190 lines
29 KiB
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190 lines
29 KiB
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<document id="EE612BCB2C433FDB6EEA4EEE9A8D380C" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629679974841" checkinUser="felipe" docAuthor="Sartori, Michel & Derleth, Pascale" docDate="2010" docId="038D783A781C7148FF66BBE69FE4F4E7" docLanguage="en" docName="zt02490p039.pdf" docOrigin="Zootaxa 2490" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kilariodes Sartori & Derleth 2010, gen. nov." docType="treatment" docVersion="2" lastPageNumber="36" masterDocId="FFB40042781E714BFFF1BD509B4BF045" masterDocTitle="The dipterous Leptophlebiidae of Borneo (Insecta, Ephemeroptera)" masterLastPageNumber="39" masterPageNumber="33" pageNumber="35" updateTime="1699095124346" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="1AEAC95429DFD8743163401D9251CFB7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0987071A04D72C740E0A8A287FFA1CF6">
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<mods:title id="F769676D53AC52309243CB76460616F1">The dipterous Leptophlebiidae of Borneo (Insecta, Ephemeroptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="2DB9E2D623EBED8428340C3158271877" type="personal">
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<mods:role id="D1AE1FA39B06646E0176896628D361CF">
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<mods:roleTerm id="279ADB93A1C1B010FA4A8850A2517640">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6A31D575C969022A1D59DAEC1B1297AF">Sartori, Michel</mods:namePart>
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</mods:name>
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<mods:name id="DC7FC9BB9C3CA4852DE8D4A61079DE46" type="personal">
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<mods:roleTerm id="117E1F50CBEABE90C0795A15E721839E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3EED82013AA25291BC120A0E141F886B">Derleth, Pascale</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E0EA6DA3C0230B3B6FCC3CC9F183F186">text</mods:typeOfResource>
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<mods:titleInfo id="5FA7ED12B83107E4F9A79CC375D85B43">
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<mods:title id="5578F46DC6D9800190B484643360713E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="99747AF32422A6320E311974BF18C747">
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<mods:date id="065A19008728B0CBD2A2F421FB61642A">2010</mods:date>
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<mods:detail id="5D95BAAA7CD6127547C5BA0233B94115" type="volume">
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<mods:number id="B47BA794A6C37353851385B8744E0990">2490</mods:number>
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</mods:detail>
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<mods:extent id="ED9358C6D85767F9CA3790B32D1D8E2D" unit="page">
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<mods:start id="DE25FA36974479950685855DB2B82D29">33</mods:start>
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<mods:end id="138019AC31FDEF77AD2F8BF6BFC9478F">39</mods:end>
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</mods:extent>
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<mods:classification id="94AC447C15849785FB2DDDA0BE060EDC">journal article</mods:classification>
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<mods:identifier id="25C572F02DDDEB35E1CFC18631E1CB4A" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="038D783A781C7148FF66BBE69FE4F4E7" LSID="urn:lsid:plazi:treatment:038D783A781C7148FF66BBE69FE4F4E7" httpUri="http://treatment.plazi.org/id/038D783A781C7148FF66BBE69FE4F4E7" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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<subSubSection id="C33E9AA7781C7149FF66BBE69ACCF695" box="[151,391,1718,1744]" pageId="2" pageNumber="35" type="nomenclature">
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<paragraph id="8B9BC92C781C7149FF66BBE69ACCF695" blockId="2.[151,391,1718,1744]" box="[151,391,1718,1744]" pageId="2" pageNumber="35">
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<heading id="D0D37E40781C7149FF66BBE69ACCF695" bold="true" box="[151,391,1718,1744]" fontSize="11" level="1" pageId="2" pageNumber="35" reason="1">
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<emphasis id="B950153E781C7149FF66BBE69ACCF695" bold="true" box="[151,391,1718,1744]" pageId="2" pageNumber="35">
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<taxonomicName id="4C24B2AF781C7149FF66BBE69A58F695" ID-CoL="588M" authority="Sartori & Derleth, 2010" authorityName="Sartori & Derleth" authorityYear="2010" box="[151,275,1718,1744]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B950153E781C7149FF66BBE69A58F695" bold="true" box="[151,275,1718,1744]" italics="true" pageId="2" pageNumber="35">Kilariodes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A263A845781C7149FEEBBBE69ACCF695" box="[282,391,1718,1744]" pageId="2" pageNumber="35" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E9AA7781C7148FF66BA509919F1F2" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35" type="materials_examined">
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<paragraph id="8B9BC92C781C7149FF66BA509801F752" blockId="2.[151,842,1792,1816]" box="[151,842,1792,1816]" pageId="2" pageNumber="35">
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<typeStatus id="549F778E781C7149FF66BA509B87F752" box="[151,204,1792,1815]" pageId="2" pageNumber="35">Type</typeStatus>
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species:
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<taxonomicName id="4C24B2AF781C7149FEDCBA519ABCF75D" authority="Sartori & Derleth, 2010" authorityName="Sartori & Derleth" authorityYear="2010" box="[301,503,1793,1816]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="marifae" status="sp. nov.">
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<emphasis id="B950153E781C7149FEDCBA519ABCF75D" box="[301,503,1793,1816]" italics="true" pageId="2" pageNumber="35">Kilariodes marifae</emphasis>
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</taxonomicName>
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<emphasis id="B950153E781C7149FE0FBA50991FF75D" bold="true" box="[510,596,1792,1816]" pageId="2" pageNumber="35">
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<taxonomicNameLabel id="A263A845781C7149FE0FBA50991FF75D" box="[510,596,1792,1816]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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by present designation
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</paragraph>
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<paragraph id="8B9BC92C781C7148FF66BA169A14F1D7" blockId="2.[151,1436,1862,2037]" lastBlockId="3.[151,1436,152,1186]" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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<emphasis id="B950153E781C7149FF66BA169BB7F725" bold="true" box="[151,252,1862,1888]" pageId="2" pageNumber="35">Nymph.</emphasis>
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Head prognathous. Clypeus well developed (
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<figureCitation id="131FD5A9781C7149FCD1BA16983CF725" box="[800,887,1862,1888]" captionStart="FIGURE 19" captionStartId="5.[151,255,1624,1648]" captionTargetBox="[282,1298,525,1591]" captionTargetId="figure-314@5.[282,1310,517,1604]" captionTargetPageId="5" captionText="FIGURE 19. Kilariodes marifae gen.nov. sp.nov. Head of female nymph, dorsal view; cp:clypeus, lb: labrum, mb: right mandible" pageId="2" pageNumber="35">Fig. 19</figureCitation>
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) as broad as the labrum. Labrum (
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<figureCitation id="131FD5A9781C7149FAE9BA169E2DF725" box="[1304,1382,1862,1888]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="2" pageNumber="35">Figs 7</figureCitation>
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,
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<figureCitation id="131FD5A9781C7149FA85BA169ED9F725" box="[1396,1426,1862,1888]" captionStart="FIGURE 19" captionStartId="5.[151,255,1624,1648]" captionTargetBox="[282,1298,525,1591]" captionTargetId="figure-314@5.[282,1310,517,1604]" captionTargetPageId="5" captionText="FIGURE 19. Kilariodes marifae gen.nov. sp.nov. Head of female nymph, dorsal view; cp:clypeus, lb: labrum, mb: right mandible" pageId="2" pageNumber="35">19</figureCitation>
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) with lateral margin rounded and distal margin clearly cordiform; anteromedian emargination well marked (
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<figureCitation id="131FD5A9781C7149FF6EBAC19BACF7EE" box="[159,231,1937,1963]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="2" pageNumber="35">Fig. 8</figureCitation>
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). Left and right mandibles similar (
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<figureCitation id="131FD5A9781C7149FD7FBAC1984CF7EE" box="[654,775,1937,1963]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="2" pageNumber="35">Figs 9–10</figureCitation>
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); outer margin very angular with a hump in the middle. Maxillae (
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<figureCitation id="131FD5A9781C7149FEE4BAE69A27F795" box="[277,364,1974,2000]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="2" pageNumber="35">Fig. 11</figureCitation>
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) stout and robust; palp 3-segmented; inner apical angle of the galea (
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<figureCitation id="131FD5A9781C7149FB37BAE69E54F795" box="[1222,1311,1974,2000]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="2" pageNumber="35">Fig. 13</figureCitation>
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) with one comb-shaped dentiseta, no subapical row of setae. Hypopharynx (
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<figureCitation id="131FD5A9781C7149FC54BA8B98B0F7B0" box="[933,1019,2011,2037]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="2" pageNumber="35">Fig. 14</figureCitation>
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) with superlinguae well developed laterally and rounded; lingua with lateral processes well developed and rounded at apex. Labium (
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<figureCitation id="131FD5A9781D7148FACABDC89ED9F0F7" box="[1339,1426,152,178]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="3" pageNumber="36">Fig. 15</figureCitation>
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) with palp 3-segmented, glossae small and rounded apically, paraglossae large, rounded laterally and inner margin slightly pointed. Metathoracic wing pads missing. Legs (
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<figureCitation id="131FD5A9781D7148FC7CBDB298A8F0B9" box="[909,995,226,252]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="3" pageNumber="36">Fig. 16</figureCitation>
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) with stout femora, ca 2 times longer than wide; claw (
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<figureCitation id="131FD5A9781D7148FE96BC589AF6F167" box="[359,445,264,290]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="3" pageNumber="36">Fig. 17</figureCitation>
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) moderately hooked, with a single row of denticles, the proximal smaller than the others. Gills present on segments I–VII, similar in shape, and each composed of a dorsal and a ventral very long, slender and unbranched lamella. No posterolateral projections on segments II–
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<collectionCode id="ED3551E9781D7148FB66BC029FE7F129" box="[1175,1196,338,364]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="36" type="Museum">V</collectionCode>
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, slightly marked on segments
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<collectionCode id="ED3551E9781D7148FEFABC289A61F1D7" box="[267,298,376,402]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="36">VI</collectionCode>
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–IX.
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</paragraph>
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<paragraph id="8B9BC92C781D7148FF35BCCD9919F1F2" blockId="3.[151,1436,152,1186]" box="[196,594,413,439]" pageId="3" pageNumber="36">Male and female adults unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C33E9AA7781D7148FF37BC929A32F262" pageId="3" pageNumber="36" type="etymology">
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<paragraph id="8B9BC92C781D7148FF37BC929A32F262" blockId="3.[151,1436,152,1186]" pageId="3" pageNumber="36">
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<emphasis id="B950153E781D7148FF37BC929A18F199" bold="true" box="[198,339,450,476]" pageId="3" pageNumber="36">Etymology.</emphasis>
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The genus name is an arbitrary combination of letter to honour Jean-Christophe “Kil” Decker (Pully,
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<collectingCountry id="F33389BC781D7148FF01BCB89ACCF247" box="[240,391,488,514]" name="Switzerland" pageId="3" pageNumber="36">Switzerland</collectingCountry>
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), great friend of us, with reference to the Latin adjective
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<emphasis id="B950153E781D7148FBBBBCB89FD2F244" box="[1098,1177,488,513]" italics="true" pageId="3" pageNumber="36">hilaris</emphasis>
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meaning joyful. The gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E9AA7781D7148FF37BF629FE4F4E7" pageId="3" pageNumber="36" type="distribution">
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<paragraph id="8B9BC92C781D7148FF37BF6298DAF209" blockId="3.[151,1436,152,1186]" box="[198,913,562,588]" pageId="3" pageNumber="36">
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<emphasis id="B950153E781D7148FF37BF629A2CF209" bold="true" box="[198,359,562,588]" pageId="3" pageNumber="36">Distribution.</emphasis>
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Island of Borneo (
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<collectingRegion id="49E007CE781D7148FDB3BF629842F209" box="[578,777,562,588]" country="Indonesia" name="Kalimantan Timur" pageId="3" pageNumber="36">East Kalimantan</collectingRegion>
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,
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<collectingCountry id="F33389BC781D7148FCE4BF6298C2F209" box="[789,905,562,588]" name="Indonesia" pageId="3" pageNumber="36">Indonesia</collectingCountry>
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)
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</paragraph>
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<paragraph id="8B9BC92C781D7148FF37BF0798DDF449" blockId="3.[151,1436,152,1186]" pageId="3" pageNumber="36">
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<emphasis id="B950153E781D7148FF37BF079A0AF234" bold="true" box="[198,321,599,625]" pageId="3" pageNumber="36">Affinities.</emphasis>
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<taxonomicName id="4C24B2AF781D7148FEBBBF089A8DF234" authorityName="Sartori & Derleth" authorityYear="2010" box="[330,454,600,625]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B950153E781D7148FEBBBF089A8DF234" box="[330,454,600,625]" italics="true" pageId="3" pageNumber="36">Kilariodes</emphasis>
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</taxonomicName>
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possesses most of the characters listed by
|
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|
<bibRefCitation id="EFB5B4DD781D7148FC39BF089F2DF237" author="Kluge, N. J." box="[968,1126,600,626]" pageId="3" pageNumber="36" pagination="35 - 43" refId="ref4061" refString="Kluge, N. J. (1994) Habrophlebiinae subfam. n. with description of a new species of Habroleptoides from the Caucasus (Ephemeroptera: Leptophlebiidae). Zoosystematica Rossica, 3, 35 - 43." type="journal article" year="1994">Kluge (1994)</bibRefCitation>
|
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|
to define members of the subfamily
|
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|
<taxonomicName id="4C24B2AF781D7148FEE6BF2D9A90F2D2" authorityName="Peters" authorityYear="1980" box="[279,475,637,663]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="subFamily" subFamily="Atalophlebiinae">Atalophlebiinae</taxonomicName>
|
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|
: hypopharynx with superlinguae laterally developed (
|
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|
<figureCitation id="131FD5A9781D7148FB93BF2D9FF0F2D2" box="[1122,1211,637,663]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="3" pageNumber="36">Fig. 14</figureCitation>
|
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|
), dorsal surface of labrum with a well defined row of setae (
|
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|
<figureCitation id="131FD5A9781D7148FD78BFF29998F2F9" box="[649,723,674,700]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="3" pageNumber="36">Fig. 7</figureCitation>
|
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|
), galea with the proximal dentiseta comb-shaped (
|
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|
<figureCitation id="131FD5A9781D7148FAC5BFF29EC5F2F9" box="[1332,1422,674,700]" captionStart="FIGURES 7–18" captionStartId="4.[151,269,1770,1794]" captionTargetBox="[169,1423,185,1739]" captionTargetId="figure-158@4.[151,1436,159,1747]" captionTargetPageId="4" captionText="FIGURES 7–18. Kilariodes marifae gen. nov. sp. nov. Fig. 7: Labrum (dorsal face at right, ventral face at left); Fig. 8: Detail of the anteromedian emargination of the labrum; Fig. 9: Left mandible; Fig. 10: Right mandible; Fig. 11: Maxilla; Fig. 12: Maxillary palp; Fig. 13: Detail of the inner apical angle of the galea; Fig. 14: Hypopharynx; Fig. 15: Labium (dorsal face at left, ventral face at right); Fig. 16: Foreleg; Fig. 17: Tarsal claw; Fig. 18: Gill VI" pageId="3" pageNumber="36">Fig. 13</figureCitation>
|
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).
|
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|
<taxonomicName id="4C24B2AF781D7148FF66BF989A58F2A4" authorityName="Sartori & Derleth" authorityYear="2010" box="[151,275,712,737]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FF66BF989A58F2A4" box="[151,275,712,737]" italics="true" pageId="3" pageNumber="36">Kilariodes</emphasis>
|
|||
|
</taxonomicName>
|
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|
has few affinities with
|
|||
|
<taxonomicName id="4C24B2AF781D7148FDDDBF98984DF2A4" authorityName="Demoulin" authorityYear="1954" box="[556,774,712,737]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FDDDBF98984DF2A4" box="[556,774,712,737]" italics="true" pageId="3" pageNumber="36">Dipterophlebiodes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
which is a member of
|
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|
<taxonomicName id="4C24B2AF781D7148FBE8BF989F97F2A7" authorityName="Banks" authorityYear="1900" box="[1049,1244,712,738]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="subFamily" subFamily="Leptophlebiinae">Leptophlebiinae</taxonomicName>
|
|||
|
. The new genus presents some similarities with
|
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|
<taxonomicName id="4C24B2AF781D7148FDF7BFBD997DF343" authorityName="Gillies" authorityYear="1951" box="[518,566,749,774]" class="Insecta" family="Leptophlebiidae" genus="Isca" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
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|
<emphasis id="B950153E781D7148FDF7BFBD997DF343" box="[518,566,749,774]" italics="true" pageId="3" pageNumber="36">Isca</emphasis>
|
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|
</taxonomicName>
|
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|
(shape of labrum, hypopharynx or even labium), but is easily separated by the shape of the mandibles, the body size and the gills in lateral position.
|
|||
|
<taxonomicName id="4C24B2AF781D7148FBF0BE429F36F36E" authorityName="Sartori & Derleth" authorityYear="2010" box="[1025,1149,786,811]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FBF0BE429F36F36E" box="[1025,1149,786,811]" italics="true" pageId="3" pageNumber="36">Kilariodes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
presents some affinities with
|
|||
|
<taxonomicName id="4C24B2AF781D7148FF25BE689944F317" authority="Ulmer, 1920" authorityName="Ulmer" authorityYear="1920" box="[212,527,824,850]" class="Insecta" family="Leptophlebiidae" genus="Hagenulodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FF25BE689A3BF314" box="[212,368,824,849]" italics="true" pageId="3" pageNumber="36">Hagenulodes</emphasis>
|
|||
|
Ulmer, 1920
|
|||
|
</taxonomicName>
|
|||
|
, an endemic genus of
|
|||
|
<collectingCountry id="F33389BC781D7148FCEBBE6898D7F317" box="[794,924,824,850]" name="Seychelles" pageId="3" pageNumber="36">Seychelles</collectingCountry>
|
|||
|
. It shares with it the lack of posterior wing pads, and to some extent, the shape of the mandibles, the hypopharynx and the tarsal claw, but it can be easily separated by the shape of the gills (
|
|||
|
<bibRefCitation id="EFB5B4DD781D7148FDC9BED29815F3D9" author="Peters, W. L. & Edmunds, G. F." box="[568,862,898,924]" pageId="3" pageNumber="36" pagination="157 - 240" refId="ref4166" refString="Peters, W. L. & Edmunds, G. F. (1970) Revision of the generic classification of the Eastern hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970">Peters & Edmunds 1970</bibRefCitation>
|
|||
|
). The combination of the following characters is therefore unique and defines the new genus: (1) presence of a hump on the outer margin of mandibles, (2) labrum with a wide anteromedian emargination, (3) maxilla without subapical row of setae, (4) hindwing pads missing, (5) gills slender with two long and unbranched lamellae.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B9BC92C781D7148FF35B9489FE4F4E7" blockId="3.[151,1436,152,1186]" pageId="3" pageNumber="36">
|
|||
|
It could be argued that
|
|||
|
<taxonomicName id="4C24B2AF781D7148FE3EB9489900F474" authorityName="Sartori & Derleth" authorityYear="2010" box="[463,587,1048,1073]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FE3EB9489900F474" box="[463,587,1048,1073]" italics="true" pageId="3" pageNumber="36">Kilariodes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
represents in fact the larval stage of either the genus
|
|||
|
<taxonomicName id="4C24B2AF781D7148FB42B9489ED7F477" authority="Grant & Peters" authorityName="Grant & Peters" authorityYear="1993" box="[1203,1436,1048,1074]" class="Insecta" family="Leptophlebiidae" genus="Sulu" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FB42B9489FACF474" box="[1203,1255,1048,1073]" italics="true" pageId="3" pageNumber="36">Sulu</emphasis>
|
|||
|
Grant & Peters
|
|||
|
</taxonomicName>
|
|||
|
or
|
|||
|
<taxonomicName id="4C24B2AF781D7148FF46B96D9AE2F412" authority="Ulmer" authorityName="Ulmer" authorityYear="1939" box="[183,425,1085,1111]" class="Insecta" family="Leptophlebiidae" genus="Simothraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FF46B96D9A1FF413" box="[183,340,1085,1110]" italics="true" pageId="3" pageNumber="36">Simothraulus</emphasis>
|
|||
|
Ulmer
|
|||
|
</taxonomicName>
|
|||
|
, but according to
|
|||
|
<bibRefCitation id="EFB5B4DD781D7148FD8FB96D98C2F412" author="Grant, P. M. & Peters, W. L." box="[638,905,1085,1111]" pageId="3" pageNumber="36" pagination="131 - 168" refId="ref3972" refString="Grant, P. M. & Peters, W. L. (1993) Description of four new genera of Thraulus group mayflies from the Eastern hemisphere and redescription of Simothraulus and Chiusanophlebia (Ephemeroptera: Leptophlebiidae: Atalophlebiinae). Transactions of the American Entomological Society, 119, 131 - 168." type="journal article" year="1993">Grant & Peters (1993)</bibRefCitation>
|
|||
|
, both genera belong to the
|
|||
|
<taxonomicName id="4C24B2AF781D7148FB3CB96D9E7CF413" authorityName="Eaton" authorityYear="1881" box="[1229,1335,1085,1110]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FB3CB96D9E7CF413" box="[1229,1335,1085,1110]" italics="true" pageId="3" pageNumber="36">Thraulus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
lineage, which possesses gills completely different from those of
|
|||
|
<taxonomicName id="4C24B2AF781D7148FCA4B933989EF439" authorityName="Sartori & Derleth" authorityYear="2010" box="[853,981,1123,1148]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FCA4B933989EF439" box="[853,981,1123,1148]" italics="true" pageId="3" pageNumber="36">Kilariodes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
. A more complete assignment of the phylogenetic position of
|
|||
|
<taxonomicName id="4C24B2AF781D7148FE4AB9D8997CF4E4" authorityName="Sartori & Derleth" authorityYear="2010" box="[443,567,1160,1185]" class="Insecta" family="Leptophlebiidae" genus="Kilariodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B950153E781D7148FE4AB9D8997CF4E4" box="[443,567,1160,1185]" italics="true" pageId="3" pageNumber="36">Kilariodes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
will be possible when the adult stages will be known.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|