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<document id="5E7289F6A21E17B5BEEF994404E2D620" ID-DOI="10.11646/zootaxa.3616.3.3" ID-GBIF-Dataset="436ae0f7-a379-4cb9-a2ef-41a28fc8a279" ID-ISSN="1175-5326" ID-Zenodo-Dep="218827" ID-ZooBank="F70AC1EE-5A70-42B8-B1F2-55C734D26FD9" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460575923727" checkinUser="plazi" docAuthor="Hisamatsu, Sadatomo" docDate="2013" docId="421787CFF66DC870D9F8FEFDFE4EFDD2" docLanguage="en" docName="zt03616p267.pdf" docOrigin="Zootaxa 3616 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pastillus eminentithorax" docType="treatment" docVersion="9" lastPageNumber="266" masterDocId="BE2EFFB7F666C87DD96FFFB8FFE1FFC5" masterDocTitle="A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea)" masterLastPageNumber="267" masterPageNumber="253" pageNumber="264" updateTime="1698323146566" updateUser="plazi">
<mods:mods id="D4FC9F5B0EF125C58345EA7DF718EFD4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="41EBB6B8841CF624E15D7B7174355C37">
<mods:title id="E160BB03C90BC4E50A08E494B7213C10">A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea)</mods:title>
</mods:titleInfo>
<mods:name id="E4DFC4CEAF33333BBA2A7B0EB73C4BB1" type="personal">
<mods:role id="528D76CC929AE344457B705D84A1A027">
<mods:roleTerm id="7A10F0FF71D8F6B76F89E4DDE992A804">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35A6F63906BCF362D93E700CE2CCFEAD">Hisamatsu, Sadatomo</mods:namePart>
</mods:name>
<mods:typeOfResource id="A754FFAD83FC4A81965885582FDDBF7F">text</mods:typeOfResource>
<mods:relatedItem id="8339E2B2FF97826CB39A48C193B4AC44" type="host">
<mods:titleInfo id="41795CFE1CDCCC2513A4DAB75D02A8BC">
<mods:title id="C33253F23C39A0BD0DB1DEE67533C33F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0F83D3C8523DDCA4A6BE05EC840640B5">
<mods:date id="CE9BE3B18A78B14812BEAE18AD0DE86A">2013</mods:date>
<mods:detail id="123A143E0622AFB068ED43B40BA71A80" type="volume">
<mods:number id="CC3D459759B920E11B6CE58D593A11F9">3616</mods:number>
</mods:detail>
<mods:detail id="0D0B8365E7F1DB46CFA8F69A18AB11AD" type="issue">
<mods:number id="C505FC6DDBB4C66BAA3FB20639762195">3</mods:number>
</mods:detail>
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<mods:start id="6D490599182CECEE96151E3D06F5628F">253</mods:start>
<mods:end id="EC90EBEBECECA7F27523BE64D0BED979">267</mods:end>
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<mods:classification id="FFDD93D69892488C0FB132DB6F523D3C">journal article</mods:classification>
<mods:identifier id="0BF0B121B092C67F6824121859F969DC" type="DOI">10.11646/zootaxa.3616.3.3</mods:identifier>
<mods:identifier id="47FA323C90A8575E6F683EF5E7B4B070" type="GBIF-Dataset">436ae0f7-a379-4cb9-a2ef-41a28fc8a279</mods:identifier>
<mods:identifier id="C3D4E8F53D8D6A7A4EBC73B82874BE62" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="36F9E87CA910EF2FF7D77203CD22621A" type="Zenodo-Dep">218827</mods:identifier>
<mods:identifier id="C6AF30FDDE61934175566322EA25D2E7" type="ZooBank">F70AC1EE-5A70-42B8-B1F2-55C734D26FD9</mods:identifier>
</mods:mods>
<treatment id="421787CFF66DC870D9F8FEFDFE4EFDD2" ID-DOI="http://doi.org/10.5281/zenodo.6150621" ID-GBIF-Taxon="119560033" ID-Zenodo-Dep="6150621" LSID="urn:lsid:plazi:treatment:421787CFF66DC870D9F8FEFDFE4EFDD2" httpUri="http://treatment.plazi.org/id/421787CFF66DC870D9F8FEFDFE4EFDD2" lastPageId="13" lastPageNumber="266" pageId="11" pageNumber="264">
<subSubSection id="82A46552F66DC876D9F8FEFDFE6CFEBA" pageId="11" pageNumber="264" type="nomenclature">
<paragraph id="CA0136D9F66DC876D9F8FEFDFD12FE9A" blockId="11.[151,755,325,383]" box="[151,755,325,351]" pageId="11" pageNumber="264">
<heading id="914981B5F66DC876D9F8FEFDFD12FE9A" bold="true" box="[151,755,325,351]" fontSize="11" level="1" pageId="11" pageNumber="264" reason="1">
<emphasis id="F8CAEACBF66DC876D9F8FEFDFD12FE9A" bold="true" box="[151,755,325,351]" pageId="11" pageNumber="264">
<taxonomicName id="0DBE4D5AF66DC876D9F8FEFDFE5EFE9A" box="[151,447,325,351]" class="Insecta" family="Nitidulidae" genus="Pastillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="264" phylum="Arthropoda" rank="species" species="eminentithorax">
<emphasis id="F8CAEACBF66DC876D9F8FEFDFE5EFE9A" bold="true" box="[151,447,325,351]" italics="true" pageId="11" pageNumber="264">Pastillus eminentithorax</emphasis>
</taxonomicName>
S-T. Hisamatsu, sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="CA0136D9F66DC876D9F8FEDFFE6CFEBA" blockId="11.[151,755,325,383]" box="[151,397,359,383]" pageId="11" pageNumber="264">
<figureCitation id="52852A5CF66DC876D9F8FEDFFF38FEBA" box="[151,217,359,383]" captionStart="FIGURE 1" captionStartId="2.[151,250,1886,1909]" captionTargetBox="[165,1427,1464,1851]" captionTargetId="figure@2.[151,1436,1455,1862]" captionTargetPageId="2" captionText="FIGURE 1. Lateral habitus of Cybocephalidae spp. A, Cybocephalus (C.) nipponicus Endrödy-Younga, 1971 (male, Tokunomori, Ehime Pref.); B, Cybocephalus (C.) politissimus Reitter, 1898 (male, Yoshiji-chô, Shimane Pref.); C, Cybocephalus (C.) chlorocapitis sp. nov. (holotype, male, Amami Is.); D, Pastillus eminentithorax sp. nov. (paratype, female, Araike, Aichi Pref.). Scale bars = 1 mm." httpUri="https://zenodo.org/record/218828/files/figure.png" pageId="11" pageNumber="264">Figs 1</figureCitation>
D, 2A, C, 3A, 8
</paragraph>
</subSubSection>
<subSubSection id="82A46552F66DC876D9F8FE08FF08FCB2" pageId="11" pageNumber="264" type="materials_examined">
<paragraph id="CA0136D9F66DC876D9F8FE08FD69FCEA" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
<emphasis id="F8CAEACBF66DC876D9F8FE08FEC3FE0D" bold="true" box="[151,290,431,456]" pageId="11" pageNumber="264">
<typeStatus id="1505887BF66DC876D9F8FE08FF33FE0D" box="[151,210,432,456]" pageId="11" pageNumber="264">Type</typeStatus>
series.
</emphasis>
<typeStatus id="1505887BF66DC876D847FE17FE74FE02" box="[296,405,431,455]" pageId="11" pageNumber="264" type="holotype">Holotype</typeStatus>
: 3, (RYUKYU:
<collectingCountry id="B2A97649F66DC876DB3AFE17FD4DFE02" box="[597,684,431,455]" name="Japan" pageId="11" pageNumber="264">JAPAN</collectingCountry>
) / Shirahama / Iriomote /
<date id="BE001019F66DC876DAA6FE17FBADFE02" box="[969,1100,431,455]" pageId="11" pageNumber="264" value="1988-10-10">10. X. 1988</date>
/ M. Sakai leg. /
<typeStatus id="1505887BF66DC876DC67FE17FA7DFE02" box="[1288,1436,431,455]" pageId="11" pageNumber="264" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="0DBE4D5AF66DC876D9C9FE6CFE57FE29" box="[166,438,468,492]" class="Insecta" family="Nitidulidae" genus="Pastillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="264" phylum="Arthropoda" rank="species" species="eminentithorax">Pastillus eminentithorax</taxonomicName>
<emphasis id="F8CAEACBF66DC876D8D2FE6BFE24FE29" bold="true" box="[445,453,467,492]" pageId="11" pageNumber="264">/</emphasis>
S-T. Hisamatsu, 2011 /.
<typeStatus id="1505887BF66DC876DBB2FE6DFCAFFE28" box="[733,846,469,493]" pageId="11" pageNumber="264" type="paratype">Paratypes</typeStatus>
: 20 exs.: [
<emphasis id="F8CAEACBF66DC876DAADFE6BFBCFFE29" bold="true" box="[962,1070,467,492]" pageId="11" pageNumber="264">MIYAGI</emphasis>
] 1 ex., Takiyama Park, Yamotochô,
<date id="BE001019F66DC876D9A3FE4FFE81FDCA" box="[204,352,503,527]" pageId="11" pageNumber="264" value="2009-08-02">2. VIII. 2009</date>
, T. Ozaki leg. [
<emphasis id="F8CAEACBF66DC876DB66FE4FFD6AFDD5" bold="true" box="[521,651,503,528]" pageId="11" pageNumber="264">SAITAMA</emphasis>
] 1Ƥ, Urawa City,
<date id="BE001019F66DC876DA37FE4FFC07FDCA" box="[856,998,503,527]" pageId="11" pageNumber="264" value="1957-06-18">18. VI. 1957</date>
, H. Kajimura leg. [
<emphasis id="F8CAEACBF66DC876DDD3FE4FFA83FDD5" bold="true" box="[1212,1378,503,528]" pageId="11" pageNumber="264">KANAGAWA</emphasis>
] 1Ƥ, Mt. Kôbô,
<date id="BE001019F66DC876D84DFDA4FE21FDF1" box="[290,448,540,564]" pageId="11" pageNumber="264" value="1989-04-22">22. IV. 1989</date>
, Y. Hirano leg. [
<emphasis id="F8CAEACBF66DC876DBF7FDA3FCE1FDF1" bold="true" box="[664,768,539,564]" pageId="11" pageNumber="264">TOKYO</emphasis>
] 1Ƥ, Sanbancho,
<date id="BE001019F66DC876DA8CFDA4FB8CFDF1" box="[995,1133,540,564]" pageId="11" pageNumber="264" value="2008-10-04">4. X. 2008</date>
, K. Masumoto leg.; 1Ƥ, Shimizudani, Megasaki City,
<date id="BE001019F66DC876D88DFD87FD91FD92" box="[482,624,575,599]" pageId="11" pageNumber="264" value="2001-03-16">16. III. 2001</date>
, Y. Hirano leg. [
<emphasis id="F8CAEACBF66DC876DA48FD87FC9AFD9D" bold="true" box="[807,891,575,600]" pageId="11" pageNumber="264">AICHI</emphasis>
] 1Ƥ, Araike, Tempaku Ward, Nagoya City,
<date id="BE001019F66DC876DC07FD87FF30FDB9" pageId="11" pageNumber="264" value="1995-05-06">6. V. 1995</date>
, H. Yoshitomi leg.; 1Ƥ, Hadenoshi-Hassawa,
<date id="BE001019F66DC876DA6AFDDCFC43FDB9" box="[773,930,612,636]" pageId="11" pageNumber="264" value="1996-04-27">27. IV. 1996</date>
, Y. Hirano leg. [
<emphasis id="F8CAEACBF66DC876DD16FDDBFAC2FDB9" bold="true" box="[1145,1315,611,636]" pageId="11" pageNumber="264">HIROSHIMA</emphasis>
] 1Ƥ, Mt. Haigamine, Kure City,
<date id="BE001019F66DC876D8F1FD3FFDC2FD5A" box="[414,547,647,671]" pageId="11" pageNumber="264" value="2002-09-08">8. IX. 2002</date>
, I. Okamoto leg. [
<emphasis id="F8CAEACBF66DC876DB9EFD3FFC96FD65" bold="true" box="[753,887,647,672]" pageId="11" pageNumber="264">SHIMANE</emphasis>
] 1Ƥ, Kushiro-chô, Masuda City,
<date id="BE001019F66DC876DD9CFD3FFA99FD5A" box="[1267,1400,647,671]" pageId="11" pageNumber="264" value="1996-05-15">15. V. 1996</date>
, Y. Hirano leg. [
<emphasis id="F8CAEACBF66DC876D84BFD14FE64FD01" bold="true" box="[292,389,684,708]" pageId="11" pageNumber="264">EHIME</emphasis>
] 1Ƥ, Ohmishima Is.,
<date id="BE001019F66DC876DB17FD14FCF3FD01" box="[632,786,684,708]" pageId="11" pageNumber="264" value="1959-07-16">16. VII. 1959</date>
, M. Miyatake leg.; 6Ƥ, Shiroyama, Matsuyama City,
<date id="BE001019F66DC876DC06FD14FF33FD22" pageId="11" pageNumber="264" value="1973-05-05">5. V. 1973</date>
, M. Sakai leg. [
<emphasis id="F8CAEACBF66DC876D8EDFD77FE02FD2D" bold="true" box="[386,483,719,744]" pageId="11" pageNumber="264">
<collectingCountry id="B2A97649F66DC876D8EDFD77FE02FD2D" box="[386,483,719,744]" name="India" pageId="11" pageNumber="264">KOCHI</collectingCountry>
</emphasis>
] 1Ƥ, Cape Ashizuri,
<date id="BE001019F66DC876DBBCFD77FC83FD22" box="[723,866,719,743]" pageId="11" pageNumber="264" value="1961-06-13">13. VI. 1961</date>
, S. Hisamatsu leg. [
<emphasis id="F8CAEACBF66DC876DD2CFD77FB3BFD2D" bold="true" box="[1091,1242,719,744]" pageId="11" pageNumber="264">NAGASAKI</emphasis>
] 1Ƥ, Unzen Spa,
<date id="BE001019F66DC876D9F8FD4CFEFAFCC9" box="[151,283,756,780]" pageId="11" pageNumber="264" value="1967-06-05">5. VI. 1967</date>
, S. Hisamatsu leg.; 1Ƥ, Mt. Fugen~Mt. Myoken,
<date id="BE001019F66DC876DA21FD4DFC28FCC9" box="[846,969,756,780]" pageId="11" pageNumber="264" value="1970-10-07">7. X. 1970</date>
, S. Hisamatsu leg. [
<emphasis id="F8CAEACBF66DC876DDC3FD4BFA81FCC9" bold="true" box="[1196,1376,755,780]" pageId="11" pageNumber="264">KAGOSHIMA</emphasis>
] 1Ƥ, Sendai City, 28~
<date id="BE001019F66DC876D83CFCAFFE00FCEA" box="[339,481,791,815]" pageId="11" pageNumber="264" value="2001-03-24">24. III. 2001</date>
, Y. Hirano leg.
</paragraph>
<paragraph id="CA0136D9F66DC876D9A6FC84FF08FCB2" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
<emphasis id="F8CAEACBF66DC876D9A6FC84FE71FC91" bold="true" box="[201,400,827,852]" pageId="11" pageNumber="264">
<typeStatus id="1505887BF66DC876D9A6FC84FEE5FC91" box="[201,260,828,852]" pageId="11" pageNumber="264">Type</typeStatus>
depository.
</emphasis>
The
<typeStatus id="1505887BF66DC876D8A2FC84FDCEFC91" box="[461,559,828,852]" pageId="11" pageNumber="264" type="holotype">holotype</typeStatus>
and 15
<typeStatus id="1505887BF66DC876DBE2FC85FD18FC90" box="[653,761,829,853]" pageId="11" pageNumber="264" type="paratype">paratypes</typeStatus>
are deposited in the EUM. Five
<typeStatus id="1505887BF66DC876DD1CFC85FB3FFC90" box="[1139,1246,829,853]" pageId="11" pageNumber="264" type="paratype">paratypes</typeStatus>
are deposited in PCYH.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F66DC876D9A6FC3CFC72FC05" pageId="11" pageNumber="264" type="etymology">
<paragraph id="CA0136D9F66DC876D9A6FC3CFC72FC05" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
<emphasis id="F8CAEACBF66DC876D9A6FC3CFEB1FC59" bold="true" box="[201,336,900,924]" pageId="11" pageNumber="264">Etymology.</emphasis>
The specific name means “projecting pronotum (eminens + thorax)”, which is derived from the strongly projecting lateral margins of the pronotum in lateral aspect.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F66DC876D9A6FC73FC00FBE9" pageId="11" pageNumber="264" type="diagnosis">
<paragraph id="CA0136D9F66DC876D9A6FC73FC00FBE9" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
<emphasis id="F8CAEACBF66DC876D9A6FC73FEDDFC21" bold="true" box="[201,316,971,996]" pageId="11" pageNumber="264">Diagnosis</emphasis>
. Body strongly convex, almost hemispherical. Antennae (
<figureCitation id="52852A5CF66DC876DAABFC74FBE4FC21" box="[964,1029,972,996]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
K) 10-segmented, with 2-segmented club; scape strongly enlarged, widest at middle. Lateral margins of pronotum in lateral view strongly projecting. Inter-metacoxal lobe of abdominal sternite III (
<figureCitation id="52852A5CF66DC876DBC9FBACFD09FBE9" box="[678,744,1044,1068]" captionStart="FIGURE 2" captionStartId="3.[151,250,1105,1128]" captionTargetBox="[155,1427,198,1078]" captionTargetId="figure@3.[151,1436,193,1085]" captionTargetPageId="3" captionText="FIGURE 2. Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) nipponicus Endrödy-Younga, 1971. A, C, Pastillus eminentithorax; B, D, Cybocephalus (C.) nipponicus. A, B, body, ventral view; C, D, scutellum. A = Mt. Kôbô, Kanagawa Pref.; B, D = Morimatsu, Ehime Pref.; C = Mt. Haigamine, Hiroshima Pref." httpUri="https://zenodo.org/record/218829/files/figure.png" pageId="11" pageNumber="264">Fig. 2</figureCitation>
A) strongly narrowed.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F66DC870D9A6FB8FFDC4FF6A" lastPageId="13" lastPageNumber="266" pageId="11" pageNumber="264" type="description">
<paragraph id="CA0136D9F66DC876D9A6FB8FFEFDFBB1" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
<emphasis id="F8CAEACBF66DC876D9A6FB8FFEBAFB95" bold="true" box="[201,347,1079,1104]" pageId="11" pageNumber="264">Description.</emphasis>
Length 1.66–1.83 (1.78) mm (n = 5), width 0.70–0.80 (0.61) mm (n = 5), height 0.63–0.70 (0.67) mm (n = 5).
</paragraph>
<paragraph id="CA0136D9F66DC876D9A8FB38FD1EFB1A" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
<emphasis id="F8CAEACBF66DC876D9A8FB38FEE0FB5D" box="[199,257,1152,1176]" italics="true" pageId="11" pageNumber="264">Male</emphasis>
. Body highly convex and shining, contractile; dorsal disc almost glabrous. Coloration black to dark brown; antennal segments 2–8, maxillary and labial palpi, prosternum, mesoventrite, procoxae, protrochanters, profemora, mesocoxae and mesotrochanters yellowish.
</paragraph>
<paragraph id="CA0136D9F66DC876D9A8FB54FB4EFA51" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
Head wide; vertex distinctly punctuate; interspaces smooth; apical margin distinctly bordered, with deep emaginations for reception of antennal scape. Labrum (
<figureCitation id="52852A5CF66DC876DA6FFAB7FCA0FAE2" box="[768,833,1295,1319]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
G) about as wide as long, with truncate apical margin. Eyes large, but scarcely visible in ventral aspect. Mentum (
<figureCitation id="52852A5CF66DC876DA56FA8CFC9DFA89" box="[825,892,1332,1356]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
J) trapezoidal. Antennal grooves long and deep. Antennae (
<figureCitation id="52852A5CF66DC876D87DFAEFFEB5FAAA" box="[274,340,1367,1391]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
K) 10-segmented, with 2-segmented club; scape strongly swollen, widest at middle; approximate ratio of each segment (n = 1) is 2.55: 1.96: 1.25: 1.11: 1.24: 1.02: 1.05: 1.00: 1.51: 4.33.
</paragraph>
<paragraph id="CA0136D9F66DC876D9A8FA18FD07FA3A" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
Pronotum strongly convex, 1.70–1.78 (1.74) times as wide as long (n = 5); lateral margins strongly projecting in lateral aspect; dorsal disc with minute punctures and short recumbent sparse setae; interspaces smooth. Scutellum (
<figureCitation id="52852A5CF66DC876D877FA5FFEBBFA3A" box="[280,346,1511,1535]" captionStart="FIGURE 2" captionStartId="3.[151,250,1105,1128]" captionTargetBox="[155,1427,198,1078]" captionTargetId="figure@3.[151,1436,193,1085]" captionTargetPageId="3" captionText="FIGURE 2. Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) nipponicus Endrödy-Younga, 1971. A, C, Pastillus eminentithorax; B, D, Cybocephalus (C.) nipponicus. A, B, body, ventral view; C, D, scutellum. A = Mt. Kôbô, Kanagawa Pref.; B, D = Morimatsu, Ehime Pref.; C = Mt. Haigamine, Hiroshima Pref." httpUri="https://zenodo.org/record/218829/files/figure.png" pageId="11" pageNumber="264">Fig. 2</figureCitation>
C) triangular, with acuminate apex.
</paragraph>
<paragraph id="CA0136D9F66DC876D9A8F9B4FE33F881" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
Elytra strongly convex in lateral aspect, conjointly 1.17–1.24 (1.19) times as long as wide (n = 5), 1.53–1.73 (1.61) times as long as pronotum (n = 5); disc (
<figureCitation id="52852A5CF66DC876DBA3F997FCF0F982" box="[716,785,1583,1607]" captionStart="FIGURE 3" captionStartId="4.[151,250,1216,1239]" captionTargetBox="[157,1430,225,1157]" captionTargetId="figure@4.[151,1436,209,1173]" captionTargetPageId="4" captionText="FIGURE 3. Elytral punctuation of Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) spp. A, Pastillus eminentithorax sp. nov.; B, Cybocephalus (C.) nipponicus Endrödy-Younga, 1971; C, D, Cybocephalus (C.) politissimus Reitter, 1898. A = female, Mt. Haigamine, Hiroshima Pref.; B = male, Morimatsu, Ehime Pref.; C = male, Yakushima Is.; D = male, Sugitate, Ehime Pref." httpUri="https://zenodo.org/record/218830/files/figure.png" pageId="11" pageNumber="264">Fig. 3</figureCitation>
A) with minute, uniform punctures; interspaces smooth. Prosternum strongly carinate at middle. Metaventrite (
<figureCitation id="52852A5CF66DC876DB9AF9ECFCD6F9A9" box="[757,823,1620,1644]" captionStart="FIGURE 2" captionStartId="3.[151,250,1105,1128]" captionTargetBox="[155,1427,198,1078]" captionTargetId="figure@3.[151,1436,193,1085]" captionTargetPageId="3" captionText="FIGURE 2. Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) nipponicus Endrödy-Younga, 1971. A, C, Pastillus eminentithorax; B, D, Cybocephalus (C.) nipponicus. A, B, body, ventral view; C, D, scutellum. A = Mt. Kôbô, Kanagawa Pref.; B, D = Morimatsu, Ehime Pref.; C = Mt. Haigamine, Hiroshima Pref." httpUri="https://zenodo.org/record/218829/files/figure.png" pageId="11" pageNumber="264">Fig. 2</figureCitation>
A) depressed, short, about as wide as metafemora; disc asetose. Inter-metacoxal distance strongly narrow, separated by 1.28 (1.28) times as wide as inter-mesocoxal distance (n = 1). Inter-mesocoxal distance strongly narrowed. Abdominal sternites (
<figureCitation id="52852A5CF66DC876DD36F924FB7FF971" box="[1113,1182,1692,1716]" captionStart="FIGURE 2" captionStartId="3.[151,250,1105,1128]" captionTargetBox="[155,1427,198,1078]" captionTargetId="figure@3.[151,1436,193,1085]" captionTargetPageId="3" captionText="FIGURE 2. Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) nipponicus Endrödy-Younga, 1971. A, C, Pastillus eminentithorax; B, D, Cybocephalus (C.) nipponicus. A, B, body, ventral view; C, D, scutellum. A = Mt. Kôbô, Kanagawa Pref.; B, D = Morimatsu, Ehime Pref.; C = Mt. Haigamine, Hiroshima Pref." httpUri="https://zenodo.org/record/218829/files/figure.png" pageId="11" pageNumber="264">Fig. 2</figureCitation>
A) covered with long, recumbent yellowish setae; inter-metacoxal area of abdominal sternite III strongly narrowed, triangular; femoral lines (
<figureCitation id="52852A5CF66DC876D98EF95CFEC8F939" box="[225,297,1764,1788]" captionStart="FIGURE 2" captionStartId="3.[151,250,1105,1128]" captionTargetBox="[155,1427,198,1078]" captionTargetId="figure@3.[151,1436,193,1085]" captionTargetPageId="3" captionText="FIGURE 2. Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) nipponicus Endrödy-Younga, 1971. A, C, Pastillus eminentithorax; B, D, Cybocephalus (C.) nipponicus. A, B, body, ventral view; C, D, scutellum. A = Mt. Kôbô, Kanagawa Pref.; B, D = Morimatsu, Ehime Pref.; C = Mt. Haigamine, Hiroshima Pref." httpUri="https://zenodo.org/record/218829/files/figure.png" pageId="11" pageNumber="264">Fig. 2</figureCitation>
A) strongly expressed beyond anterior margin of abdominal sternite III. Legs (
<figureCitation id="52852A5CF66DC876DDB5F95CFAC1F939" box="[1242,1312,1764,1788]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
D–F) with protibiae slender, inner and outer margin slightly sinuate; metatibiae flat and wide, strongly widened from basal third to apex. Claws simple.
</paragraph>
<paragraph id="CA0136D9F66DC876D9A6F8F7FC32F811" blockId="11.[151,1437,431,2004]" pageId="11" pageNumber="264">
Male genitalia with tegmen (
<figureCitation id="52852A5CF66DC876DB7BF8F7FDB9F8A2" box="[532,600,1871,1895]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
A) in ventral view constricted at middle; anterior margin of tegmen with two projections. Median lobe (
<figureCitation id="52852A5CF66DC876D89AF8CCFDD9F849" box="[501,568,1908,1932]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
B) in ventral view subparallel-sided from base to middle, abruptly attenuate anteriorly; anterior margin more or less truncate in ventral aspect; apico-lateral corners with tuft of long setae in ventral aspect. Internal sac sclerites of male genitalia (
<figureCitation id="52852A5CF66DC876DB99F804FCD9F811" box="[758,824,1980,2004]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="11" pageNumber="264">Fig. 8</figureCitation>
C) as figured.
</paragraph>
<caption id="9EC16651F66AC871D9F8F95BFEE1F893" httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="12" pageNumber="265" targetBox="[164,1431,178,1730]" targetPageId="12">
<paragraph id="CA0136D9F66AC871D9F8F95BFEE1F893" blockId="12.[151,1436,1763,1878]" pageId="12" pageNumber="265">
<emphasis id="F8CAEACBF66AC871D9F8F95BFEF3F93C" bold="true" box="[151,274,1763,1786]" pageId="12" pageNumber="265">FIGURE 8.</emphasis>
<taxonomicName id="0DBE4D5AF66AC871D877F95CFDF1F93F" box="[280,528,1764,1786]" class="Insecta" family="Nitidulidae" genus="Pastillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="265" phylum="Arthropoda" rank="species" species="eminentithorax" status="sp. nov.">
<emphasis id="F8CAEACBF66AC871D877F95CFDF1F93F" box="[280,528,1764,1786]" italics="true" pageId="12" pageNumber="265">Pastillus eminentithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3F957B0F66AC871DB79F95DFD85F93F" box="[534,612,1765,1786]" pageId="12" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A–C = holotype, Amami Is.; D–K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="CA0136D9F66AC871D9A8F83BFAB0F827" blockId="12.[151,1436,1922,2018]" pageId="12" pageNumber="265">
<emphasis id="F8CAEACBF66AC871D9A8F83BFEFBF85E" box="[199,282,1923,1947]" italics="true" pageId="12" pageNumber="265">Female</emphasis>
. Apical margin of abdominal tergite and sternite VII rounded. Ovipositor (
<figureCitation id="52852A5CF66AC871DD39F83AFB76F85F" box="[1110,1175,1922,1946]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="12" pageNumber="265">
Fig.
<date id="BE001019F66AC871DDE5F83AFB76F85F" box="[1162,1175,1922,1946]" pageId="12" pageNumber="265">8</date>
</figureCitation>
I) elongate and slender; coxites feebly sclerotized, deeply cleft and appearing separated, apex of stylus with 3~4 setae; 5~6 setae present along apico-lateral margin. Spermatheca (
<figureCitation id="52852A5CF66AC871DB07F872FD48F827" box="[616,681,1994,2018]" captionStart="FIGURE 8" captionStartId="12.[151,250,1763,1786]" captionTargetBox="[164,1431,178,1730]" captionTargetId="figure@12.[151,1436,163,1749]" captionTargetPageId="12" captionText="FIGURE 8. Pastillus eminentithorax sp. nov. A, tegmen, ventral view; B, median lobe, ventral view; C, sclerite of internal sac; D, left protibia; E, left mesotibia; F, left metatibia; G, labrum; H, spermatheca, spermathecal gland, and sperm duct; I, ovipositor; J, mentum; K, right antenna. A – C = holotype, Amami Is.; D – K = paratype, Ohmishima Is., Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218835/files/figure.png" pageId="12" pageNumber="265">Fig. 8</figureCitation>
H) C-shaped, spermathecal gland distinct, sperm duct coiled.
</paragraph>
<paragraph id="CA0136D9F66BC870D9A6FF2FFDC4FF6A" blockId="13.[151,1436,151,535]" box="[201,549,151,176]" pageId="13" pageNumber="266">
<emphasis id="F8CAEACBF66BC870D9A6FF2FFEACFF75" bold="true" box="[201,333,151,176]" pageId="13" pageNumber="266">Bionomics.</emphasis>
Biology unknown.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F66BC870D9A6FF03FAEBFF11" box="[201,1290,187,212]" pageId="13" pageNumber="266" type="distribution">
<paragraph id="CA0136D9F66BC870D9A6FF03FAEBFF11" blockId="13.[151,1436,151,535]" box="[201,1290,187,212]" pageId="13" pageNumber="266">
<emphasis id="F8CAEACBF66BC870D9A6FF03FEBAFF11" bold="true" box="[201,347,187,212]" pageId="13" pageNumber="266">Distribution</emphasis>
. This species is known from Honshû, Shikoku, Kyûshû, and Iriomote Island,
<collectingCountry id="B2A97649F66BC870DDACFF05FAE7FF11" box="[1219,1286,189,212]" name="Japan" pageId="13" pageNumber="266">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F66BC870D9A6FF58FE4EFDD2" pageId="13" pageNumber="266" type="discussion">
<paragraph id="CA0136D9F66BC870D9A6FF58FEF7FE69" blockId="13.[151,1436,151,535]" pageId="13" pageNumber="266">
<emphasis id="F8CAEACBF66BC870D9A6FF58FED3FF3D" bold="true" box="[201,306,224,248]" pageId="13" pageNumber="266">Remark.</emphasis>
This new species is similar to
<taxonomicName id="0DBE4D5AF66BC870DBF9FF58FBD7FF32" authority="Endrody-Younga" authorityName="Endrody-Younga" box="[662,1078,223,248]" class="Insecta" family="Nitidulidae" genus="Pastillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="conflexus">
<emphasis id="F8CAEACBF66BC870DBF9FF58FC8AFF3D" box="[662,875,224,248]" italics="true" pageId="13" pageNumber="266">Pastillus conflexus</emphasis>
Endrödy-Younga
</taxonomicName>
, known from
<collectingCountry id="B2A97649F66BC870DDB8FF67FA8DFF32" box="[1239,1388,223,247]" name="Sierra Leone" pageId="13" pageNumber="266">Sierra Leone</collectingCountry>
, but can be distinguished from the latter by the ten-segmented antennae and peculiar shape of male genitalia.
<taxonomicName id="0DBE4D5AF66BC870DC55FEBCFEA0FE85" class="Insecta" family="Nitidulidae" genus="Pastillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="eminentithorax" status="sp. nov.">
<emphasis id="F8CAEACBF66BC870DC55FEBCFEA0FE85" italics="true" pageId="13" pageNumber="266">Pastillus eminentithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3F957B0F66BC870D827FE90FE7EFEFA" box="[328,415,296,319]" pageId="13" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
is widely distributed in
<collectingCountry id="B2A97649F66BC870DBDEFE90FD15FEFA" box="[689,756,296,319]" name="Japan" pageId="13" pageNumber="266">Japan</collectingCountry>
. Twenty-one specimens were examined, with only one male found, which was collected on Iriomote Is., Okinawa Pref. This preliminary data on sex ratios may indicate a parthenogenetic lifestyle of the species. However, further examination and field data are needed to corroborate this hypothesis.
</paragraph>
<paragraph id="CA0136D9F66BC870D9A6FE00FE4EFDD2" blockId="13.[151,1436,151,535]" pageId="13" pageNumber="266">
<taxonomicName id="0DBE4D5AF66BC870D9A6FE00FECAFE15" box="[201,299,440,464]" class="Insecta" family="Nitidulidae" genus="Pastillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="F8CAEACBF66BC870D9A6FE00FECAFE15" box="[201,299,440,464]" italics="true" pageId="13" pageNumber="266">Pastillus</emphasis>
</taxonomicName>
is a small genus, consisting of three species, with previously known distributions limited to localities in the Afrotropical Region. This new species represents the fourth species of the genus, and is the first recorded from Palaearctic Region.
</paragraph>
</subSubSection>
</treatment>
</document>
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