Mandible (
<figureCitationid="13562A58ED4C3828FDA2FB23FD09FAC9"box="[615,716,1261,1288]"captionStart="FIGURE 361"captionStartId="542.[136,245,1907,1933]"captionTargetBox="[219,1354,205,1847]"captionTargetId="figure-21@542.[191,1397,185,1870]"captionTargetPageId="542"captionText="FIGURE 361. Ammonixys quadridens gen. et sp. nov., female. A, habitus, dorsal; B, habitus, ventrolateral; C, leg 5 and urosome, ventral; D, anterior part of prosome, ventral; E, rostrum; F, antennule; G, antenna; H, labrum; I, mandible. Scale bars: A, B, 0.2 mm; C, D, 0.05 mm; E–I, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592036"httpUri="https://zenodo.org/record/4592036/files/figure.png"pageId="541"pageNumber="542">Fig.361I</figureCitation>
) with coxal gnathobase bearing 4 teeth on distal half of medial margin and pectinate proximal half of medial margin: palp consisting of basis and 2-segmented endopod; basis with 1 seta mediodistally; exopod completely fused with basis, represented by outer lobe of basis and pair of large setae at base of lobe; endopod 2-segmented, incompletely articulated from basis, with 1 broad seta on first segment and 4 setaeon second. Maxillule (
<figureCitationid="13562A58ED4C3828FDCDFA33FDB9F9D9"box="[520,636,1533,1560]"captionStart="FIGURE 362"captionStartId="543.[136,245,1465,1491]"captionTargetBox="[144,1414,230,1423]"captionTargetId="figure-194@543.[136,1451,197,1433]"captionTargetPageId="543"captionText="FIGURE 362. Ammonixys quadridens gen. et sp. nov., female. A, maxillule; B, maxilla; C, maxilliped; D, leg 1; E, leg 2; F, leg 3; G, leg 4. Scale bars: A–C, 0.01 mm; D–G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592038"httpUri="https://zenodo.org/record/4592038/files/figure.png"pageId="541"pageNumber="542">Fig. 362A</figureCitation>
) armedwith 6 setae on arthrite, 1 on coxal epipodite, 1 on basis, 4 on exopod and 3 on endopod; coxal endite absent. Maxilla (
<figureCitationid="13562A58ED4C3828FF55F9ADFECEF9BF"box="[144,267,1635,1662]"captionStart="FIGURE 362"captionStartId="543.[136,245,1465,1491]"captionTargetBox="[144,1414,230,1423]"captionTargetId="figure-194@543.[136,1451,197,1433]"captionTargetPageId="543"captionText="FIGURE 362. Ammonixys quadridens gen. et sp. nov., female. A, maxillule; B, maxilla; C, maxilliped; D, leg 1; E, leg 2; F, leg 3; G, leg 4. Scale bars: A–C, 0.01 mm; D–G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592038"httpUri="https://zenodo.org/record/4592038/files/figure.png"pageId="541"pageNumber="542">Fig. 362B</figureCitation>
) 4-segmented; syncoxa bearing 2 endites, with 3 setae on first endite and 2 on second; basis with strong claw plus 1 seta; small endopod with 1 seta on first segment and 3 setaeon second. Maxilliped (
<figureCitationid="13562A58ED4C3828FDB6F907FD21F925"box="[627,740,1737,1764]"captionStart="FIGURE 362"captionStartId="543.[136,245,1465,1491]"captionTargetBox="[144,1414,230,1423]"captionTargetId="figure-194@543.[136,1451,197,1433]"captionTargetPageId="543"captionText="FIGURE 362. Ammonixys quadridens gen. et sp. nov., female. A, maxillule; B, maxilla; C, maxilliped; D, leg 1; E, leg 2; F, leg 3; G, leg 4. Scale bars: A–C, 0.01 mm; D–G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592038"httpUri="https://zenodo.org/record/4592038/files/figure.png"pageId="541"pageNumber="542">Fig. 362C</figureCitation>
<figureCitationid="13562A58ED4C3828FEF6F8C3FE57F8E9"box="[307,402,1805,1832]"captionStart="FIGURE 362"captionStartId="543.[136,245,1465,1491]"captionTargetBox="[144,1414,230,1423]"captionTargetId="figure-194@543.[136,1451,197,1433]"captionTargetPageId="543"captionText="FIGURE 362. Ammonixys quadridens gen. et sp. nov., female. A, maxillule; B, maxilla; C, maxilliped; D, leg 1; E, leg 2; F, leg 3; G, leg 4. Scale bars: A–C, 0.01 mm; D–G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.4592038"httpUri="https://zenodo.org/record/4592038/files/figure.png"pageId="541"pageNumber="542">Fig. 362</figureCitation>
D-G) biramous with 3-segmented exopods; endopods 3-segmented inlegs 1–3, but 2- segmented in leg 4. Protopod 2-segmented in leg 1, but unsegmented in legs 2–4. Inner coxal seta absent in legs 1–4. Leg 1 lacking innerdistal element on basis. Outer seta on basis present only in leg 3. Endopod of leg 1 strongly curved inwards. First exopodal segment of legs 3 and 4 and second exopodal segment of legs 2 and 4 each bearing spiniform outer distal process. Outer setae on third exopodal segment of legs 2–4 vestigial. Armature formula for legs 1–4 as follows:
<taxonomicNameLabelid="A22A57B4ED4C3828FCEFFD65FC1DFD04"box="[810,984,683,709]"pageId="541"pageNumber="542"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicNameid="4C6D4D5EED4C3828FB38FD62FA6EFD04"authority="Kim & Boxshall, 2020"authorityName="Kim & Boxshall"authorityYear="2020"box="[1277,1451,683,709]"class="Copepoda"family="Notodelphyidae"genus="Doroixys"higherTaxonomySource="GBIF"kingdom="Animalia"order="Cyclopoida"pageId="541"pageNumber="542"phylum="Arthropoda"rank="species"species="obesa"status="sp. nov.">