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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1034.59413" ID-GBIF-Dataset="13846ea1-7b83-4f5e-9690-acc423f5a086" ID-PMC="PMC8093189" ID-Pensoft-Pub="1313-2970-1034-1" ID-Pensoft-UUID="2DB5C14D37835632AB3585A3AECC3B1C" ID-PubMed="33958928" ID-ZooBank="A2937A0DFF04468FB2DB6AC4D68ED997" ModsDocID="1313-2970-1034-1" checkinTime="1619530777733" checkinUser="pensoft" docAuthor="Zhang, Jianshuang, Yu, Hao & Li, Shuqiang" docDate="2021" docId="2FE4A8C400ED5CA3A665AFE983608CB0" docLanguage="en" docName="ZooKeys 1034: 1-163" docOrigin="ZooKeys 1034" docPubDate="2021-04-26" docSource="http://dx.doi.org/10.3897/zookeys.1034.59413" docTitle="Clubiona subdidentata Yu & Li 2021, sp. nov." docType="treatment" docUuid="B4B90700-E09E-430A-9B58-C80D6CF84084" docUuidSource="ZooBank" docVersion="4" id="2DB5C14D37835632AB3585A3AECC3B1C" lastPageNumber="1" masterDocId="2DB5C14D37835632AB3585A3AECC3B1C" masterDocTitle="Taxonomic studies on the sac spider genus Clubiona (Araneae, Clubionidae) from Xishuangbanna Rainforest, China" masterLastPageNumber="163" masterPageNumber="1" pageNumber="1" updateTime="1668150202507" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic studies on the sac spider genus Clubiona (Araneae, Clubionidae) from Xishuangbanna Rainforest, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Jianshuang</mods:namePart>
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<mods:affiliation>School of Life Sciences</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yu, Hao</mods:namePart>
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<mods:affiliation>Guizhou Normal University, Guiyang, Guizhou, China</mods:affiliation>
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<mods:nameIdentifier type="email">insect1986@126.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
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<mods:affiliation>School of Biological Sciences, Guizhou Education University, Guiyang, Guizhou, China</mods:affiliation>
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<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1034</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>163</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1034.59413</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1034.59413</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1034-1</mods:identifier>
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<mods:identifier type="ZooBank">A2937A0DFF04468FB2DB6AC4D68ED997</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2DB5C14D37835632AB3585A3AECC3B1C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181652928" LSID="urn:lsid:zoobank.org:act:B4B90700-E09E-430A-9B58-C80D6CF84084" httpUri="http://treatment.plazi.org/id/2FE4A8C400ED5CA3A665AFE983608CB0" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/B4B90700-E09E-430A-9B58-C80D6CF84084" authority="Yu & Li" authorityName="Yu & Li" authorityYear="2021" class="Arachnida" family="Clubionidae" genus="Clubiona" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clubiona subdidentata" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subdidentata" status="sp. nov.">Clubiona subdidentata Yu & Li</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Holotype female of Clubiona subdidentata sp. nov., epigyne (A-E) and habitus (F, G) A intact, ventral view B cleared, ventral view C cleared, ventral view D cleared, dorsal view E cleared, dorsal view F dorsal view G ventral view. Abbreviations: AM = atrial membrane; BS = bursa; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; SP = spermatheca. Scale bars: 0.1 mm (equal for A-E); 1 mm (equal for F, G)." figureDoi="10.3897/zookeys.1034.59413.figure15" httpUri="https://binary.pensoft.net/fig/535424" pageId="0" pageNumber="1">Figs 15</figureCitation>
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<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Clubiona spp. of the C. corticalis group, epigyne, intact, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure74" httpUri="https://binary.pensoft.net/fig/535483" pageId="0" pageNumber="1">, 74F</figureCitation>
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<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Clubiona spp. of the C. corticalis group, epigyne, cleared, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure82" httpUri="https://binary.pensoft.net/fig/535491" pageId="0" pageNumber="1">, 82F</figureCitation>
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<figureCitation captionStart="Figure 90" captionStartId="F90" captionText="Figure 90. Clubiona spp. of the C. corticalis group, vulva, cleared, dorsal view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: BS = bursa; CD = copulatory duct; FD = fertilisation duct; SH = spermathecal head; SP = spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure90" httpUri="https://binary.pensoft.net/fig/535499" pageId="0" pageNumber="1">, 90F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="holotype">
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<paragraph pageId="0" pageNumber="1">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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♀ (IZCAS-Ar 34749), China: Yunnan Province: Xishuangbanna: Mengla County: Xiaolongha Village:
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<geoCoordinate degrees="22" direction="north" minutes="5.017" orientation="latitude" precision="1" value="22.083616">22°5.017'N</geoCoordinate>
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,
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<geoCoordinate degrees="100" direction="east" minutes="22.084" orientation="longitude" precision="1" value="100.368065">100°22.084'E</geoCoordinate>
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, ca. 1118 m, 24.VII.2012, Q.Y. Zhao and Z.G. Chen leg.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Other material examined.</emphasis>
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1♀ (YHCLU0073), same data as holotype
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specific name is taken from its similarity to
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<taxonomicName class="Arachnida" family="Clubionidae" kingdom="Animalia" lsidName="C. didentata" order="Araneae" pageId="0" pageNumber="1" rank="species" species="didentata">
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<emphasis italics="true" pageId="0" pageNumber="1">C. didentata</emphasis>
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</taxonomicName>
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; adjective.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The female of
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<taxonomicName class="Arachnida" family="Clubionidae" kingdom="Animalia" lsidName="C. subdidentata" order="Araneae" pageId="0" pageNumber="1" rank="species" species="subdidentata">
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<emphasis italics="true" pageId="0" pageNumber="1">C. subdidentata</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished from all other members of the
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<taxonomicName class="Arachnida" family="Clubionidae" kingdom="Animalia" lsidName="C. corticalis" order="Araneae" pageId="0" pageNumber="1" rank="species" species="corticalis">
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<emphasis italics="true" pageId="0" pageNumber="1">C. corticalis</emphasis>
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</taxonomicName>
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group with the exception of
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<taxonomicName class="Arachnida" family="Clubionidae" kingdom="Animalia" lsidName="C. didentata" order="Araneae" pageId="0" pageNumber="1" rank="species" species="didentata">
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<emphasis italics="true" pageId="0" pageNumber="1">C. didentata</emphasis>
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</taxonomicName>
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(
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4679.2.1" author="Yu, H" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="201 - 230" refId="B74" refString="Yu, H, Li, S, 2019b. On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China. Zootaxa 4679 (2): 201 - 230, DOI: https://doi.org/10.11646/zootaxa.4679.2.1" title="On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China." url="https://doi.org/10.11646/zootaxa.4679.2.1" volume="4679" year="2019 b">Yu and Li 2019b</bibRefCitation>
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: 207, figs 6A-D; Figs
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<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Clubiona spp. of the C. corticalis group, epigyne, intact, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure74" httpUri="https://binary.pensoft.net/fig/535483" pageId="0" pageNumber="1">74E</figureCitation>
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,
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<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Clubiona spp. of the C. corticalis group, epigyne, cleared, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure82" httpUri="https://binary.pensoft.net/fig/535491" pageId="0" pageNumber="1">82E</figureCitation>
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,
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<figureCitation captionStart="Figure 90" captionStartId="F90" captionText="Figure 90. Clubiona spp. of the C. corticalis group, vulva, cleared, dorsal view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: BS = bursa; CD = copulatory duct; FD = fertilisation duct; SH = spermathecal head; SP = spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure90" httpUri="https://binary.pensoft.net/fig/535499" pageId="0" pageNumber="1">90E</figureCitation>
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) by having an atrial membrane (atrial membrane is absent in almost all species of the
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<taxonomicName class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" lsidName="Clubiona corticalis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="corticalis">
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<emphasis italics="true" pageId="0" pageNumber="1">Clubiona corticalis</emphasis>
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</taxonomicName>
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group) and similar vulva but can be recognised by the nearly equilateral triangular atrial membrane (Figs
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Holotype female of Clubiona subdidentata sp. nov., epigyne (A-E) and habitus (F, G) A intact, ventral view B cleared, ventral view C cleared, ventral view D cleared, dorsal view E cleared, dorsal view F dorsal view G ventral view. Abbreviations: AM = atrial membrane; BS = bursa; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; SP = spermatheca. Scale bars: 0.1 mm (equal for A-E); 1 mm (equal for F, G)." figureDoi="10.3897/zookeys.1034.59413.figure15" httpUri="https://binary.pensoft.net/fig/535424" pageId="0" pageNumber="1">15A-C</figureCitation>
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,
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<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Clubiona spp. of the C. corticalis group, epigyne, intact, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure74" httpUri="https://binary.pensoft.net/fig/535483" pageId="0" pageNumber="1">74F</figureCitation>
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,
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<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Clubiona spp. of the C. corticalis group, epigyne, cleared, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure82" httpUri="https://binary.pensoft.net/fig/535491" pageId="0" pageNumber="1">82F</figureCitation>
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) (vs. tongue- shaped in
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<taxonomicName class="Arachnida" family="Clubionidae" kingdom="Animalia" lsidName="C. didentata" order="Araneae" pageId="0" pageNumber="1" rank="species" species="didentata">
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<emphasis italics="true" pageId="0" pageNumber="1">C. didentata</emphasis>
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</taxonomicName>
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; Figs
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<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Clubiona spp. of the C. corticalis group, epigyne, intact, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure74" httpUri="https://binary.pensoft.net/fig/535483" pageId="0" pageNumber="1">74E</figureCitation>
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,
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<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Clubiona spp. of the C. corticalis group, epigyne, cleared, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure82" httpUri="https://binary.pensoft.net/fig/535491" pageId="0" pageNumber="1">82E</figureCitation>
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) and by the distinct copulatory ducts (Figs
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Holotype female of Clubiona subdidentata sp. nov., epigyne (A-E) and habitus (F, G) A intact, ventral view B cleared, ventral view C cleared, ventral view D cleared, dorsal view E cleared, dorsal view F dorsal view G ventral view. Abbreviations: AM = atrial membrane; BS = bursa; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; SP = spermatheca. Scale bars: 0.1 mm (equal for A-E); 1 mm (equal for F, G)." figureDoi="10.3897/zookeys.1034.59413.figure15" httpUri="https://binary.pensoft.net/fig/535424" pageId="0" pageNumber="1">15D, E</figureCitation>
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,
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<figureCitation captionStart="Figure 90" captionStartId="F90" captionText="Figure 90. Clubiona spp. of the C. corticalis group, vulva, cleared, dorsal view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: BS = bursa; CD = copulatory duct; FD = fertilisation duct; SH = spermathecal head; SP = spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure90" httpUri="https://binary.pensoft.net/fig/535499" pageId="0" pageNumber="1">90F</figureCitation>
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) (copulatory ducts absent in
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<taxonomicName class="Arachnida" family="Clubionidae" kingdom="Animalia" lsidName="C. didentata" order="Araneae" pageId="0" pageNumber="1" rank="species" species="didentata">
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<emphasis italics="true" pageId="0" pageNumber="1">C. didentata</emphasis>
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</taxonomicName>
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; Fig.
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<figureCitation captionStart="Figure 90" captionStartId="F90" captionText="Figure 90. Clubiona spp. of the C. corticalis group, vulva, cleared, dorsal view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: BS = bursa; CD = copulatory duct; FD = fertilisation duct; SH = spermathecal head; SP = spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure90" httpUri="https://binary.pensoft.net/fig/535499" pageId="0" pageNumber="1">90E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Female.</emphasis>
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Holotype (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Holotype female of Clubiona subdidentata sp. nov., epigyne (A-E) and habitus (F, G) A intact, ventral view B cleared, ventral view C cleared, ventral view D cleared, dorsal view E cleared, dorsal view F dorsal view G ventral view. Abbreviations: AM = atrial membrane; BS = bursa; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; SP = spermatheca. Scale bars: 0.1 mm (equal for A-E); 1 mm (equal for F, G)." figureDoi="10.3897/zookeys.1034.59413.figure15" httpUri="https://binary.pensoft.net/fig/535424" pageId="0" pageNumber="1">15F, G</figureCitation>
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): Total length 3.63; carapace 1.45 long, 1.14 wide; opisthosoma 2.18 long, 1.32 wide. Carapace uniformly greyish white, without any pattern or markings; ocular region distinctly narrowed, cervical groove and radial grooves indistinct; tegument smooth, all setae detached in ethanol. Eyes: AER slightly recurved, PER wider than AER and almost straight in dorsal view. Eye sizes and interdistances: AME 0.09, ALE 0.09, PME 0.09, PLE 0.07, AME-AME 0.06, AME-ALE 0.07, PME-PME 0.19, PME-PLE 0.10, MOQL 0.21, MOQA 0.23, MOQP 0.38. Chelicerae light orange, with three promarginal and two retromarginal teeth. Sternum pale brown, 0.80 long, 0.65 wide. Labium and endites coloured as chelicerae. Legs greyish white, uniformly coloured. Leg measurements: I 2.73 (0.81, 1.15, 0.55, 0.23), II 3.15 (0.93, 1.25, 0.74, 0.22), III 2.85 (0.93, 0.95, 0.70, 0.29), IV 3.39 (1.19, 1.32, 1.05, 0.39). Abdomen oval, nearly pure white, with inconspicuous anterior setal tufts, dorsum with two pairs of inconspicuous muscle depressions; venter without pattern.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Epigyne (Figs
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Holotype female of Clubiona subdidentata sp. nov., epigyne (A-E) and habitus (F, G) A intact, ventral view B cleared, ventral view C cleared, ventral view D cleared, dorsal view E cleared, dorsal view F dorsal view G ventral view. Abbreviations: AM = atrial membrane; BS = bursa; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; SP = spermatheca. Scale bars: 0.1 mm (equal for A-E); 1 mm (equal for F, G)." figureDoi="10.3897/zookeys.1034.59413.figure15" httpUri="https://binary.pensoft.net/fig/535424" pageId="0" pageNumber="1">15A-E</figureCitation>
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,
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<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Clubiona spp. of the C. corticalis group, epigyne, intact, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure74" httpUri="https://binary.pensoft.net/fig/535483" pageId="0" pageNumber="1">74F</figureCitation>
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,
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<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Clubiona spp. of the C. corticalis group, epigyne, cleared, ventral view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: A = atrium; AM = atrial membrane; CO = copulatory opening. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure82" httpUri="https://binary.pensoft.net/fig/535491" pageId="0" pageNumber="1">82F</figureCitation>
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,
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<figureCitation captionStart="Figure 90" captionStartId="F90" captionText="Figure 90. Clubiona spp. of the C. corticalis group, vulva, cleared, dorsal view A C. xiaokong sp. nov., holotype B C. zhaoi sp. nov., holotype C C. kai D C. tiane E C. didentata F C. subdidentata sp. nov., holotype. Abbreviations: BS = bursa; CD = copulatory duct; FD = fertilisation duct; SH = spermathecal head; SP = spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1034.59413.figure90" httpUri="https://binary.pensoft.net/fig/535499" pageId="0" pageNumber="1">90F</figureCitation>
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). Epigynal plate ca. 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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wider than long, margin not delimited; spermathecae and bursae prominently visible through epigynal plate. Atrium small, anteriorly covered by an atrial membrane. Atrial membrane shaped nearly like an equilateral triangle, with a blunt apex. Copulatory openings small, located at basolateral atrial borders. Copulatory ducts distinct, extend transversally, connecting to posteriorly located bursae. Spermathecae small, consisting of a bean-shaped proximal part and an acicular distal part, with short fertilisation ducts terminally. Bursae reniform, close together, ca. 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide, bursal surface hyaline and smooth, inside pigmented and sclerotised.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Male.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known only from the type locality, Xishuangbanna, Yunnan, China.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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