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<document id="800244874B006956E738FE0CF3E42B9D" ID-DOI="10.1111/j.1096-3642.2012.00863.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5409181" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630350074602" checkinUser="marcus" docAuthor="Criscione, Francesco, Law, Margot Louisa & Köhler, Frank" docDate="2012" docId="03E9878DFFDCFF9471AEA6345BA5F97A" docLanguage="en" docName="zoj863.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00863.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Exiligada Iredale 1939" docType="treatment" docVersion="4" lastPageNumber="696" masterDocId="FFD0FFF5FFDAFF93710AA04F5F3FFFCD" masterDocTitle="Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species" masterLastPageNumber="722" masterPageNumber="689" pageNumber="695" updateTime="1699241497941" updateUser="plazi">
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<mods:title id="E9275919F304E57DB4CA112C48514778">Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species</mods:title>
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<mods:namePart id="A96B77695B0EA92C1A2FB8A2E06AEEA0">Criscione, Francesco</mods:namePart>
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<mods:namePart id="2E4C2398E325C552EFDFFDC353F9657B">Law, Margot Louisa</mods:namePart>
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<mods:namePart id="0C14F9F540BA8B77E66210E988969A32">Köhler, Frank</mods:namePart>
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<mods:title id="C8C01C1BE7C45D4C6AEDE9C72981270A">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="032F859137B53F5AEE45263FD2FC9D8E">2012</mods:date>
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<mods:title id="00C62387B24C31D9164889A1F652C028">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="84B7B2C826E6DD3B60C9ACA5B0D8B1EE">2012-11-26</mods:number>
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<mods:number id="14D56935B901F7A234AFE28852AC4E61">166</mods:number>
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<mods:number id="1179FBACE8B08B06D291884C31C028C0">4</mods:number>
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<mods:start id="E099223ADCCD53D3E6B6E38E96EE3701">689</mods:start>
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<mods:identifier id="2A030CC020C9C6BFAFD0205FBEC4D461" type="DOI">10.1111/j.1096-3642.2012.00863.x</mods:identifier>
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<treatment id="03E9878DFFDCFF9471AEA6345BA5F97A" LSID="urn:lsid:plazi:treatment:03E9878DFFDCFF9471AEA6345BA5F97A" httpUri="http://treatment.plazi.org/id/03E9878DFFDCFF9471AEA6345BA5F97A" lastPageId="7" lastPageNumber="696" pageId="6" pageNumber="695">
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<subSubSection id="C35A6510FFDCFF9571AEA6345DDDF95C" box="[164,738,1659,1681]" pageId="6" pageNumber="695" type="nomenclature">
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<paragraph id="8BFF369BFFDCFF9571AEA6345DDDF95C" blockId="6.[164,738,1659,1681]" box="[164,738,1659,1681]" pageId="6" pageNumber="695">
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<heading id="D0B781F7FFDCFF9571AEA6345DDDF95C" box="[164,738,1659,1681]" centered="true" fontSize="8" level="2" pageId="6" pageNumber="695" reason="8">
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<treatmentCitationGroup id="AB5011B5FFDCFF9571AEA6345DDDF95C" box="[164,738,1659,1681]" pageId="6" pageNumber="695">
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<treatmentCitation id="0AE1108AFFDCFF9571AEA6345D3EF95C" author="Iredale T" box="[164,513,1659,1681]" page="68" pageId="6" pageNumber="695" year="1939">
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<taxonomicName id="4C404D18FFDCFF9571AEA6345D3EF95C" ID-CoL="7P3TH" authority="IREDALE, 1939: 68" authorityName="Iredale" authorityPageNumber="68" authorityYear="1939" box="[164,513,1659,1681]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="695" phylum="Mollusca" rank="genus">
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<emphasis id="B934EA89FFDCFF9571AEA6345E22F942" box="[164,285,1659,1680]" italics="true" pageId="6" pageNumber="695">EXILIGADA</emphasis>
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<bibRefCitation id="EFD14B6AFFDCFF95702DA6345D3EF95C" author="Iredale T" box="[295,513,1659,1681]" pageId="6" pageNumber="695" pagination="1 - 88" refId="ref21430" refString="Iredale T. 1939. A review of the land Mollusca of Western Australia. Records of the Western Australian Museum and Art Gallery 2: 1 - 88." type="journal article" year="1939">IREDALE, 1939: 68</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="0AE1108AFFDCFF95731AA6345DDDF95C" author="Solem A" box="[528,738,1659,1681]" page="670" pageId="6" pageNumber="695" year="1984">
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<bibRefCitation id="EFD14B6AFFDCFF95731AA6345DDDF95C" author="Solem A" box="[528,738,1659,1681]" pageId="6" pageNumber="695" pagination="427 - 705" refId="ref22502" refString="Solem A. 1984. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). IV. Taxa from the Kimberley, Westraltrachia Iredale, 1933 and related genera. Records of the Western Australian Museum Supplement 17: 427 - 705." type="journal article" year="1984">SOLEM, 1984: 670</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A6510FFDCFF94719AA6ED5BA5F97A" lastPageId="7" lastPageNumber="696" pageId="6" pageNumber="695" type="diagnosis">
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<paragraph id="8BFF369BFFDCFF95719AA6ED5CFCF9CD" blockId="6.[144,759,1697,1903]" lastBlockId="6.[806,1422,1269,1903]" pageId="6" pageNumber="695">
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<emphasis id="B934EA89FFDCFF95719AA6ED5E5EF97B" box="[144,353,1697,1719]" italics="true" pageId="6" pageNumber="695">Generic diagnosis:</emphasis>
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Shell (
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<figureCitation id="137B2A1EFFDCFF9570BEA6EE5D26F97A" box="[436,537,1697,1719]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="8.[144,223,954,973]" captionStartId-1="9.[164,243,1300,1319]" captionStartId-2="9.[826,905,1233,1252]" captionTargetBox-0="[144,1425,196,924]" captionTargetBox-1="[171,774,199,1267]" captionTargetBox-2="[834,1435,197,1200]" captionTargetId-0="figure-472@8.[144,1425,196,924]" captionTargetId-1="figure-225@9.[168,777,197,1270]" captionTargetId-2="figure-294@9.[830,1439,194,1203]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="9" captionText-0="Figure 4. Photographs of living specimens of Exiligada species (courtesy Vince Kessner, Adelaide River). A, E. negriensis Iredale, 1939. B, E. pallida sp. nov. C, E. nodulicaudata sp. nov. D, E. punctata sp. nov. E, E. qualis Iredale, 1939. F, E. monochroma sp. nov. G, E. unistriata sp. nov. H, E. floraevallis sp. nov. I, E. rivifontis sp. nov." captionText-1="Figure 5. Shells of Exiligada species. A, E. negriensis (holotype AM C.64865). B, E. brabyi sp. nov. (paratype C.443279). C, E. calciphila sp. nov. (paratype C.462723). D, E. floraevallis sp. nov. (paratype WAM S49204). E, E. gregoriana sp. nov. (paratype AM C.462820). F, E. limbunya sp. nov. (paratype AM C.470205). G, E. longicauda sp. nov. (paratype WAM S49228). H, E. monochroma sp. nov. (paratype AM C.469772)." captionText-2="Figure 6. Shells of Exiligada species. A, E. montejinni sp. nov. (paratype AM C.469982). B, E. nodulicauda sp. nov. (paratype WAM S49160). C, E. pallida sp. nov. (paratype WAM S49215). D, E. punctata sp. nov. (paratype WAM S49221). E, E. qualis Iredale, 1939 (holotype AM C.64866). F, E. rivifontis sp. nov. (WAM S49225). G, E. unistriata sp. nov. (paratype AM C.470460). Scale bar = 2 mm. Note that foot produces from some shells." figureDoi-0="http://doi.org/10.5281/zenodo.5409193" figureDoi-1="http://doi.org/10.5281/zenodo.5409197" figureDoi-2="http://doi.org/10.5281/zenodo.5409195" httpUri-0="https://zenodo.org/record/5409193/files/figure.png" httpUri-1="https://zenodo.org/record/5409197/files/figure.png" httpUri-2="https://zenodo.org/record/5409195/files/figure.png" pageId="6" pageNumber="695">Figs 4–6</figureCitation>
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): moderately large, subglobose, periphery slightly angulated to rounded, spire weakly to moderately elevated, thin to moderately thick; umbilicus narrowly winding, open to almost completely concealed by columellar reflection; protoconch and teleoconch with moderately fine radial growth lines, fine micropustulations or microdepressions sometimes present. Last whorl moderate to wide in cross-section, periphery uniformly rounded. Apertural lip reflected, moderately expanded, parietal wall thin; no internal lip nodes. Shell colour variable, background whitish, yellowish or brownish horn, without or with one to several moderately wide to thin brown-reddish spiral bands, sometimes randomly to regularly dashed, with or without dark grey punctuations.
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</paragraph>
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<paragraph id="8BFF369BFFDCFF947235A6475D42F9B7" blockId="6.[806,1422,1269,1903]" lastBlockId="7.[164,779,932,1658]" lastPageId="7" lastPageNumber="696" pageId="6" pageNumber="695">Genitalia: typically camaenid with prostate and uterus being fused forming spermoviduct and lacking diverticulum and stimulatory organs; development of gonads depending on seasonal activity and maturity. Penial sheath well developed, extending entire length, usually thickened distally. Epiphallus forming a tube surrounded by thick layers of muscle, well separated from penis, distally connected to penis through a narrow duct in a zigzag configuration; epiphallus variable in length, three times shorter to two times longer than penis. Penial retractor muscle inserting on vas-epiphallic junction, with fibres continuing along epiphallus, and attached to apical part of penis. Vas deferens entering penial sheath basally to almost halfway up, continuing apically within sheath until reflexing into epiphallus. Penis ten to three times longer than wide, not coiled or coiled inside sheath, with moderately thick wall; inner penial wall structure variable, usually consisting of several irregularly shaped, often corrugated, wide longitudinal pilasters or of fewer regularly shaped, smooth, narrower ones. Pilaster often running along entire wall length, but sometimes interrupted. In some species finer sculptures, such as irregular tubercles or regular nodules, can be present. Sculptures of posterior part of wall often different from that of anterior part. Vagina four times shorter to slightly longer than penis, usually three times shorter to as long as free oviduct, but up to more than two times longer in some species; bursa copulatrix relatively simple, usually as long as free oviduct and sometimes slightly shorter, reaching anterior end of spermoviduct, with or without welldifferentiated end. Albumen gland large. Talon embedded in albumen gland within proximal portion. Hermaphroditic duct tightly undulating.</paragraph>
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<caption id="DF3F6613FFDDFF9471AEA3685B38FCB8" ID-DOI="http://doi.org/10.5281/zenodo.5409191" ID-Zenodo-Dep="5409191" httpUri="https://zenodo.org/record/5409191/files/figure.png" pageId="7" pageNumber="696" startId="7.[164,243,807,826]" targetBox="[166,1441,198,774]" targetPageId="7">
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<paragraph id="8BFF369BFFDDFF9471AEA3685B38FCB8" blockId="7.[164,1441,807,886]" pageId="7" pageNumber="696">
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<emphasis id="B934EA89FFDDFF9471AEA3685E33FCF6" bold="true" box="[164,268,807,827]" pageId="7" pageNumber="696">Figure 3.</emphasis>
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Diagrammatic representation of pair–wise comparisons of shell parameters in
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<taxonomicName id="4C404D18FFDDFF94756AA3675A26FCF6" box="[1120,1305,808,827]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B934EA89FFDDFF94756AA3675BF9FCF6" box="[1120,1222,808,827]" italics="true" pageId="7" pageNumber="696">Exiligada</emphasis>
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species
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</taxonomicName>
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by means of pair-wise one–way ANOVA (H, left; D, right). Full squares: null hypothesis that group means are equal rejected with statistical significance (P
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<emphasis id="B934EA89FFDDFF9470BFA32C5EFEFCB8" box="[437,449,867,885]" italics="true" pageId="7" pageNumber="696"><</emphasis>
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0.05). Crosses: null hypothesis not rejected (P
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<emphasis id="B934EA89FFDDFF9472B0A32C5CF9FCB8" box="[954,966,867,885]" italics="true" pageId="7" pageNumber="696">></emphasis>
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0.05).
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</paragraph>
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</caption>
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<paragraph id="8BFF369BFFDDFF9471AEA6FE5DE0F90B" blockId="7.[164,735,1712,1734]" box="[164,735,1712,1734]" pageId="7" pageNumber="696">
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<emphasis id="B934EA89FFDDFF9471AEA6FE5EB1F90B" box="[164,398,1713,1734]" italics="true" pageId="7" pageNumber="696">Aestivating strategy:</emphasis>
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Aestivating as ‘free sealers’.
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</paragraph>
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<paragraph id="8BFF369BFFDDFF9471AEA6B25DE9F8A2" blockId="7.[164,779,1789,1903]" pageId="7" pageNumber="696">
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<emphasis id="B934EA89FFDDFF9471AEA6B25E30F8DF" box="[164,271,1789,1810]" italics="true" pageId="7" pageNumber="696">Remarks:</emphasis>
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If not stated otherwise,
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<typeStatus id="54FB8839FFDDFF94736CA6B25DEDF8DE" box="[614,722,1789,1811]" pageId="7" pageNumber="696" type="holotype">holotypes</typeStatus>
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are dissected wet specimens, whose shell was cracked to permit access to soft tissues. References to size are intended as relative to other congeneric species.
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</paragraph>
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<paragraph id="8BFF369BFFDDFF947259A3EB5BA5F97A" blockId="7.[826,1441,932,1719]" pageId="7" pageNumber="696">
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Typical combination of features separating
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<taxonomicName id="4C404D18FFDDFF947468A3EB5C4CFC1A" authorityName="Iredale" authorityYear="1939" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
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<emphasis id="B934EA89FFDDFF947468A3EB5C4CFC1A" italics="true" pageId="7" pageNumber="696">Exiligada</emphasis>
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</taxonomicName>
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from other camaenids are shell shape of the ‘chloritid type’, presence of dashed shell bands (where present), and in particular shape and zigzag configuration of epiphallus. Amongst other camaenid genera occurring in EK and NT,
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<taxonomicName id="4C404D18FFDDFF947588A4725A07FB9F" authorityName="Solem" authorityYear="1985" box="[1154,1336,1085,1106]" class="Gastropoda" family="Camaenidae" genus="Mesodontrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
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<emphasis id="B934EA89FFDDFF947588A4725A07FB9F" box="[1154,1336,1085,1106]" italics="true" pageId="7" pageNumber="696">Mesodontrachia</emphasis>
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</taxonomicName>
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is most similar in shell shape and size but differs by slightly lower spire and by whorls being never angulated as in some but not all
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<taxonomicName id="4C404D18FFDDFF9472FDA4D65BFFFB62" box="[1015,1216,1177,1199]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B934EA89FFDDFF9472FDA4D65B59FB63" box="[1015,1126,1177,1198]" italics="true" pageId="7" pageNumber="696">Exiligada</emphasis>
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species
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</taxonomicName>
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; shell very different from all other genera by larger size and much higher spire, narrower umbilicus with greater extent of cover by columellar reflection; protoconch microsculpture similar to
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<taxonomicName id="4C404D18FFDDFF9472B9A55C5B7DFAE5" authorityName="Solem" authorityYear="1984" box="[947,1090,1299,1320]" class="Gastropoda" family="Camaenidae" genus="Prototrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
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<emphasis id="B934EA89FFDDFF9472B9A55C5B7DFAE5" box="[947,1090,1299,1320]" italics="true" pageId="7" pageNumber="696">Prototrachia</emphasis>
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</taxonomicName>
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and some
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<taxonomicName id="4C404D18FFDDFF9475C8A55C5AA2FAE4" box="[1218,1437,1299,1321]" class="Gastropoda" family="Camaenidae" genus="Ordtrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="species" species="undetermined">
|
|||
|
<emphasis id="B934EA89FFDDFF9475C8A55C5A7EFAE5" box="[1218,1345,1299,1320]" italics="true" pageId="7" pageNumber="696">Ordtrachia</emphasis>
|
|||
|
species
|
|||
|
</taxonomicName>
|
|||
|
; teleoconch sculpture similar to
|
|||
|
<taxonomicName id="4C404D18FFDDFF9475B2A57D5A51FA8A" authorityName="Solem" authorityYear="1985" box="[1208,1390,1330,1351]" class="Gastropoda" family="Camaenidae" genus="Mesodontrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF9475B2A57D5A51FA8A" box="[1208,1390,1330,1351]" italics="true" pageId="7" pageNumber="696">Mesodontrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
but differing from remaining genera by absence of radial ribs and spiral keels; shell colour usually lighter; colour pattern very distinct from all other genera by presence of spiral bands, but species with no banding most similar to
|
|||
|
<taxonomicName id="4C404D18FFDDFF947501A5845BFEFA2D" authorityName="Solem" authorityYear="1985" box="[1035,1217,1483,1504]" class="Gastropoda" family="Camaenidae" genus="Mesodontrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF947501A5845BFEFA2D" box="[1035,1217,1483,1504]" italics="true" pageId="7" pageNumber="696">Mesodontrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
.
|
|||
|
<collectingRegion id="4984F879FFDDFF9475D2A5845A22FA2D" box="[1240,1309,1483,1504]" country="Turkey" name="Bursa" pageId="7" pageNumber="696">Bursa</collectingRegion>
|
|||
|
copulatrix similar to that of
|
|||
|
<taxonomicName id="4C404D18FFDDFF94750EA5A65B85FA33" authorityName="Solem" authorityYear="1985" box="[1028,1210,1513,1534]" class="Gastropoda" family="Camaenidae" genus="Mesodontrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF94750EA5A65B85FA33" box="[1028,1210,1513,1534]" italics="true" pageId="7" pageNumber="696">Mesodontrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and
|
|||
|
<taxonomicName id="4C404D18FFDDFF9475FEA5A65A4CFA33" authorityName="Solem" authorityYear="1984" box="[1268,1395,1513,1534]" class="Gastropoda" family="Camaenidae" genus="Ordtrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF9475FEA5A65A4CFA33" box="[1268,1395,1513,1534]" italics="true" pageId="7" pageNumber="696">Ordtrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
but much shorter than in
|
|||
|
<taxonomicName id="4C404D18FFDDFF94754FA6475BD9F9D0" authorityName="Iredale" authorityYear="1939" box="[1093,1254,1544,1565]" class="Gastropoda" family="Camaenidae" genus="Torresitrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF94754FA6475BD9F9D0" box="[1093,1254,1544,1565]" italics="true" pageId="7" pageNumber="696">Torresitrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
; penis differing from that of
|
|||
|
<taxonomicName id="4C404D18FFDDFF9472D9A6695BB6F9F6" authorityName="Solem" authorityYear="1985" box="[979,1161,1574,1595]" class="Gastropoda" family="Camaenidae" genus="Mesodontrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF9472D9A6695BB6F9F6" box="[979,1161,1574,1595]" italics="true" pageId="7" pageNumber="696">Mesodontrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
by presence of epiphallus; epiphallus differing from
|
|||
|
<taxonomicName id="4C404D18FFDDFF94759AA60A5A0EF997" authorityName="Iredale" authorityYear="1939" box="[1168,1329,1605,1626]" class="Gastropoda" family="Camaenidae" genus="Torresitrachia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="genus">
|
|||
|
<emphasis id="B934EA89FFDDFF94759AA60A5A0EF997" box="[1168,1329,1605,1626]" italics="true" pageId="7" pageNumber="696">Torresitrachia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
in shape, thickness of wall, and by absence of flagellum; penial wall differing by presence of main stimulatory pilaster in some
|
|||
|
<taxonomicName id="4C404D18FFDDFF9472C3A6EE5BA5F97A" box="[969,1178,1697,1719]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="696" phylum="Mollusca" rank="species" species="undetermined">
|
|||
|
<emphasis id="B934EA89FFDDFF9472C3A6EE5B07F97B" box="[969,1080,1697,1718]" italics="true" pageId="7" pageNumber="696">Exiligada</emphasis>
|
|||
|
species.
|
|||
|
</taxonomicName>
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|