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<document id="9BACB725DC96D3671E70EC58DF9A1C6B" ID-CLB-Dataset="30549" ID-DOI="10.11646/zootaxa.4392.3.8" ID-GBIF-Dataset="9028dbc4-eb37-446d-a625-4d2ae43dccc4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1196271" ID-ZooBank="308B8770-4638-4914-8F93-508622567801" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520855525095" checkinUser="plazi" docAuthor="Agassiz, David &amp; Kallies, Axel" docDate="2018" docId="03C06C57FA71FFE201C4FD15FBB67F7E" docLanguage="en" docName="zootaxa.4392.3.8.pdf" docOrigin="Zootaxa 4392 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Osmanthedon Agassiz &amp; Kallies 2018, gen. nov." docType="treatment" docVersion="7" lastPageNumber="590" masterDocId="FFF9142FFA70FFE00153FFDAFFE27C11" masterDocTitle="A new genus and species of myrmecophile clearwing moth (Lepidoptera: Sesiidae) from East Africa" masterLastPageNumber="594" masterPageNumber="588" pageNumber="589" updateTime="1698529746767" updateUser="ExternalLinkService">
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<mods:title id="2E4DFFB929C3B3BB2C722E8849DE2678">A new genus and species of myrmecophile clearwing moth (Lepidoptera: Sesiidae) from East Africa</mods:title>
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<mods:namePart id="30208D12D0697F41A7CCA5B58C658218">Agassiz, David</mods:namePart>
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<mods:namePart id="B75E20E04675E72A2C95AB45C605B972">Kallies, Axel</mods:namePart>
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<treatment id="03C06C57FA71FFE201C4FD15FBB67F7E" ID-DOI="http://doi.org/10.5281/zenodo.5959209" ID-GBIF-Taxon="142212293" ID-Zenodo-Dep="5959209" LSID="urn:lsid:plazi:treatment:03C06C57FA71FFE201C4FD15FBB67F7E" httpUri="http://treatment.plazi.org/id/03C06C57FA71FFE201C4FD15FBB67F7E" lastPageId="2" lastPageNumber="590" pageId="1" pageNumber="589">
<subSubSection id="C3738ECAFA71FFE101C4FD15FDF17EFB" box="[151,531,719,746]" pageId="1" pageNumber="589" type="nomenclature">
<paragraph id="8BD6DD41FA71FFE101C4FD15FDF17EFB" blockId="1.[151,531,719,746]" box="[151,531,719,746]" pageId="1" pageNumber="589">
<heading id="D09E6A2DFA71FFE101C4FD15FDF17EFB" bold="true" box="[151,531,719,746]" fontSize="11" level="1" pageId="1" pageNumber="589" reason="1">
<emphasis id="B91D0153FA71FFE101C4FD15FDF17EFB" bold="true" box="[151,531,719,746]" pageId="1" pageNumber="589">
<taxonomicName id="4C69A6C2FA71FFE101C4FD15FE7F7EFB" ID-CoL="B8YZ3" authority="Kallies" authorityName="Agassiz &amp; Kallies" authorityYear="2018" box="[151,413,719,746]" class="Insecta" family="Sesiidae" genus="Osmanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D0153FA71FFE101C4FD15FEA27EF8" bold="true" box="[151,320,719,745]" italics="true" pageId="1" pageNumber="589">Osmanthedon</emphasis>
Kallies
</taxonomicName>
<taxonomicNameLabel id="A22EBC28FA71FFE100F6FD0AFDF17EFB" box="[421,531,720,746]" pageId="1" pageNumber="589" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3738ECAFA71FFE101C4FCCFFCB57F38" box="[151,855,788,810]" pageId="1" pageNumber="589" type="materials_examined">
<paragraph id="8BD6DD41FA71FFE101C4FCCFFCB57F38" blockId="1.[151,855,788,810]" box="[151,855,788,810]" pageId="1" pageNumber="589">
Type species:
<taxonomicName id="4C69A6C2FA71FFE10075FCCEFD087F3B" authority="Agassiz &amp; Kallies" authorityName="Agassiz &amp; Kallies" authorityYear="2018" box="[294,746,788,810]" class="Insecta" family="Sesiidae" genus="Osmanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="domaticola" status="sp. nov.">
<emphasis id="B91D0153FA71FFE10075FCCEFDC97F3B" box="[294,555,788,810]" italics="true" pageId="1" pageNumber="589">Osmanthedon domaticola</emphasis>
Agassiz &amp; Kallies
</taxonomicName>
<taxonomicNameLabel id="A22EBC28FA71FFE103BCFCCEFCB57F38" box="[751,855,788,809]" pageId="1" pageNumber="589" rank="species">
<emphasis id="B91D0153FA71FFE103BCFCCEFCB57F38" bold="true" box="[751,855,788,809]" pageId="1" pageNumber="589">spec. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3738ECAFA71FFE101C4FC8EFE9B799B" pageId="1" pageNumber="589" type="description">
<paragraph id="8BD6DD41FA71FFE101C4FC8EFC547F7D" blockId="1.[151,1437,852,1418]" box="[151,950,852,877]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE101C4FC8EFECB7F7C" bold="true" box="[151,297,852,877]" pageId="1" pageNumber="589">Description.</emphasis>
Small moths, with a wingspan of
<quantity id="4C9170A4FA71FFE103F4FC8EFCD77F7D" box="[679,821,852,877]" metricMagnitude="-2" metricUnit="m" metricValue="1.725" metricValueMax="1.95" metricValueMin="1.5" pageId="1" pageNumber="589" unit="mm" value="17.25" valueMax="19.5" valueMin="15.0">1519.5 mm</quantity>
(
<figureCitation id="1352C1C4FA71FFE10210FC8EFC4A7F7D" box="[835,936,852,877]" captionStart="FIGURES16" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[203,1354,243,1932]" captionTargetId="figure@3.[198,1393,193,1960]" captionTargetPageId="3" captionText="FIGURES16. Osmanthedon domaticola gen. etspec.nov. 1.Male,holotype (alar expanse15 mm).2. Female,paratype (alar expanse 18.5 mm). 3. Venation. 4. Male genitalia, phallus removed. 5. Male genitalia, phallus.6. Female genitalia." httpUri="https://zenodo.org/record/1196273/files/figure.png" pageId="1" pageNumber="589">Figs 1, 2</figureCitation>
).
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194FCAFFBDB7FE3" blockId="1.[151,1437,852,1418]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE10194FCAFFEE67F9F" box="[199,260,885,910]" italics="true" pageId="1" pageNumber="589">Head</emphasis>
. Proboscis minute, perhaps not functional; labial palpus, basal and second palpomeres rough scaled, distal margin of second palpomere with bristly scales and extending beyond frons, apical palpomere smooth, 2/3 as long as second; frons smooth. Antenna thick, in particular in male, apically abruptly ending, not tapering, ventrally covered with short ciliae, which are slightly longer than the width of the flagellum.
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194FC26FAAC784D" blockId="1.[151,1437,852,1418]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE10194FC26FED17802" box="[199,307,1020,1043]" italics="true" pageId="1" pageNumber="589">Forewing</emphasis>
. Transparent areas densely covered in semitransparent scales. Venation (
<figureCitation id="1352C1C4FA71FFE10527FC20FB587802" box="[1140,1210,1018,1043]" captionStart="FIGURES16" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[203,1354,243,1932]" captionTargetId="figure@3.[198,1393,193,1960]" captionTargetPageId="3" captionText="FIGURES16. Osmanthedon domaticola gen. etspec.nov. 1.Male,holotype (alar expanse15 mm).2. Female,paratype (alar expanse 18.5 mm). 3. Venation. 4. Male genitalia, phallus removed. 5. Male genitalia, phallus.6. Female genitalia." httpUri="https://zenodo.org/record/1196273/files/figure.png" pageId="1" pageNumber="589">Fig. 3</figureCitation>
): forewing with M3 and Cu1 separate, R1 and R2 not fused, R4 and R5 with a short common stalk, hindwing veins M3, Cu1 and Cu2 closely approximated; M3 and Cu1 arise from a common point from the cross vein, without a common stalk.
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194FBBCFE7078B0" blockId="1.[151,1437,852,1418]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE10194FBBCFED3786E" box="[199,305,1126,1151]" italics="true" pageId="1" pageNumber="589">Abdomen</emphasis>
. Anal tuft small in male, in female only laterally developed, terminal segment dorsally and ventrally with very short scales.
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194FB73FEDF7956" blockId="1.[151,1437,852,1418]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE10194FB73FE8F78D3" box="[199,365,1193,1218]" italics="true" pageId="1" pageNumber="589">Male genitalia</emphasis>
(
<figureCitation id="1352C1C4FA71FFE1002EFB73FE0778D3" box="[381,485,1193,1218]" captionStart="FIGURES16" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[203,1354,243,1932]" captionTargetId="figure@3.[198,1393,193,1960]" captionTargetPageId="3" captionText="FIGURES16. Osmanthedon domaticola gen. etspec.nov. 1.Male,holotype (alar expanse15 mm).2. Female,paratype (alar expanse 18.5 mm). 3. Venation. 4. Male genitalia, phallus removed. 5. Male genitalia, phallus.6. Female genitalia." httpUri="https://zenodo.org/record/1196273/files/figure.png" pageId="1" pageNumber="589">Figs 4, 5</figureCitation>
). Typical of
<taxonomicName id="4C69A6C2FA71FFE1032BFB73FCCA78D3" authorityName="Niculescu" authorityYear="1964" box="[632,808,1193,1218]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
(setae bifurcate, scopula androconialis present, phallus without coecum penis); however, the vinculum and tegumen tightly fused, tegumen ventrally membranous, without distinct gnathos, valva with a soft bulbous crista near its centre; base of the valva with well-developed scale or sensilla sockets; ventral margin of the valve with a distinct processus half way; phallus with a distinctly hooked and large cornutus.
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194FA8AFE9B799B" blockId="1.[151,1437,852,1418]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE10194FA8AFE657978" box="[199,391,1360,1385]" italics="true" pageId="1" pageNumber="589">Female genitalia</emphasis>
(
<figureCitation id="1352C1C4FA71FFE100C5FA8AFE397978" box="[406,475,1360,1385]" captionStart="FIGURES16" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[203,1354,243,1932]" captionTargetId="figure@3.[198,1393,193,1960]" captionTargetPageId="3" captionText="FIGURES16. Osmanthedon domaticola gen. etspec.nov. 1.Male,holotype (alar expanse15 mm).2. Female,paratype (alar expanse 18.5 mm). 3. Venation. 4. Male genitalia, phallus removed. 5. Male genitalia, phallus.6. Female genitalia." httpUri="https://zenodo.org/record/1196273/files/figure.png" pageId="1" pageNumber="589">Fig. 6</figureCitation>
). Antrum very short, ductus seminalis close to ostium, ductus bursae close to antrum distinctly expanded.
</paragraph>
</subSubSection>
<subSubSection id="C3738ECAFA71FFE20194FA6EFBCC7F3C" lastPageId="2" lastPageNumber="590" pageId="1" pageNumber="589" type="diagnosis">
<paragraph id="8BD6DD41FA71FFE10194FA6EFB7579DD" blockId="1.[199,1175,1460,1485]" box="[199,1175,1460,1485]" pageId="1" pageNumber="589">
<emphasis id="B91D0153FA71FFE10194FA6EFEA379DC" bold="true" box="[199,321,1460,1485]" pageId="1" pageNumber="589">Diagnosis.</emphasis>
This genus is unique based on at least three autapomorphic character states:
</paragraph>
<paragraph id="8BD6DD41FA71FFE101C4FA2DFC897A44" blockId="1.[151,1436,1527,1654]" pageId="1" pageNumber="589">
(1) Hindwing veins M3, Cu1 and Cu2 closely approximated (well separated in all other
<taxonomicName id="4C69A6C2FA71FFE10535FA2DFAF97A01" authorityName="Niculescu" authorityYear="1964" box="[1126,1307,1527,1552]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
). (2) Valva with a soft bulbous crista near its centre (with a distinctly sclerotized ridge-like crista or a dense field of setae near the base of the valva in all other
<taxonomicName id="4C69A6C2FA71FFE103F9F9E6FCBD7A44" authorityName="Niculescu" authorityYear="1964" box="[682,863,1596,1621]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA71FFE101C4F984FAB07A67" blockId="1.[151,1436,1527,1654]" box="[151,1362,1629,1654]" pageId="1" pageNumber="589">
(3) Valva with sacculus with well-developed scale or sensilla sockets (without in all other
<taxonomicName id="4C69A6C2FA71FFE105C2F987FAA47A67" authorityName="Niculescu" authorityYear="1964" box="[1169,1350,1629,1654]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194F97BFBE07B2F" blockId="1.[151,1436,1696,1854]" pageId="1" pageNumber="589">
In addition to these unique characters,
<emphasis id="B91D0153FA71FFE103D9F97AFCC47AA8" box="[650,806,1696,1721]" italics="true" pageId="1" pageNumber="589">Osmanthedon</emphasis>
differs from most
<taxonomicName id="4C69A6C2FA71FFE10551F97AFB507AA8" authorityName="Niculescu" authorityYear="1964" box="[1026,1202,1696,1721]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
by the shape of the antennae, which are distinctly thickened and terminate abruptly in
<emphasis id="B91D0153FA71FFE102C5F91BFBD07ACB" box="[918,1074,1729,1754]" italics="true" pageId="1" pageNumber="589">Osmanthedon</emphasis>
, while they are thinner, clavate and tapering apically in most other
<taxonomicName id="4C69A6C2FA71FFE10374F939FD387AED" authorityName="Niculescu" authorityYear="1964" box="[551,730,1763,1788]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
, including
<taxonomicName id="4C69A6C2FA71FFE10204F939FB7A7AEA" authority="Hubner [1819]" authorityName="Hubner [" authorityYear="1819" box="[855,1176,1763,1788]" class="Insecta" family="Sesiidae" genus="Synanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA71FFE10204F939FC0A7AED" box="[855,1000,1763,1788]" italics="true" pageId="1" pageNumber="589">Synanthedon</emphasis>
<bibRefCitation id="EFF8A0B0FA71FFE102BCF939FB7A7AEA" box="[1007,1176,1763,1788]" pageId="1" pageNumber="589" refId="ref3981" refString="Hubner, J. ([1819]) Verzeichniss bekannter Schmettlinge. Verlag J. Hubner, Augsburg, pp. 2 - 11, 17 - 176." type="book chapter">Hübner [1819]</bibRefCitation>
</taxonomicName>
. However, some South American species currently assigned to
<taxonomicName id="4C69A6C2FA71FFE10307F8DEFC937B0C" authority="Butler, 1874" authorityName="Butler" authorityYear="1874" box="[596,881,1796,1821]" class="Insecta" family="Sesiidae" genus="Euryphrissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA71FFE10307F8DEFD3F7B0C" box="[596,733,1796,1821]" italics="true" pageId="1" pageNumber="589">Euryphrissa</emphasis>
<bibRefCitation id="EFF8A0B0FA71FFE103B0F8DEFC937B0C" box="[739,881,1796,1821]" pageId="1" pageNumber="589" refId="ref3500" refString="Butler, A. G. (1874) Notes on the Aegeriidae, with Descriptions of new Genera and Species. The Annals and Magazine of Natural History (Fourth Series), 14, 407 - 411. https: // doi. org / 10.1080 / 00222937408681004" type="journal article">Butler, 1874</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91D0153FA71FFE1022FF8DFFC0E7B0D" box="[892,1004,1797,1820]" italics="true" pageId="1" pageNumber="589">Carmenta</emphasis>
<bibRefCitation id="EFF8A0B0FA71FFE102A0F8DEFB797B0C" box="[1011,1179,1796,1821]" pageId="1" pageNumber="589" refId="ref3593" refString="Edwards, H. (1881) New Genera and Species of the Family Aegeridae. Papilio, 1 (10), 179 - 208, pl. 4." type="journal article">Edwards, 1881</bibRefCitation>
and
<taxonomicName id="4C69A6C2FA71FFE10582F8DFFF2D7B2F" authority="Le Cerf, 1917" authorityName="Le Cerf" authorityYear="1917" class="Insecta" family="Sesiidae" genus="Aegerina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA71FFE10582F8DFFADA7B0D" box="[1233,1336,1797,1820]" italics="true" pageId="1" pageNumber="589">Aegerina</emphasis>
<bibRefCitation id="EFF8A0B0FA71FFE1046CF8DFFF2D7B2F" pageId="1" pageNumber="589" refId="ref4118" refString="Le Cerf, F. (1917) Contributions a l'etude des Aegeriidae. Description et Iconographie d'Especes et de Formes nouvelles ou peu connues. In: Oberthur, C. (Ed.), Etudes de Lepidopterologie Comparee, 14. Impr. Oberthur, Rennes, pp. 137 - 388, pls 475 - 481." type="journal article">Le Cerf, 1917</bibRefCitation>
</taxonomicName>
have similar antennae (
<bibRefCitation id="EFF8A0B0FA71FFE1008AF8FFFC147B2F" box="[473,1014,1829,1854]" pageId="1" pageNumber="589" refId="ref3696" refString="Eichlin, T. D. (2003) Carmenta guayaba, a new clearwing moth from Peru (Lepidoptera: Sesiidae). Tropical Lepidoptera, 11 (1 - 2), 44 - 45." type="journal article">Eichlin 2003; Bartsch, personal communication</bibRefCitation>
).
</paragraph>
<paragraph id="8BD6DD41FA71FFE10194F8B1FB537B95" blockId="1.[199,1201,1899,1924]" box="[199,1201,1899,1924]" pageId="1" pageNumber="589">
Other characters that distinguish
<emphasis id="B91D0153FA71FFE1036AF8B1FD377B95" box="[569,725,1899,1924]" italics="true" pageId="1" pageNumber="589">Osmanthedon</emphasis>
from most
<taxonomicName id="4C69A6C2FA71FFE10205F8B1FBE47B95" authorityName="Niculescu" authorityYear="1964" box="[854,1030,1899,1924]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
are as follows:
</paragraph>
<paragraph id="8BD6DD41FA71FFE201C4F874FA957CA1" blockId="1.[151,1436,1965,2031]" lastBlockId="2.[151,1436,151,613]" lastPageId="2" lastPageNumber="590" pageId="1" pageNumber="589">
(1) Forewing veins R1 and R2 not fused (often fused distally, or occasionally completely fused in
<taxonomicName id="4C69A6C2FA71FFE10585F877FA697BD7" authorityName="Niculescu" authorityYear="1964" box="[1238,1419,1965,1990]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
). (2) Forewing veins R4 and R5 with a short common stalk (longer in typical
<taxonomicName id="4C69A6C2FA71FFE102A4F808FB4E7BFA" authorityName="Niculescu" authorityYear="1964" box="[1015,1196,2002,2027]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
). (3) Hindwing veins M3 and Cu1 arise from a common point from the cross vein (with a common stalk in most
<taxonomicName id="4C69A6C2FA72FFE205E4FF4DFA8E7CA1" authorityName="Niculescu" authorityYear="1964" box="[1207,1388,151,176]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA72FFE201C4FF66FE677CE7" blockId="2.[151,1436,151,613]" pageId="2" pageNumber="590">
(3) Hindwing veins M3 and Cu1 arise from a common point from the cross vein (with a common stalk in most
<taxonomicName id="4C69A6C2FA72FFE20197FF07FE9B7CE7" authorityName="Niculescu" authorityYear="1964" box="[196,377,221,246]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA72FFE201C4FF25FE277D8C" blockId="2.[151,1436,151,613]" pageId="2" pageNumber="590">
(4) Male genitalia with vinculum and tegumen tightly fused (connected by a thin membrane in most
<taxonomicName id="4C69A6C2FA72FFE20194FEFAFE997D28" authorityName="Niculescu" authorityYear="1964" box="[199,379,288,313]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), without any distinct gnathos lobes (well developed in most
<taxonomicName id="4C69A6C2FA72FFE2053DFEFAFAC07D28" authorityName="Niculescu" authorityYear="1964" box="[1134,1314,288,313]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), tegumen ventrally membranous (sclerotized in most
<taxonomicName id="4C69A6C2FA72FFE2039EFE9BFC607D4B" authorityName="Niculescu" authorityYear="1964" box="[717,898,321,346]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), ventral margin of the valva with a distinct processus (lacking in most
<taxonomicName id="4C69A6C2FA72FFE20353FEB8FD577D6A" authorityName="Niculescu" authorityYear="1964" box="[512,693,354,379]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
); phallus with a distinctly hooked and large cornutus (without in most
<taxonomicName id="4C69A6C2FA72FFE20057FE5EFE5B7D8C" authorityName="Niculescu" authorityYear="1964" box="[260,441,388,413]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA72FFE201C4FE7FFD3B7E10" blockId="2.[151,1436,151,613]" pageId="2" pageNumber="590">
(5) Female genitalia with antrum very short (long and well-developed in most
<taxonomicName id="4C69A6C2FA72FFE2054AFE7FFB2F7DAF" authorityName="Niculescu" authorityYear="1964" box="[1049,1229,421,446]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), ductus seminalis close to ostium (well removed in most
<taxonomicName id="4C69A6C2FA72FFE2032FFE1CFCD27DCE" authorityName="Niculescu" authorityYear="1964" box="[636,816,454,479]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), ductus bursae distinctly expanded in the section close to the antrum (simple in most
<taxonomicName id="4C69A6C2FA72FFE2034BFE32FD2F7E10" authorityName="Niculescu" authorityYear="1964" box="[536,717,488,513]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA72FFE201C4FDD3FE6A7E74" blockId="2.[151,1436,151,613]" pageId="2" pageNumber="590">
(6) The anal tuft in the female is only laterally developed, terminal segment dorsally and ventrally with very short scales (anal tuft in females very prominent, terminal segment dorsally and ventrally with long scales in most
<taxonomicName id="4C69A6C2FA72FFE20194FD96FE9E7E74" authorityName="Niculescu" authorityYear="1964" box="[199,380,588,613]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
).
</paragraph>
<paragraph id="8BD6DD41FA72FFE20194FD54FBCC7F3C" blockId="2.[151,1437,654,880]" pageId="2" pageNumber="590">
The new genus shares some characters with
<taxonomicName id="4C69A6C2FA72FFE203ECFD55FC067EB6" authority="Strand, 1914" authorityName="Strand" authorityYear="1914" box="[703,996,654,679]" class="Insecta" family="Sesiidae" genus="Camaegeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE203ECFD55FCA87EB7" box="[703,842,655,678]" italics="true" pageId="2" pageNumber="590">Camaegeria</emphasis>
Strand, 1914
</taxonomicName>
, which also lacks the sclerotized crista sacculi and has an undeveloped anal tuft in the female. However, the two genera are not similar otherwise (comp.
<bibRefCitation id="EFF8A0B0FA72FFE201C4FD0BFE6E7EFB" box="[151,396,721,746]" pageId="2" pageNumber="590" refId="ref3420" refString="Bartsch, D. &amp; Berg, J. (2012) New species and review of the Afrotropical clearwing moth genus Camaegeria Strand, 1914 (Lepidoptera: Sesiidae: Synanthedonini). Zootaxa, 3181, 28 - 46." type="journal article">Bartsch &amp; Berg 2012</bibRefCitation>
). Superficially, the new genus is also somewhat similar to
<taxonomicName id="4C69A6C2FA72FFE20573FD08FB9D7EF8" authorityName="Le Cerf" authorityYear="1917" box="[1056,1151,722,745]" class="Insecta" family="Sesiidae" genus="Osminia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE20573FD08FB9D7EF8" box="[1056,1151,722,745]" italics="true" pageId="2" pageNumber="590">Osminia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69A6C2FA72FFE205E4FD0BFF317F1A" authority="Bartsch, 2015" authorityName="Bartsch" authorityYear="2015" class="Insecta" family="Sesiidae" genus="Coccophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE205E4FD0BFADA7EFB" box="[1207,1336,721,746]" italics="true" pageId="2" pageNumber="590">Coccophila</emphasis>
<bibRefCitation id="EFF8A0B0FA72FFE2046CFD0BFF317F1A" pageId="2" pageNumber="590" refId="ref3327" refString="Bartsch, D. (2015) New taxa of southern African Sesiini (Lepidoptera: Sesiidae). Zootaxa, 3956 (3), 428 - 436. https: // doi. org / 10.11646 / zootaxa. 3956.3.7" type="journal article">Bartsch, 2015</bibRefCitation>
</taxonomicName>
. These genera, however, belong to the tribes
<taxonomicName id="4C69A6C2FA72FFE2039EFD28FCDF7F1A" authorityName="Duckworth &amp; Eichlin" authorityYear="1977" box="[717,829,754,779]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Osminiini">Osminiini</taxonomicName>
and
<taxonomicName id="4C69A6C2FA72FFE20227FD28FC217F1A" authorityName="Boisduval" authorityYear="1828" box="[884,963,754,779]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="tribe" tribe="Sesiini">Sesiini</taxonomicName>
, respectively, and differ fundamentally by the genitalia and wing venation (
<bibRefCitation id="EFF8A0B0FA72FFE20356FCCEFD787F3D" box="[517,666,788,813]" pageId="2" pageNumber="590" refId="ref3659" refString="Eichlin, T. D. (1998) Western Hemisphere Clearwing Moths of the Tribe Osminiini (Lepidoptera: Sesiidae: Sesiinae). Holarctic Lepidoptera, 5 (1), 23 - 33." type="journal article">Eichlin 1998</bibRefCitation>
;
<bibRefCitation id="EFF8A0B0FA72FFE203F6FCCEFCD57F3D" box="[677,823,788,813]" pageId="2" pageNumber="590" refId="ref4011" refString="Kallies, A. (2004) Sesiidae (Lepidoptera, Sesioidea). Esperiana Memoir, 1, 185 - 188." type="journal article">Kallies 2004</bibRefCitation>
;
<bibRefCitation id="EFF8A0B0FA72FFE20211FCCEFC387F3C" box="[834,986,788,813]" pageId="2" pageNumber="590" refId="ref3327" refString="Bartsch, D. (2015) New taxa of southern African Sesiini (Lepidoptera: Sesiidae). Zootaxa, 3956 (3), 428 - 436. https: // doi. org / 10.11646 / zootaxa. 3956.3.7" type="journal article">Bartsch 2015</bibRefCitation>
,
<bibRefCitation id="EFF8A0B0FA72FFE202B6FCCEFBC07F3C" box="[997,1058,788,813]" pageId="2" pageNumber="590" refId="ref3369" refString="Bartsch, D. (2016) Revisionary checklist of the southern African Osminiini (Lepidoptera: Sesiidae). Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 9, 229 - 265. https: // doi. org / 10.18476 / sbna. v 9. a 15" type="journal article">2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3738ECAFA72FFE20194FCEFFBB67F7E" pageId="2" pageNumber="590" type="etymology">
<paragraph id="8BD6DD41FA72FFE20194FCEFFBB67F7E" blockId="2.[151,1437,654,880]" pageId="2" pageNumber="590">
<emphasis id="B91D0153FA72FFE20194FCEFFEAC7F5F" bold="true" box="[199,334,821,846]" pageId="2" pageNumber="590">Etymology.</emphasis>
The name derives from the generic names
<taxonomicName id="4C69A6C2FA72FFE20263FCECFC6D7F5C" authorityName="Le Cerf" authorityYear="1917" box="[816,911,822,845]" class="Insecta" family="Sesiidae" genus="Osminia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE20263FCECFC6D7F5C" box="[816,911,822,845]" italics="true" pageId="2" pageNumber="590">Osminia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69A6C2FA72FFE20296FCEFFBB47F5F" authorityName="Hubner [" authorityYear="1819" box="[965,1110,821,846]" class="Insecta" family="Sesiidae" genus="Synanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE20296FCEFFBB47F5F" box="[965,1110,821,846]" italics="true" pageId="2" pageNumber="590">Synanthedon</emphasis>
</taxonomicName>
, referring to the similarity of the new genus to species of
<taxonomicName id="4C69A6C2FA72FFE20082FC8DFDD27F7F" authorityName="Le Cerf" authorityYear="1917" box="[465,560,855,878]" class="Insecta" family="Sesiidae" genus="Osminia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE20082FC8DFDD27F7F" box="[465,560,855,878]" italics="true" pageId="2" pageNumber="590">Osminia</emphasis>
</taxonomicName>
while indicating its relationship to
<taxonomicName id="4C69A6C2FA72FFE202EFFC8CFBAF7F7E" authorityName="Hubner [" authorityYear="1819" box="[956,1101,854,879]" class="Insecta" family="Sesiidae" genus="Synanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B91D0153FA72FFE202EFFC8CFBAF7F7E" box="[956,1101,854,879]" italics="true" pageId="2" pageNumber="590">Synanthedon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>