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92 lines
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<document id="9503AE2CC4C3FBD5CA729F49F912652A" ID-DOI="http://dx.doi.org/10.3897/zookeys.482.8343" ID-GBIF-Dataset="4fdb2d6d-b69c-4e87-9fb3-1de661126876" ID-PMC="PMC4337222" ID-Pensoft-Pub="1313-2970-482-91" ID-PubMed="25709532" ID-ZooBank="21C3B7D0B18743EABB38175C704D7550" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-482-91" ModsDocOrigin="ZooKeys 482" ModsDocTitle="Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898 (Coleoptera, Scarabaeidae, Cetoniinae)" checkinTime="1451244653051" checkinUser="pensoft" docAuthor="Rojkoff, Sebastien & Perissinotto, Renzo" docDate="2015" docId="15C68E87E7B91A6EDF5A17F22916B7F6" docLanguage="en" docName="ZooKeys 482: 91-142" docOrigin="ZooKeys 482" docSource="http://dx.doi.org/10.3897/zookeys.482.8343" docTitle="Atrichelaphinis (Eugeaphinis) bomboesbergica Rojkoff & Perissinotto, 2015, sp. n." docType="treatment" docUuid="CE128E12-51B9-4143-8AA3-72E301FBFCB0" docUuidSource="ZooBank" docVersion="6" lastPageNumber="112" masterDocId="822C54210D6A7F60FFE2FFC9FFA1A039" masterDocTitle="Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898 (Coleoptera, Scarabaeidae, Cetoniinae)" masterLastPageNumber="142" masterPageNumber="91" pageNumber="112" updateTime="1701316131199" updateUser="plazi">
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<mods:titleInfo id="89B7C91BBA7B808E396F302CF721321D">
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<mods:title id="8A3B665F33B6B0E3CC54EEC095CEE28A">Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898 (Coleoptera, Scarabaeidae, Cetoniinae)</mods:title>
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<mods:namePart id="DBAE7F772D4492A5A9F62A5440D10607">Rojkoff, Sebastien</mods:namePart>
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<mods:name id="43AA9FBBF5ED4DF294551DB00B3BD5EA" type="personal">
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<mods:namePart id="93B9917507698813F98293E1FA9C1E82">Perissinotto, Renzo</mods:namePart>
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<mods:typeOfResource id="993DC1EDABCB4A49EC541D14A5FFB8F8">text</mods:typeOfResource>
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<mods:title id="BB5576935D5079A64633FDCD147C71C9">ZooKeys</mods:title>
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<mods:part id="8D063A57445FCD84ABE4C5C035AE9766">
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<mods:date id="1727E6C4E74B0D6B6AF19ADA71580725">2015</mods:date>
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<mods:detail id="44B9F06601C71E4A23BD0F9E1813160B" type="volume">
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<mods:number id="DAF532D66990DC6B303911F97FFD113F">482</mods:number>
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</mods:detail>
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<mods:start id="B8A15C40967474A76A3288E0636084A3">91</mods:start>
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<mods:end id="5EF9A75BA2342A6DE68F6E5E9361BAD4">142</mods:end>
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<mods:url id="188149874BF03DDCE6F59F6FE507E8D9">http://dx.doi.org/10.3897/zookeys.482.8343</mods:url>
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<mods:classification id="E5789614D284903AF0CF7DF2EF55C373">journal article</mods:classification>
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<mods:identifier id="C920DB3C77896ED339CE3064B0E5A4F3" type="DOI">http://dx.doi.org/10.3897/zookeys.482.8343</mods:identifier>
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<mods:identifier id="51A99C8FA69CC7AACCAB108C420E091B" type="Pensoft-Pub">1313-2970-482-91</mods:identifier>
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<mods:identifier id="471DF1677E18A46ACBECAB41FEE7A0AC" type="ZooBank">21C3B7D0B18743EABB38175C704D7550</mods:identifier>
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</mods:mods>
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<treatment id="15C68E87E7B91A6EDF5A17F22916B7F6" ID-GBIF-Taxon="152058288" LSID="urn:lsid:zoobank.org:act:CE128E12-51B9-4143-8AA3-72E301FBFCB0" httpUri="http://treatment.plazi.org/id/15C68E87E7B91A6EDF5A17F22916B7F6" lastPageNumber="112" pageId="21" pageNumber="112">
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<subSubSection id="F8553457DEEBA5829E0A1DA2B6DD49FC" pageId="21" pageNumber="112" type="multiple">
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<paragraph id="F5717541B4AD58857538EDA78E74FA30" pageId="21" pageNumber="112">Taxon classification Animalia Coleoptera Scarabaeidae</paragraph>
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</subSubSection>
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<subSubSection id="AF776ECCB954F22B4B7B07E5718E9A5A" pageId="21" pageNumber="112" type="nomenclature">
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<paragraph id="FF0F3207BC1D9B95FE23007E3B47B204" pageId="21" pageNumber="112">
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<taxonomicName id="A6D2DD05B2CAA35A2713571D4395C9DF" ID-CoL="5VWCN" LSID="http://zoobank.org/CE128E12-51B9-4143-8AA3-72E301FBFCB0" class="Insecta" family="Scarabaeidae" genus="Atrichelaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichelaphinis (Eugeaphinis) bomboesbergica" order="Coleoptera" pageId="21" pageNumber="112" phylum="Arthropoda" rank="species" species="bomboesbergica" subGenus="Eugeaphinis">Atrichelaphinis (Eugeaphinis) bomboesbergica</taxonomicName>
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<taxonomicNameLabel id="B64AAF19D90986DB55B1A7FE9F312DD6" pageId="21" pageNumber="112">sp. n.</taxonomicNameLabel>
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Figure 12
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</paragraph>
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</subSubSection>
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<subSubSection id="732DFF00FD7407CF4B4DAE9FA27FBA00" pageId="21" pageNumber="112" type="type specimens">
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<paragraph id="610C6C43A8B2E114DB114A3E9C7E5D54" pageId="21" pageNumber="112">Type specimens.</paragraph>
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<paragraph id="F3188E4D18F0D818A2592D7EC7D1E6FF" pageId="21" pageNumber="112">Holotype male: South Africa EC, Hofmeyr, 28-XI-2011, R. Perissinotto & L. Clennell leg (ISAM). Paratypes: 1♂, same data as holotype (ISAM, PCRP); 1♂, same data as holotype, but 10-XII-2011 (PCRP); 8♂ 3♀, same data as holotype, but 24-XII-2011 (TMSA, PCSR); 2♂ 1♀, same data as holotype, but 26-XII-2011 (PCRP); 8♀, same data as holotype, but 18-XII-2010 (PCRP, PCSR).</paragraph>
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</subSubSection>
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<subSubSection id="84E6DF8315FB8697655D24D597E26385" pageId="21" pageNumber="112" type="description">
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<paragraph id="0167EC45ED688E525EDDBDCBEF69C364" pageId="21" pageNumber="112">Description</paragraph>
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<paragraph id="789171B714BA65DB210A84944766A945" pageId="21" pageNumber="112">(n = 25). Size: length ♂, 9.4-11.7 mm; ♀, 10.1-12.8 mm; width ♂, 5.8-6.7 mm; ♀, 6.2 to 7.8 mm.</paragraph>
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<paragraph id="B8C30962D6762B6FBCA314F8CB916CC2" pageId="21" pageNumber="112">Body. Dorsal surface slightly shiny, ground colour from ochraceous to light-brown, with many black/dark brown markings and small white maculae; scale-like setae present and particularly well developed on pronotum, more extensive in male than in female.</paragraph>
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<paragraph id="40D83EAAAF2D2AEA233E7B94B13C9EC6" pageId="21" pageNumber="112">Head. Anterior margin of clypeus sharply upturned, particularly in male, sligthly bilobed apically, anterior angles weakly rounded, lateral declivity visible from above; large crescent to horseshoe punctures, particularly dense on frons and vertex; scale-type setae particularly long and dense from frons to vertex; antennae with pedicel and flagellum reddish-brown, but clubs dark brown to black, club notably longer in male than in female.</paragraph>
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<paragraph id="1FF02B173F7603CEF9CCA46D4DAE9C7B" pageId="21" pageNumber="112">Pronotum. With black markings not covering more than half of total area and particularly developed on anterior part of disc, on both sides of medial line; anterior margin tectiform; disc moderately tuberculate in front; with scale-like setae and round punctures diffuse but widespread thoughout surface, setae more dense and longer on lateral margins; lateral margins and angles smoothly rounded with ante-scutellar arch relatively straight.</paragraph>
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<paragraph id="97E9F66EF7E39C595054D3D4B0EAFDCC" pageId="21" pageNumber="112">Scutellum. With apex from weakly rounded to acute; lateral margins from straight to weakly concave, with shallow and narrow lateral grooves; prominent oblong medial black mark extending from base to middle of disc; exhibiting few fine punctures on apical half but no scale-like setae.</paragraph>
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<paragraph id="9795D7A8E3E64EB7F3D566A4C9F2EC81" pageId="21" pageNumber="112">Elytra. Weakly tricostate, with costae barely visible in apical part; sutural costa bulging out towards middle of elytral length; striae partly geminate and with coarse horseshoe sculpture; black marking most developed around humeral and apical calluses and in mid area of lateral half; apical sutural end virtually straight in male but curving outwards in female.</paragraph>
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<paragraph id="619873B138A8FD404F6C56CF5975B0E7" pageId="21" pageNumber="112">Pygidium. Brown to reddish at centre, becoming dark brown to black towards lateral and lower margins; fine sculture and dense cover of scale-like setae throughout surface; exhibiting 2-3 pairs of depressed areas close to lateral margins.</paragraph>
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<paragraph id="F505B7068CD8FAD211A1414237FF0111" pageId="21" pageNumber="112">Underside. Dark brown to black with white scattered maculae, particularly on metasternum and lateral margins of abdominal sternites; densely covered with long white setae, replaced in mid area of metasternum and abdominal sternites by few scattered scale-like setae; coarse and scattered horseshoe sculpture throughout, except on central areas of sternum and abdominal sternites; abdominal sternites weakly concave at middle in male, slightly convex in female; mesosternal apophysis ochraceous, small and rounded, with no projections extending between mesocoxae.</paragraph>
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<paragraph id="0CAA871EDE6CF4380AF3EA19B48ADFAC" pageId="21" pageNumber="112">Legs. Tibia and femora light brown, with dark brown to black tips and joints; scattered white maculae present on both dorsal and ventral sides; tarsi dark brown to black; protibia unidentate but broadening remarkably towards apex, forming spade-like structure; numerous long setae throughout surface and scale-like setae at joints; metatibial spurs thin and acute in male, slightly enlarged and blunt in female.</paragraph>
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<paragraph id="CA0DE8ABCB5747A6FA8A7C3010971767" pageId="21" pageNumber="112">Aedeagus. Parameres virtually straight from base to apical convergence, forming then a perfectly round apex, with slight indent at centre (dorsal view); apical margin curving downwards, but no ventral folding or projections visible in lateral view.</paragraph>
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</subSubSection>
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<subSubSection id="612477B2A23177513E2D00F95EA89FEE" pageId="21" pageNumber="112" type="derivatio nominis">
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<paragraph id="59BA4E6F2FD4A40D9A60CDB84841797B" pageId="21" pageNumber="112">Derivatio nominis.</paragraph>
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<paragraph id="3B8EB8176EEF3D0E8FBEE301804ACBFA" pageId="21" pageNumber="112">The species is named after the Bamboesberg mountain range of the Eastern Cape Province of South Africa, where it was discovered on its south-western slopes.</paragraph>
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</subSubSection>
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<subSubSection id="BA234C667E8C51B1F27F7B9BE6F756BF" pageId="21" pageNumber="112" type="remarks">
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<paragraph id="B543510184863261B93747A9A4127DD2" pageId="21" pageNumber="112">Remarks.</paragraph>
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<paragraph id="A3377DB660F94D8837CD1130BCCC81D2" pageId="21" pageNumber="112">
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This new species represents the southernmost extension of the genus distribution range in the Afrotropical Region.
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<taxonomicName id="39B6808257A175A03B9CC6EAE9F1AF64" class="Insecta" family="Scarabaeidae" genus="Atrichelaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichelaphinis bamboesbergica" order="Coleoptera" pageId="21" pageNumber="112" phylum="Arthropoda" rank="species" species="bamboesbergica">Atrichelaphinis bamboesbergica</taxonomicName>
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appears to be restricted to a small area of the eastern Karoo semiarid region, where its larval stages develop exclusively in the dung middens of the antbear,
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<taxonomicName id="ABFFEA87946321EB2EAAD43B3DACF133" class="Mammalia" family="Orycteropodidae" genus="Orycteropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orycteropus afer" order="Tubulidentata" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="afer">Orycteropus afer</taxonomicName>
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(Pallas, 1766). Adults have a relatively short life span (2-3 weeks) and appear to be unable to feed, as none has yet been observed either on fruits, flowers or sap flows.
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</paragraph>
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<caption id="B4B70277F2FE49D7CDAA8417E5B05566" pageId="21" pageNumber="112">
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<paragraph id="1D2ADF1B16F927BC95F993DBE1E0A101" pageId="21" pageNumber="112">
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Figure 12.
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<taxonomicName id="C1E2860E5FEE3866D9EDE47EF460C7A1" class="Insecta" family="Scarabaeidae" genus="Atrichelaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichelaphinis (Eugeaphinis) bomboesbergica" order="Coleoptera" pageId="21" pageNumber="112" phylum="Arthropoda" rank="species" species="bomboesbergica" subGenus="Eugeaphinis">Atrichelaphinis (Eugeaphinis) bomboesbergica</taxonomicName>
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sp. n., holotype male (ISAM). A Dorsal view B ventral view C parameres in dorsal view D parameres in lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document>
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