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<document id= "48E11B378413A0E77773F43E3676C06D" ID-CLB-Dataset= "39813" ID-DOI= "10.11646/zootaxa.3990.2.2" ID-GBIF-Dataset= "1740ec81-c55e-44ce-9a6d-cd62acd5d898" ID-ISSN= "1175-5326" ID-Zenodo-Dep= "242947" ID-ZooBank= "BDF1BC11-4848-4858-99DC-0336A5FBEE2B" IM.materialsCitations_approvedBy= "felipe" IM.metadata_approvedBy= "felipe" IM.tables_requiresApprovalFor= "existingObjects,plazi" IM.taxonomicNames_approvedBy= "felipe" checkinTime= "1461267666231" checkinUser= "plazi" docAuthor= "Helwerda, Renate A." docDate= "2015" docId= "03FA878C9501FE5BFF26E1DE02A6CCC2" docLanguage= "en" docName= "zt03990p220.pdf" docOrigin= "Zootaxa 3990 (2)" docStyle= "DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId= "647186512141C8FC8976D5BCC54AEB7D" docStyleName= "Zootaxa.2013-.journal_article" docStyleVersion= "9" docTitle= "Ringicula" docType= "treatment" docVersion= "11" lastPageNumber= "214" masterDocId= "FFC3FFF49510FE4AFFB1E574036DCA1D" masterDocTitle= "Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines" masterLastPageNumber= "220" masterPageNumber= "197" pageNumber= "214" updateTime= "1732492386731" updateUser= "ExternalLinkService" >
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<mods:title id= "88920594D364577AB82366803FC2DF62" > Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines</mods:title>
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<mods:name id= "7FBCBDA611F6E750BF55963CD5700044" type= "personal" >
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<mods:namePart id= "EFD943B7A8BBD3DCE24491FAE139B1BF" > Helwerda, Renate A.</mods:namePart>
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<mods:title id= "1376801BFC8CD4A6219A206971D609AA" > Zootaxa</mods:title>
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<mods:part id= "E22EF8BA0FBB7216894A2BA95AE60834" >
<mods:date id= "9D1E786B8CF2C50F54E5B0C196D25C1B" > 2015</mods:date>
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<mods:number id= "E4A28E1723D6CFE100E32DFC12069E30" > 3990</mods:number>
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<mods:detail id= "C41D7E5C408FA81E31A7F974CCA9AE48" type= "issue" >
<mods:number id= "75C872B02D6921FF2E4E3772DE96E57C" > 2</mods:number>
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<treatment id= "03FA878C9501FE5BFF26E1DE02A6CCC2" ID-DOI= "http://doi.org/10.5281/zenodo.6116545" ID-GBIF-Taxon= "119638250" ID-Zenodo-Dep= "6116545" LSID= "urn:lsid:plazi:treatment:03FA878C9501FE5BFF26E1DE02A6CCC2" httpUri= "http://treatment.plazi.org/id/03FA878C9501FE5BFF26E1DE02A6CCC2" lastPageNumber= "214" pageId= "17" pageNumber= "214" scope_class= "Gastropoda" scope_family= "Ringiculidae" scope_subClass= "Heterobranchia" >
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<subSubSection id= "C34965119501FE5BFF26E1DE0278CEFA" pageId= "17" pageNumber= "214" type= "nomenclature" >
<paragraph id= "8BEC369A9501FE5BFF26E1DE0235CEDE" blockId= "17.[151,344,1194,1255]" box= "[151,344,1194,1219]" pageId= "17" pageNumber= "214" >
<heading id= "D0A481F69501FE5BFF26E1DE0235CEDE" bold= "true" box= "[151,344,1194,1219]" fontSize= "10" level= "2" pageId= "17" pageNumber= "214" reason= "2" >
<emphasis id= "B927EA889501FE5BFF26E1DE0235CEDE" bold= "true" box= "[151,344,1194,1219]" pageId= "17" pageNumber= "214" >
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<taxonomicName id= "4C534D199501FE5BFF26E1DE0264CEDE" ID-CoL= "7PFTM" box= "[151,265,1194,1219]" class= "Gastropoda" family= "Ringiculidae" genus= "Ringicula" kingdom= "Animalia" pageId= "17" pageNumber= "214" phylum= "Mollusca" rank= "genus" >
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<emphasis id= "B927EA889501FE5BFF26E1DE0264CEDE" bold= "true" box= "[151,265,1194,1219]" italics= "true" pageId= "17" pageNumber= "214" > Ringicula</emphasis>
</taxonomicName>
spec 3
</emphasis>
</heading>
</paragraph>
<paragraph id= "8BEC369A9501FE5BFF26E1BA0278CEFA" blockId= "17.[151,344,1194,1255]" box= "[151,277,1230,1255]" pageId= "17" pageNumber= "214" >
(
<figureCitation id= "13682A1F9501FE5BFF2FE1BA0260CEFA" box= "[158,269,1230,1255]" captionStart= "FIGURE 29 – captionStartId= "16.[151,250,1798,1820]" captionTargetBox= "[234,1351,195,1775]" captionTargetId= "figure@16.[234,1352,195,1776]" captionTargetPageId= "16" captionText= "FIGURE 29 – httpUri= "https://zenodo.org/record/242953/files/figure.png" pageId= "17" pageNumber= "214" > Figure 33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id= "C34965119501FE5BFF26E0620198CFA2" pageId= "17" pageNumber= "214" type= "materials_examined" >
<paragraph id= "8BEC369A9501FE5BFF26E062014BCF32" blockId= "17.[151,1436,1302,1759]" box= "[151,550,1302,1327]" pageId= "17" pageNumber= "214" >
<emphasis id= "B927EA889501FE5BFF26E062026BCF32" bold= "true" box= "[151,262,1302,1327]" pageId= "17" pageNumber= "214" > Material.</emphasis>
Tiep2 (1)
<collectionCode id= "ED42AE5F9501FE5BFECDE06302D1CF32" box= "[380,444,1303,1327]" httpUri= "http://grbio.org/cool/47nz-wbm0" name= "National Museum of Natural History, Naturalis" pageId= "17" pageNumber= "214" > RGM</collectionCode>
961.834.
</paragraph>
<paragraph id= "8BEC369A9501FE5BFF76E04E0198CFA2" blockId= "17.[151,1436,1302,1759]" pageId= "17" pageNumber= "214" >
<emphasis id= "B927EA889501FE5BFF76E04E02F7CF4E" bold= "true" box= "[199,410,1338,1363]" pageId= "17" pageNumber= "214" > Characterization.</emphasis>
Shell small; H
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,
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W
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mm; protoconch slightly elevated, 1.2 whorl, DP
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; nucleus small, DN
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; teleoconch with distinct subsutural ridge and groove; additional sculpture of 16 spiral grooves on the body whorl; outer lip very thick with swollen lower half; parietal callus with lamella; columella with two folds; lower fold bears two knobs.
</paragraph>
</subSubSection>
<subSubSection id= "C34965119501FE5BFF76E0BE02A6CCC2" pageId= "17" pageNumber= "214" type= "discussion" >
<paragraph id= "8BEC369A9501FE5BFF76E0BE02A6CCC2" blockId= "17.[151,1436,1302,1759]" pageId= "17" pageNumber= "214" >
<emphasis id= "B927EA889501FE5BFF76E0BE0256CFFE" bold= "true" box= "[199,315,1482,1507]" pageId= "17" pageNumber= "214" > Remarks.</emphasis>
This specimen is similar to
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<taxonomicName id= "4C534D199501FE5BFD38E0BF002BCFFF" box= "[649,838,1483,1507]" class= "Gastropoda" family= "Ringiculidae" genus= "Ringicula" kingdom= "Animalia" pageId= "17" pageNumber= "214" phylum= "Mollusca" rank= "species" species= "opima" >
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<emphasis id= "B927EA889501FE5BFD38E0BF002BCFFF" box= "[649,838,1483,1507]" italics= "true" pageId= "17" pageNumber= "214" > Ringicula opima</emphasis>
</taxonomicName>
nov. spec., but its spire is smaller and it has fewer whorls. The thought arose that this specimen is therefore a deviant specimen of this species, with a teratologically small size. However, its apertural callus is well developed, but it does not extend as far onto the base as in
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<taxonomicName id= "4C534D199501FE5BFF26E342023ECC53" box= "[151,339,1590,1614]" class= "Gastropoda" family= "Ringiculidae" genus= "Ringicula" kingdom= "Animalia" pageId= "17" pageNumber= "214" phylum= "Mollusca" rank= "species" species= "opima" >
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<emphasis id= "B927EA889501FE5BFF26E342023ECC53" box= "[151,339,1590,1614]" italics= "true" pageId= "17" pageNumber= "214" > Ringicula opima</emphasis>
</taxonomicName>
nov. spec.. It also bears two knobs on the lower fold of the columella and its outer lip is only slightly projected beyond the columella. The rest of the apertural characters were obscured by sediment, but they were revealed by making a 3D reconstruction of a CT-scan of this shell. This shows that the columellar folds are also shaped differently. All in all, this specimen differs from
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<taxonomicName id= "4C534D199501FE5BFC8DE3D7009BCCA7" box= "[828,1014,1699,1723]" class= "Gastropoda" family= "Ringiculidae" genus= "Ringicula" kingdom= "Animalia" pageId= "17" pageNumber= "214" phylum= "Mollusca" rank= "species" species= "opima" >
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<emphasis id= "B927EA889501FE5BFC8DE3D7009BCCA7" box= "[828,1014,1699,1723]" italics= "true" pageId= "17" pageNumber= "214" > Ringicula opima</emphasis>
</taxonomicName>
nov. spec. in so many characters, that it is treated separately here.
</paragraph>
</subSubSection>
</treatment>
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