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161 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.241.3802" ID-GBIF-Dataset="5ef2e002-5b7e-44b8-a18e-dd5458da7186" ID-PMC="PMC3559108" ID-Pensoft-Pub="1313-2970-241-33" ID-PubMed="23372412" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-241-33" ModsDocOrigin="ZooKeys 241" ModsDocTitle="Synopsis of the Argentinian scarab genus Pseudogeniates Ohaus (Coleoptera, Scarabaeidae, Rutelinae)" checkinTime="1451248563233" checkinUser="pensoft" docAuthor="Jameson, Mary Liz & Ocampo, Federico" docDate="2012" docId="A3712DC185CF4D70D68F072ACCAC19BD" docLanguage="en" docName="ZooKeys 241: 33-53" docOrigin="ZooKeys 241" docSource="http://dx.doi.org/10.3897/zookeys.241.3802" docTitle="Pseudogeniates cordobaensis Soula 2009" docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="B070FFBFFFB1FFA2FFD1D153D71DFF83" masterDocTitle="Synopsis of the Argentinian scarab genus Pseudogeniates Ohaus (Coleoptera, Scarabaeidae, Rutelinae)" masterLastPageNumber="53" masterPageNumber="33" pageNumber="39" updateTime="1668154706406" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Synopsis of the Argentinian scarab genus Pseudogeniates Ohaus (Coleoptera, Scarabaeidae, Rutelinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jameson, Mary Liz</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ocampo, Federico</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>241</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>53</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.241.3802</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.241.3802</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-241-33</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152037872" LSID="urn:lsid:plazi:treatment:A3712DC185CF4D70D68F072ACCAC19BD" httpUri="http://treatment.plazi.org/id/A3712DC185CF4D70D68F072ACCAC19BD" lastPageId="8" lastPageNumber="41" pageId="6" pageNumber="39">
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<subSubSection pageId="6" pageNumber="39" type="nomenclature">
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<paragraph pageId="6" pageNumber="39">
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<taxonomicName LSID="http://species-id.net/wiki/Pseudogeniates_cordobaensis" authority="Soula, 2009" authorityName="Soula" authorityYear="2009" class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates cordobaensis" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="cordobaensis">Pseudogeniates cordobaensis Soula, 2009</taxonomicName>
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Figs 1, 5, 6, 8, 11, 14, 16, 19, 22
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="39" type="reference_group">
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<paragraph pageId="6" pageNumber="39">
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates cordobaensis" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="cordobaensis">Pseudogeniates cordobaensis</taxonomicName>
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<bibRefCitation author="Soula, M" journalOrPublisher="Photos de toutes le especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: Pelidnotina). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT; Saintry, France" pageId="14" pageNumber="47" title="Les coleopteres du nouveau monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3." year="2009">Soula 2009</bibRefCitation>
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: 122.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="39" type="material examined">
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<paragraph pageId="6" pageNumber="39">Material examined.</paragraph>
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<paragraph pageId="6" pageNumber="39">
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Holotype male, from
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<normalizedToken originalValue="Ohaus’s">Ohaus's</normalizedToken>
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type series of
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates intermedius" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="intermedius">Pseudogeniates intermedius</taxonomicName>
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, at ZMHB labeled: a) "Argentina S. d. Cordoba J. Hubrich S." (typeset, white label), b) male symbol, c) "Pseudogeniates intermedius cotype Ohs." (
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<normalizedToken originalValue="Ohaus’s">Ohaus's</normalizedToken>
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handwritten, red label), d) "SYNTYPUS Pseudogeniates intermedius Ohaus, 1914 labelled by MNHUS 2007" (typeset, red label), f) "Paralectotype 2009 Pseudogeniates intermedius Oh. Soula det." (typeset and handwritten, red label), g) "Holotype 2009 Pseudogeniates cordobaensis Soula Soula" (handwritten and typeset, red label).
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<bibRefCitation author="Soula, M" journalOrPublisher="Photos de toutes le especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: Pelidnotina). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT; Saintry, France" pageId="14" pageNumber="47" title="Les coleopteres du nouveau monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3." year="2009">Soula (2009)</bibRefCitation>
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based the description on one male specimen.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39" type="description">
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<paragraph pageId="6" pageNumber="39">Description</paragraph>
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<paragraph lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39">
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(based on 10 males and 17 females). Length 13.8-17.0 mm. Widest width 6.8-8.0 mm. Color: Dorsum, venter, and appendages rufous-castaneous (Fig. 1)
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<pageBreakToken pageId="7" pageNumber="40" start="start">.</pageBreakToken>
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Head (Fig. 16): Frons densely punctate to confluently punctate (mid-disc), rugopunctate laterally and apically; punctures small and moderately large, mixed. Clypeus with dorsal surface densely punctate to confluently punctate (mid-disc), rugopunctate laterally and apically, punctures small and large (mixed), some setose; setae short to moderate, rufous, sparse; shape broadly semicircular, apex and sides broadly reflexed, with or without constriction basolaterally; disc (frontal view) densely rugopunctate, punctures small and large, mixed, some setose (setae short to moderately long, rufous, sparse). Labrum setose; setae moderately long and short, rufous. Maxilla (Fig. 5) with galea distinct, with 1 elongate, bristle-like, medial seta. Mentum (Fig. 8) nearly as long as wide or slightly longer than wide, form pentagonal, inner apex projects anteriorly, with inner shelf. Antenna 10-segmented with 3 segmented club; club slightly longer than stem. Pronotum: Medial line obsolete. Surface (disc and mediolaterally) moderately densely imbricate with sparse, short, testaceous setae. Margin with short, rufous setae. Elytra: Sutural stria impressed from base of scutellum to apex. Elytral apex weakly spiniform or quadrate. Discal striae weakly impressed, punctate; 5 on disc mesad of humerus, 5 laterad of humerus; discal stria 1 (adjacent to sutural stria) extending from base to apical umbone; striae 2-3 and 4-5 paired, extending from near base to apical umbone, stria 5 faintly impressed and incomplete; humeral striae 1-4 faintly impressed, incomplete, indicated by shallow punctures, stria 5 moderately impressed, extending from base of metacoxa to subapex; small to moderate in size, some longitudinal (Fig. 5). Intervals with moderately dense, random punctures, punctures minute to moderate in size; interval 1 broader than others. Pygidium: Disc and apex sparsely punctate; base and sides moderately densely imbricate, nearly obsolete; punctures minute to small. Legs: Protarsomere 5 of male subequal to tarsomeres 2-4. Modified proclaw of male subequal in length to tarsomeres 2-4; internoapical tooth present. Protibia (Fig. 11) (male and female) with teeth evenly separated; basal 1/3 weakly dilated (male) or moderately dilated (female). Mesotibia (male) with sides subparallel, weakly divergent towards apex (male) or weakly divergent (female); apex oblique with 10-11 moderately long spinulae; external edge with weak carina in basal 1/3, 1 carina in apical 1/3. Metatrochanter (male) weakly produced beyond posterior border of femur. Metacoxal corner square (female). Metacoxa at middle weakly produced posteriorly. Metatibia of male (Fig. 14) with sides subparallel, weakly divergent towards apex; external edge with 1 carina in basal 1/3 (faint), 1 carina in apical 1/3 (faint); apex without corbel, with 2 inner spurs (spurs equal in width in male, weakly curved at apex), inner apex with 28 short, stout spinulae. Metatibia of female divergent at apical 1/5; external edge with 1 carina in basal 1/3, 1 carina in apical 1/3; apex without corbel, with 2 inner spurs, inner apex with 28 short, stout spinulae. Parameres: Ventral plate nearly as long as dorsal plate with sides converging to a quadrate apex (Fig. 19).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="41" pageId="7" pageNumber="40" type="diagnosis">
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<paragraph pageId="7" pageNumber="40">Diagnosis.</paragraph>
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<paragraph lastPageId="8" lastPageNumber="41" pageId="7" pageNumber="40">
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates cordobaensis" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="cordobaensis">Pseudogeniates cordobaensis</taxonomicName>
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is separated from other species in the genus by the pentagonal form of the mentum (width subequal to length) that has the inner apex projecting anteriorly and has an inner shelf (Fig. 8). In comparison, the form of mentum is longer than wide in
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates intermedius" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="intermedius">Pseudogeniates intermedius</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates richterianus" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="richterianus">Pseudogeniates richterianus</taxonomicName>
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(Figs 9-10). In
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates intermedius" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="intermedius">Pseudogeniates intermedius</taxonomicName>
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, the inner apex of the mentum does not project anteriorly, but does possess
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<pageBreakToken pageId="8" pageNumber="41" start="start">an</pageBreakToken>
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inner shelf (Fig. 9); in
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates richterianus" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="richterianus">Pseudogeniates richterianus</taxonomicName>
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, the inner apex projects anteriorly, but does not possess an inner shelf (Fig. 10). The ventral plate of the male parameres in
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates cordob-aensis" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="cordob-aensis">Pseudogeniates cordob-aensis</taxonomicName>
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is nearly as long as dorsal plate with sides converging to a quadrate apex (Fig. 19). The ventral plate of
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates richterianus" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="richterianus">Pseudogeniates richterianus</taxonomicName>
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is about half the length of the dorsal plate (Fig. 21), whereas in
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates intermedius" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="intermedius">Pseudogeniates intermedius</taxonomicName>
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the ventral plate is nearly as long as the dorsal plate, but the sides converge with a weak constriction preapically and a rounded apex (Fig. 20).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="41" type="distribution">
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<paragraph pageId="8" pageNumber="41">Distribution</paragraph>
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<paragraph pageId="8" pageNumber="41">
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(Fig. 22).
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates cordobaensis" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="cordobaensis">Pseudogeniates cordobaensis</taxonomicName>
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is distributed in the Monte de Sierras y Bolsones in the Monte eco-region in Argentina. The distribution of this species coincides with the Montane Forest region (
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<bibRefCitation author="Navarro, FR" journalOrPublisher="Revista Chilena de Historia Natural" pageId="14" pageNumber="47" pagination="507 - 522" title="Can insect data be used to infer areas of endemism? An example from the Yungas of Argentina." url="10.4067/S0716-078X2009000400006" volume="82" year="2009">Navarro et al. 2009</bibRefCitation>
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) and the Yungas Forest area of endemism in Argentina (
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<bibRefCitation author="Szumik, C" journalOrPublisher="Cladistics" pageId="14" pageNumber="47" pagination="317 - 329" title="Detecting areas of endemism with a taxonomically diverse data set: plants, mammals, reptiles, amphibians, birds, and insects from Argentina." url="10.1111/j.1096-0031.2011.00385.x" volume="28" year="2012">Szumik et al. 2012</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="41" type="locality data">
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<paragraph pageId="8" pageNumber="41">Locality data.</paragraph>
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<paragraph pageId="8" pageNumber="41">
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27 specimens from IAZA, WSUC, and ZMHB. ARGENTINA (27): CATAMARCA (26): Salar de Pipanaco,
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<normalizedToken originalValue="Pío">Pio</normalizedToken>
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Brizuela (37 km S
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<normalizedToken originalValue="Andalgalá">Andalgala</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-27.826113">27°49'34"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-66.24638">66°14'47"W</geoCoordinate>
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, 751 m),
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<normalizedToken originalValue="Quirós">Quiros</normalizedToken>
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(2 km S on RN 157,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-28.812027">28°48'43.3"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-65.10628">65°06'22.6"W</geoCoordinate>
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),
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<normalizedToken originalValue="CÓRDOBA">CORDOBA</normalizedToken>
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(1): Huerta Grande (Sierra de
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<normalizedToken originalValue="Córdoba">Cordoba</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="41" type="temporal data">
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<paragraph pageId="8" pageNumber="41">Temporal data.</paragraph>
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<paragraph pageId="8" pageNumber="41">December (9).</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="41" type="remarks">
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<paragraph pageId="8" pageNumber="41">Remarks.</paragraph>
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<paragraph pageId="8" pageNumber="41">
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The holotype specimen for this species was part of the type series for
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates intermedius" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="intermedius">Pseudogeniates intermedius</taxonomicName>
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, a series that included three specimens from Santiago del Estero and one specimen (=
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<taxonomicName class="Insecta" family="Rutelidae" genus="Pseudogeniates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeniates cordobaensis" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="cordobaensis">Pseudogeniates cordobaensis</taxonomicName>
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) from Huerta Grande in the Sierra de
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<normalizedToken originalValue="Cordóba">Cordoba</normalizedToken>
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,
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<normalizedToken originalValue="Córdoba">Cordoba</normalizedToken>
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Province (
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<bibRefCitation author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="14" pageNumber="47" pagination="299 - 304" title="Neue Coleoptera lamellicornia aus Argentinien. (Col.) IV." url="10.1002/mmnd.191419140307" volume="1914" year="1914">Ohaus 1914</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="41" type="natural history">
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<paragraph pageId="8" pageNumber="41">Natural history.</paragraph>
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<paragraph pageId="8" pageNumber="41">This species is attracted to black lights. It was recorded from an elevation of 751 m.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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