treatments-xml/data/EE/05/53/EE05535497354343FF23FE76C564C0B7.xml

<document id="9F92F47574FC5D68A64188330A5B540A" ID-DOI="10.5281/zenodo.178845" ID-GBIF-Dataset="41e4b4a1-34ba-4984-8bb0-812131adda08" ID-ISSN="1175-5326" ID-Zenodo-Dep="178845" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459927027021" checkinUser="plazi" docAuthor="Wu, Shu-Ping, Lin, Yao-Sung &amp; Hwang, Chung-Chi" docDate="2007" docId="EE05535497354343FF23FE76C564C0B7" docLanguage="en" docName="zt01608p068.pdf" docOrigin="Zootaxa 1608" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Satsuma longkiauwensis Wu, Lin &amp; Hwang, 2007, sp. nov." docType="treatment" docVersion="10" lastPageNumber="65" masterDocId="123C2B2C97374345FFB4FF8CC72FC53D" masterDocTitle="A new Satsuma species (Pulmonata: Camaenidae) endemic to Taiwan" masterLastPageNumber="68" masterPageNumber="59" pageNumber="61" updateTime="1698229628092" updateUser="plazi">
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<mods:title id="B482ED6B3F98B34F9B41A0F1E8344413">A new Satsuma species (Pulmonata: Camaenidae) endemic to Taiwan</mods:title>
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<mods:namePart id="A6288ECD50C7EA9A552ACF5DE606CBCF">Wu, Shu-Ping</mods:namePart>
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<mods:namePart id="DA0AD4CB3BDCD97DBA205DE2DCC6A1BA">Lin, Yao-Sung</mods:namePart>
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<mods:namePart id="235F300CF39115F805D20DA5A47E018B">Hwang, Chung-Chi</mods:namePart>
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<mods:title id="8E5E0B60379CD0FFF63B3759F7653A8B">Zootaxa</mods:title>
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<mods:part id="8492A29272CD586B88160ECF1F800E36">
<mods:date id="1341DC02434C8C8620E2986B54828EAA">2007</mods:date>
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<treatment id="EE05535497354343FF23FE76C564C0B7" ID-DOI="http://doi.org/10.5281/zenodo.6247439" ID-GBIF-Taxon="119374428" ID-Zenodo-Dep="6247439" LSID="urn:lsid:plazi:treatment:EE05535497354343FF23FE76C564C0B7" httpUri="http://treatment.plazi.org/id/EE05535497354343FF23FE76C564C0B7" lastPageId="6" lastPageNumber="65" pageId="2" pageNumber="61">
<subSubSection id="2EB6B1C997354347FF23FE76C637C701" pageId="2" pageNumber="61" type="nomenclature">
<paragraph id="6613E24297354347FF23FE76C509C729" blockId="2.[151,550,506,572]" box="[151,550,506,532]" pageId="2" pageNumber="61">
<heading id="3D5B552E97354347FF23FE76C509C729" bold="true" box="[151,550,506,532]" fontSize="11" level="1" pageId="2" pageNumber="61" reason="1">
<emphasis id="54D83E5097354347FF23FE76C509C729" bold="true" box="[151,550,506,532]" pageId="2" pageNumber="61">
<taxonomicName id="A1AC99C197354347FF23FE76C6EEC729" ID-CoL="7P54Z" box="[151,449,506,532]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="61" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097354347FF23FE76C6EEC729" bold="true" box="[151,449,506,532]" italics="true" pageId="2" pageNumber="61">Satsuma longkiauwensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FEB832B97354347FE7CFE76C509C729" box="[456,550,506,532]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6613E24297354347FF23FDAEC637C701" blockId="2.[151,550,506,572]" box="[151,280,546,572]" pageId="2" pageNumber="61">
(
<figureCitation id="FE97FEC797354347FF2BFDAEC620C701" box="[159,271,546,572]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 5" captionStartId-0="3.[151,255,1899,1923]" captionStartId-1="4.[151,255,1678,1702]" captionStartId-2="5.[151,255,1772,1796]" captionTargetBox-0="[215,1408,206,1867]" captionTargetBox-1="[311,1356,191,1632]" captionTargetBox-2="[371,1216,1175,1747]" captionTargetId-0="figure@3.[173,1443,168,1868]" captionTargetId-1="figure@4.[226,1360,142,1654]" captionTargetId-2="figure@5.[371,1217,1175,1748]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Satsuma longkiauwensis sp. nov. A – D. shell of holotype (TMMT 0701). Scale bar = 10 mm. A. Apertural view; B. Lateral view; C. Basal view; D. Top view; E. Living specimen (paratype TMMT 0704). Photograph by Chung- Yi Hsiao." captionText-1="FIGURE 3. Reproductive system of Satsuma longkiauwensis sp. nov. (paratype TMMT 0705) A. Whole genitalia, vas deferens is cut to unfold the genitalia; B. Verge and penial caecum; C. Interior of genitalia. Scale bar for A and C = 10 mm, scale bar for B = 1 mm." captionText-2="FIGURE 5. Jaw of Satsuma longkiauwensis sp. nov. (paratype, TMMT 0705). Scale bar = 600 µm." httpUri-0="https://zenodo.org/record/178847/files/figure.png" httpUri-1="https://zenodo.org/record/178848/files/figure.png" httpUri-2="https://zenodo.org/record/178849/files/figure.png" pageId="2" pageNumber="61">Figs. 2–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997354347FF23FDFEC6E1C6C9" pageId="2" pageNumber="61" type="materials_examined">
<paragraph id="6613E24297354347FF23FDFEC6E1C6C9" blockId="2.[151,1437,626,2024]" pageId="2" pageNumber="61">
<emphasis id="54D83E5097354347FF23FDFEC662C7B1" bold="true" box="[151,333,626,652]" pageId="2" pageNumber="61">
<typeStatus id="B9175CE097354347FF23FDFEC7FDC7B1" box="[151,210,626,652]" pageId="2" pageNumber="61">Type</typeStatus>
material.
</emphasis>
<emphasis id="54D83E5097354347FEECFDFEC6DFC7B6" box="[344,496,626,651]" italics="true" pageId="2" pageNumber="61">
<typeStatus id="B9175CE097354347FEECFDFEC6A0C7B6" box="[344,399,626,651]" pageId="2" pageNumber="61">Type</typeStatus>
locality
</emphasis>
: 
<collectingCountry id="1EBBA2D297354347FDB6FDFEC557C7B1" box="[514,632,626,652]" name="Taiwan" pageId="2" pageNumber="61">TAIWAN</collectingCountry>
: Pingtung County, Shuangliu, 
<geoCoordinate id="0398848597354347FC51FDFEC356C7B1" box="[997,1145,626,652]" direction="north" orientation="latitude" pageId="2" pageNumber="61" precision="1" value="22.218695">22º13’7.3”N</geoCoordinate>
<geoCoordinate id="0398848597354347FB37FDFEC21CC7B1" box="[1155,1331,626,652]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="1" value="120.793304">120º47’35.9”E</geoCoordinate>
, alt. 
<quantity id="A1544FA797354347FAC4FDFEC79EC789" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="61" unit="m" value="280.0">280 m</quantity>
; 
<collectionCode id="00BD7A8797354347FF0BFD16C638C789" box="[191,279,666,692]" pageId="2" pageNumber="61">TMMT</collectionCode>
0 701 (
<typeStatus id="B9175CE097354347FED9FD16C6F4C789" box="[365,475,666,692]" pageId="2" pageNumber="61" type="holotype">holotype</typeStatus>
): adult with decaying soft body in ethanol, collected by S.-P. Wu, 
<date id="1212C48297354347FB43FD16C2B8C789" box="[1271,1431,666,692]" pageId="2" pageNumber="61" value="2005-10-20">20 Oct. 2005</date>
; 
<collectionCode id="00BD7A8797354347FF23FD4EC7C0C7E1" box="[151,239,706,732]" pageId="2" pageNumber="61">TMMT</collectionCode>
0 705 (
<typeStatus id="B9175CE097354347FEF3FD4EC69BC7E1" box="[327,436,706,732]" pageId="2" pageNumber="61" type="paratype">paratype</typeStatus>
): adult dissected for genitalia, 
<taxonomicName id="A1AC99C197354347FC9CFD4EC45EC7E1" box="[808,881,706,732]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="2" pageNumber="64" phylum="Bryophyta" rank="genus">radula</taxonomicName>
and jaw, remaining soft parts in ethanol, collected by S.-P. Wu, 
<date id="1212C48297354347FECBFD66C50BC639" box="[383,548,746,772]" pageId="2" pageNumber="61" value="2005-09-13">13 Sept. 2005</date>
; 
<collectionCode id="00BD7A8797354347FD85FD66C5A6C639" box="[561,649,746,772]" pageId="2" pageNumber="61">TMMT</collectionCode>
0 704 (
<typeStatus id="B9175CE097354347FD6FFD66C468C639" box="[731,839,746,772]" pageId="2" pageNumber="61" type="paratype">paratype</typeStatus>
): sub-adult dissected for 
<taxonomicName id="A1AC99C197354347FBDFFD66C39BC639" box="[1131,1204,746,772]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="2" pageNumber="64" phylum="Bryophyta" rank="genus">radula</taxonomicName>
and jaw, remaining soft parts in ethanol, collected by S.-P. Wu, 
<date id="1212C48297354347FD27FC9EC418C611" box="[659,823,786,812]" pageId="2" pageNumber="61" value="2005-09-13">13 Sept. 2005</date>
; 
<collectionCode id="00BD7A8797354347FCF6FC9EC4B5C611" box="[834,922,786,812]" pageId="2" pageNumber="61">TMMT</collectionCode>
0 706 (
<typeStatus id="B9175CE097354347FC5EFC9EC378C611" box="[1002,1111,786,812]" pageId="2" pageNumber="61" type="paratype">paratype</typeStatus>
): sub-adult soft body in ethanol, collected by S.-P. Wu, 
<date id="1212C48297354347FE6BFCB6C5ACC669" box="[479,643,826,852]" pageId="2" pageNumber="61" value="2005-09-13">13 Sept. 2005</date>
; 
<collectionCode id="00BD7A8797354347FD3BFCB6C5C8C669" box="[655,743,826,852]" pageId="2" pageNumber="61">TMMT</collectionCode>
0 707 (
<typeStatus id="B9175CE097354347FC83FCB6C48BC669" box="[823,932,826,852]" pageId="2" pageNumber="61" type="paratype">paratype</typeStatus>
): adult shell, collected by S.-P. Wu, 
<date id="1212C48297354347FAF0FCB6C7FAC641" pageId="2" pageNumber="61" value="2004-02-21">21 Feb. 2004</date>
; 
<collectionCode id="00BD7A8797354347FF55FCEEC616C641" box="[225,313,866,892]" pageId="2" pageNumber="61">BMNH</collectionCode>
20060795 and 
<collectionCode id="00BD7A8797354347FE5AFCEEC516C641" LSID="urn:lsid:biocol.org:col:34784" box="[494,569,866,892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="2" pageNumber="61">ANSP</collectionCode>
413696 (
<typeStatus id="B9175CE097354347FD1CFCEEC40FC641" box="[680,800,866,892]" pageId="2" pageNumber="61" type="paratype">paratypes</typeStatus>
): adult shell, collected by S.-P. Wu, 
<date id="1212C48297354347FB71FCEEC24EC641" box="[1221,1377,866,892]" pageId="2" pageNumber="61" value="2005-10-20">20 Oct. 2005</date>
. 
<emphasis id="54D83E5097354347FF23FC06C612C69E" box="[151,317,906,931]" italics="true" pageId="2" pageNumber="61">Other locality</emphasis>
: 
<collectingCountry id="1EBBA2D297354347FEE4FC06C6E9C699" box="[336,454,906,932]" name="Taiwan" pageId="2" pageNumber="61">TAIWAN</collectingCountry>
: Pingtung County, Mudan, 
<geoCoordinate id="0398848597354347FCA6FC06C499C699" box="[786,950,906,932]" direction="north" orientation="latitude" pageId="2" pageNumber="61" precision="1" value="22.132055">22º07’55.4”N</geoCoordinate>
<geoCoordinate id="0398848597354347FC0BFC06C35EC699" box="[959,1137,906,932]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="1" value="120.788475">120º47’18.5”E</geoCoordinate>
, alt. 
<quantity id="A1544FA797354347FB1AFC06C3D0C699" box="[1198,1279,906,932]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="61" unit="m" value="225.0">225 m</quantity>
; empty adult shells: 
<collectionCode id="00BD7A8797354347FF5FFC3EC66FC6F1" box="[235,320,946,972]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="61">NMNS</collectionCode>
5415-001–003 (three 
<typeStatus id="B9175CE097354347FDF9FC3EC590C6F1" box="[589,703,946,972]" pageId="2" pageNumber="61" type="paratype">paratypes</typeStatus>
) and 
<collectionCode id="00BD7A8797354347FCB2FC3EC471C6F1" box="[774,862,946,972]" pageId="2" pageNumber="61">TMMT</collectionCode>
0702–0703 (two 
<typeStatus id="B9175CE097354347FB81FC3EC384C6F1" box="[1077,1195,946,972]" pageId="2" pageNumber="61" type="paratype">paratypes</typeStatus>
): collected by C.-C. Hwang, 
<date id="1212C48297354347FF4EFC56C6E5C6C9" box="[250,458,986,1012]" pageId="2" pageNumber="61" value="1997-08-26" valueMax="1997-08-27" valueMin="1997-08-26">26–27 Aug. 1997</date>
.
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997354347FF72FB8EC347C151" pageId="2" pageNumber="61" type="diagnosis">
<paragraph id="6613E24297354347FF72FB8EC347C151" blockId="2.[151,1437,626,2024]" pageId="2" pageNumber="61">
<emphasis id="54D83E5097354347FF72FB8EC669C121" bold="true" box="[198,326,1026,1052]" pageId="2" pageNumber="61">Diagnosis.</emphasis>
A large-sized, dextral 
<taxonomicName id="A1AC99C197354347FDE7FB8EC599C126" box="[595,694,1026,1051]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="61" phylum="Mollusca" rank="genus">
<emphasis id="54D83E5097354347FDE7FB8EC599C126" box="[595,694,1026,1051]" italics="true" pageId="2" pageNumber="61">Satsuma</emphasis>
</taxonomicName>
with light brown to olive background with depressed base and open umbilicus; Reproductive system with robust flagellum, swollen distal vagina, truncated oval verge, thick and muscular sheath tying male genitalia and expansive base of spermathecal stalk.
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997354347FF72FBF6C539C181" pageId="2" pageNumber="61" type="etymology">
<paragraph id="6613E24297354347FF72FBF6C539C181" blockId="2.[151,1437,626,2024]" pageId="2" pageNumber="61">
<emphasis id="54D83E5097354347FF72FBF6C67CC1A9" bold="true" box="[198,339,1146,1172]" pageId="2" pageNumber="61">Etymology.</emphasis>
Longkiauw (Langjiao) is the ancient name of nowadays Hengchun Peninsula and originates from the aboriginal race Paiwan.
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997354340FF72FB46C43CC15C" lastPageId="5" lastPageNumber="64" pageId="2" pageNumber="61" type="description">
<paragraph id="6613E24297354347FF72FB46C4F4C368" blockId="2.[151,1437,626,2024]" pageId="2" pageNumber="61">
<emphasis id="54D83E5097354347FF72FB46C670C1D9" bold="true" box="[198,351,1226,1252]" pageId="2" pageNumber="61">Description.</emphasis>
<emphasis id="54D83E5097354347FEDDFB47C68BC1D9" box="[361,420,1227,1252]" italics="true" pageId="2" pageNumber="61">Shell</emphasis>
(
<figureCitation id="FE97FEC797354347FE03FB46C52CC1D9" box="[439,515,1226,1252]" captionStart="FIGURE 2" captionStartId="3.[151,255,1899,1923]" captionTargetBox="[215,1408,206,1867]" captionTargetId="figure@3.[173,1443,168,1868]" captionTargetPageId="3" captionText="FIGURE 2. Satsuma longkiauwensis sp. nov. A – D. shell of holotype (TMMT 0701). Scale bar = 10 mm. A. Apertural view; B. Lateral view; C. Basal view; D. Top view; E. Living specimen (paratype TMMT 0704). Photograph by Chung- Yi Hsiao." httpUri="https://zenodo.org/record/178847/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
): large, dextral, semi-spherical, thick, rigid; height 
<quantity id="A1544FA797354347FBC5FB46C3D9C1D9" box="[1137,1270,1226,1252]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.3" metricValueMin="2.4" pageId="2" pageNumber="61" unit="mm" value="28.5" valueMax="33.0" valueMin="24.0">24–33 mm</quantity>
; width 
<quantity id="A1544FA797354347FAE6FB46C7E7C032" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.1" metricValueMin="3.4" pageId="2" pageNumber="61" unit="mm" value="37.5" valueMax="41.0" valueMin="34.0">34–41 mm</quantity>
; 5 7/8 –6 1/8 whorls. Apex obtuse. Whorls strongly inflated. Periphery bluntly angulated. Base of shell bulged outwards and flattened behind peristome; Surface of periostracum smooth, glossy, with faint oblique and spiral striations. Color of shell light brown to olive with red-brown stain. Peripheral band thin, sharp, redbrown. Surface above peripheral band brown, gradually lighter towards suture; area immediately below peripheral band yellowish white; basal band light brown colored, broad, gradually smear towards umbilicus. Umbilicus uncolored. Aperture diagonal, ovate. Peristome in lateral view concave. Lip thick, expanded; reflected at outer and inferior parts. Columellar lip oblique, straight; dilation of columellar lip not covering umbilicus. Lip color dark purple-red, glossy. Parietal callus thin, semitransparent. Umbilicus open, broad, deep. 
<typeStatus id="B9175CE097354347FF69F9B7C660C368" box="[221,335,1595,1621]" pageId="2" pageNumber="61" type="holotype">Holotype</typeStatus>
: height 
<quantity id="A1544FA797354347FE1EF9B7C6D0C368" box="[426,511,1595,1621]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="61" unit="mm" value="28.0">28 mm</quantity>
, width 
<quantity id="A1544FA797354347FDE1F9B7C585C368" box="[597,682,1595,1621]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="61" unit="mm" value="39.0">39 mm</quantity>
, 6 1/8 whorls (
<figureCitation id="FE97FEC797354347FCFFF9B7C4BEC368" box="[843,913,1595,1621]" captionStart="FIGURE 2" captionStartId="3.[151,255,1899,1923]" captionTargetBox="[215,1408,206,1867]" captionTargetId="figure@3.[173,1443,168,1868]" captionTargetPageId="3" captionText="FIGURE 2. Satsuma longkiauwensis sp. nov. A – D. shell of holotype (TMMT 0701). Scale bar = 10 mm. A. Apertural view; B. Lateral view; C. Basal view; D. Top view; E. Living specimen (paratype TMMT 0704). Photograph by Chung- Yi Hsiao." httpUri="https://zenodo.org/record/178847/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
A–D).
</paragraph>
<paragraph id="6613E24297354340FF72F9EAC56AC5E7" blockId="2.[151,1437,626,2024]" lastBlockId="5.[151,1437,152,418]" lastPageId="5" lastPageNumber="64" pageId="2" pageNumber="61">
<emphasis id="54D83E5097354347FF72F9EAC697C342" box="[198,440,1638,1663]" italics="true" pageId="2" pageNumber="61">Reproductive system</emphasis>
(
<figureCitation id="FE97FEC797354347FE7EF9EAC53CC3BD" box="[458,531,1638,1664]" captionStart="FIGURE 3" captionStartId="4.[151,255,1678,1702]" captionTargetBox="[311,1356,191,1632]" captionTargetId="figure@4.[226,1360,142,1654]" captionTargetPageId="4" captionText="FIGURE 3. Reproductive system of Satsuma longkiauwensis sp. nov. (paratype TMMT 0705) A. Whole genitalia, vas deferens is cut to unfold the genitalia; B. Verge and penial caecum; C. Interior of genitalia. Scale bar for A and C = 10 mm, scale bar for B = 1 mm." httpUri="https://zenodo.org/record/178848/files/figure.png" pageId="2" pageNumber="61">Fig. 3</figureCitation>
): Spermathecal stalk as long as spermoviduct, swollen at basal half, slender till spermatheca, with a constriction dividing base into two expansions of equal length; basal expansion bacillary, with ten to twelve strong, corrugated folds along inside; distal expansion shuttle-shaped, with seven thin, strong zigzag folds inside. Spermatheca oval. Free oviduct long, about same length as basal expansion of spermathecal stalk. Proximal vagina muscular, smooth externally, gross, weaker than basal expansion of spermathecal stalk. Internal folds in proximal vagina continued with those of spermathecal stalk counting nine in number, strongly elevated, lamellate, smooth, gradually weakened towards atrium. Subsequent part of vagina gradually slender towards a sudden swell distally, with ten thin, wiggly folds inside. Distal 1/5 of vagina abruptly thickened, muscular, depressed, pale brown in color, with wide, flattened folds inside. Male genitalia twisted, wrapped by a thin, semi-translucent, muscular sheath. Flagellum moderate, robust, tapering, with thickened base. Epiphallus as strong as base of flagellum with four to six unequally strong pilasters along inside, terminated with a shortly projecting, truncated oval, finely wrinkled verge entering penis. Penial caecum (appendix) not definite, with only an angled expansion surrounding verge. Pilasters around verge weak, wiggly, numbered 15–19. Proximal part of penis strong, thickened with 13 smooth, well-defined, strong pilasters. Subsequent part of penis slender with nine low, wiggly, corrugated pilasters inside. Distal part of penis extended again, about same length as distal swell of vagina, with weak, zigzag, corrugated pilasters internally. Atrium short, finely wrinkled inside.
</paragraph>
<caption id="32D3B2CA97344346FF23F8E7C7D4C2F1" httpUri="https://zenodo.org/record/178847/files/figure.png" pageId="3" pageNumber="62" targetBox="[215,1408,206,1867]" targetPageId="3">
<paragraph id="6613E24297344346FF23F8E7C7D4C2F1" blockId="3.[151,1436,1899,1996]" pageId="3" pageNumber="62">
<emphasis id="54D83E5097344346FF23F8E7C635C2BE" bold="true" box="[151,282,1899,1923]" pageId="3" pageNumber="62">FIGURE 2.</emphasis>
<taxonomicName id="A1AC99C197344346FE96F8E7C508C2BF" box="[290,551,1899,1922]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="62" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097344346FE96F8E7C508C2BF" box="[290,551,1899,1922]" italics="true" pageId="3" pageNumber="62">Satsuma longkiauwensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FEB832B97344346FD9BF8E7C5A9C2BE" box="[559,646,1899,1923]" pageId="3" pageNumber="62" rank="species">
<emphasis id="54D83E5097344346FD9BF8E7C5A9C2BE" bold="true" box="[559,646,1899,1923]" pageId="3" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
A–D. shell of holotype (TMMT 0701). Scale bar = 10 mm. A. Apertural view; B. Lateral view; C. Basal view; D. Top view; E. Living specimen (paratype TMMT 0704). Photograph by Chung- Yi Hsiao.
</paragraph>
</caption>
<caption id="32D3B2CA97334341FF23F902C6E4C3D7" httpUri="https://zenodo.org/record/178848/files/figure.png" pageId="4" pageNumber="63" targetBox="[311,1356,191,1632]" targetPageId="4">
<paragraph id="6613E24297334341FF23F902C6E4C3D7" blockId="4.[151,1436,1678,1770]" pageId="4" pageNumber="63">
<emphasis id="54D83E5097334341FF23F902C635C39B" bold="true" box="[151,282,1678,1702]" pageId="4" pageNumber="63">FIGURE 3.</emphasis>
Reproductive system of 
<taxonomicName id="A1AC99C197334341FD9CF902C401C398" box="[552,814,1678,1701]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="63" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097334341FD9CF902C401C398" box="[552,814,1678,1701]" italics="true" pageId="4" pageNumber="63">Satsuma longkiauwensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FEB832B97334341FC82F902C4A2C39B" box="[822,909,1678,1702]" pageId="4" pageNumber="63" rank="species">
<emphasis id="54D83E5097334341FC82F902C4A2C39B" bold="true" box="[822,909,1678,1702]" pageId="4" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
(paratype TMMT 0705) A. Whole genitalia, vas deferens is cut to unfold the genitalia; B. Verge and penial caecum; C. Interior of genitalia. Scale bar for A and C = 10 mm, scale bar for B = 1 mm.
</paragraph>
</caption>
<paragraph id="6613E24297324340FF72FF64C4C2C49F" blockId="5.[151,1437,152,418]" pageId="5" pageNumber="64">
<emphasis id="54D83E5097324340FF72FF64C6AFC43C" box="[198,384,232,257]" italics="true" pageId="5" pageNumber="64">
<taxonomicName id="A1AC99C197324340FF72FF64C634C43C" box="[198,283,232,257]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="5" pageNumber="64" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
and jaw
</emphasis>
: 
<taxonomicName id="A1AC99C197324340FE3BFF64C6CDC43F" box="[399,482,232,258]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="5" pageNumber="64" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
ribbon length 
<quantity id="A1544FA797324340FD3AFF64C5C4C43F" box="[654,747,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="5" pageNumber="64" unit="mm" value="7.6">7.6 mm</quantity>
, width 
<quantity id="A1544FA797324340FCF4FF64C4B1C43F" box="[832,926,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="64" unit="mm" value="2.6">2.6 mm</quantity>
. 
<taxonomicName id="A1AC99C197324340FC1CFF64C4D4C43F" box="[936,1019,232,258]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="5" pageNumber="64" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
formula 56 + C + 56. Central tooth with blunt, wide, curved mesocone and two weak, symmetrical ectocones (Fig. 4A). Lateral teeth with narrower, outwardly directed mesocone and weak endocone and ectocone (Fig. 4B). Marginal teeth with more outwardly directed mesocone and prominently forked, pointed endocone and ectocone (Fig. 4C). Jaw dark brown, curved with high, wide vertical ribs, reduced on margins (
<figureCitation id="FE97FEC797324340FC22FE04C4F2C49F" box="[918,989,392,418]" captionStart="FIGURE 5" captionStartId="5.[151,255,1772,1796]" captionTargetBox="[371,1216,1175,1747]" captionTargetId="figure@5.[371,1217,1175,1748]" captionTargetPageId="5" captionText="FIGURE 5. Jaw of Satsuma longkiauwensis sp. nov. (paratype, TMMT 0705). Scale bar = 600 µm." httpUri="https://zenodo.org/record/178849/files/figure.png" pageId="5" pageNumber="64">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="6613E24297324340FF23FBA9C43CC15C" blockId="5.[151,1436,1061,1121]" pageId="5" pageNumber="64">
<emphasis id="54D83E5097324340FF23FBA9C63AC100" bold="true" box="[151,277,1061,1085]" pageId="5" pageNumber="64">FIGURE 4</emphasis>
. 
<taxonomicName id="A1AC99C197324340FE91FBA9C65EC101" box="[293,369,1061,1084]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="5" pageNumber="64" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
of 
<taxonomicName id="A1AC99C197324340FE2EFBA9C58DC101" box="[410,674,1061,1084]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097324340FE2EFBA9C58DC101" box="[410,674,1061,1084]" italics="true" pageId="5" pageNumber="64">Satsuma longkiauwensis</emphasis>
</taxonomicName>
<emphasis id="54D83E5097324340FD18FBA9C42AC100" bold="true" box="[684,773,1061,1085]" pageId="5" pageNumber="64">
<taxonomicNameLabel id="4FEB832B97324340FD18FBA9C42AC100" box="[684,773,1061,1085]" pageId="5" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<typeStatus id="B9175CE097324340FCA1FBA9C45AC101" box="[789,885,1061,1084]" pageId="5" pageNumber="64" type="paratype">paratype</typeStatus>
, TMMT 0705). A. Central tooth; B. Lateral tooth (15th, left); C. Marginal tooth (45th, left). Scale bar = 20 µm.
</paragraph>
</subSubSection>
<caption id="32D3B2CA97324340FF23F960C3EEC23E" box="[151,1217,1772,1796]" httpUri="https://zenodo.org/record/178849/files/figure.png" pageId="5" pageNumber="64" targetBox="[371,1216,1175,1747]" targetPageId="5">
<paragraph id="6613E24297324340FF23F960C3EEC23E" blockId="5.[151,1217,1772,1796]" box="[151,1217,1772,1796]" pageId="5" pageNumber="64">
<emphasis id="54D83E5097324340FF23F960C63DC239" bold="true" box="[151,274,1772,1796]" pageId="5" pageNumber="64">FIGURE 5</emphasis>
. Jaw of 
<taxonomicName id="A1AC99C197324340FED9F961C55DC239" box="[365,626,1773,1796]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097324340FED9F961C55DC239" box="[365,626,1773,1796]" italics="true" pageId="5" pageNumber="64">Satsuma longkiauwensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FEB832B97324340FDCDF960C5E0C239" box="[633,719,1772,1796]" pageId="5" pageNumber="64" rank="species">
<emphasis id="54D83E5097324340FDCDF960C5E0C239" bold="true" box="[633,719,1772,1796]" pageId="5" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
(paratype, TMMT 0705). Scale bar = 600 µm.
</paragraph>
</caption>
<subSubSection id="2EB6B1C997324340FF72F8BBC571C2CC" pageId="5" pageNumber="64" type="distribution">
<paragraph id="6613E24297324340FF72F8BBC571C2CC" blockId="5.[151,1436,1847,2033]" pageId="5" pageNumber="64">
<emphasis id="54D83E5097324340FF72F8BBC670C26C" bold="true" box="[198,351,1847,1873]" pageId="5" pageNumber="64">Distribution</emphasis>
(
<figureCitation id="FE97FEC797324340FEC5F8BBC697C26C" box="[369,440,1847,1873]" captionStart="FIGURE 1" captionStartId="1.[151,255,1764,1788]" captionTargetBox="[195,1394,322,1730]" captionTargetId="figure@1.[186,1402,317,1737]" captionTargetPageId="1" captionText="FIGURE 1. Map of Taiwan showing the distribution of Satsuma longkiauwensis sp. nov. Solid asterisk, type locality; empty asterisk, locality of paratypes NMNS 5415 - 001 – 003 and TMMT 0702 – 0703; grey area around asterisks, distribution of Satsuma longkiauwensis; bold black line, Central Range; thin line, administrative boundary." httpUri="https://zenodo.org/record/178846/files/figure.png" pageId="5" pageNumber="64">Fig. 1</figureCitation>
): All specimens were collected in mountainous lowland between the river basin of the rivers Fengangxi and Sichungxi. Living individuals were collected only in Shouangliu, the 
<typeStatus id="B9175CE097324340FB5BF8D3C20EC244" box="[1263,1313,1887,1913]" pageId="5" pageNumber="64">type</typeStatus>
locality. 
<emphasis id="54D83E5097324340FF72F80BC61CC29C" bold="true" box="[198,307,1927,1953]" pageId="5" pageNumber="64">Ecology:</emphasis>
Live specimens were observed active in the field in September. They were nocturnal and herbivorous feeding on decaying fallen leaves and gramineous shoots (
<figureCitation id="FE97FEC797324340FC16F823C4C4C2F4" box="[930,1003,1967,1993]" captionStart="FIGURE 2" captionStartId="3.[151,255,1899,1923]" captionTargetBox="[215,1408,206,1867]" captionTargetId="figure@3.[173,1443,168,1868]" captionTargetPageId="3" captionText="FIGURE 2. Satsuma longkiauwensis sp. nov. A – D. shell of holotype (TMMT 0701). Scale bar = 10 mm. A. Apertural view; B. Lateral view; C. Basal view; D. Top view; E. Living specimen (paratype TMMT 0704). Photograph by Chung- Yi Hsiao." httpUri="https://zenodo.org/record/178847/files/figure.png" pageId="5" pageNumber="64">Fig. 2</figureCitation>
E). They were found on ground and crawling on rocks or lower tree trunks.
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997314343FF72FF1BC417C787" pageId="6" pageNumber="65" type="discussion">
<paragraph id="6613E24297314343FF72FF1BC7C2C77F" blockId="6.[151,1437,151,1418]" pageId="6" pageNumber="65">
<emphasis id="54D83E5097314343FF72FF1BC66DC58C" bold="true" box="[198,322,151,177]" pageId="6" pageNumber="65">Remarks:</emphasis>
The reproductive system of 
<taxonomicName id="A1AC99C197314343FD15FF14C4ECC58C" box="[673,963,152,177]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097314343FD15FF14C4ECC58C" box="[673,963,152,177]" italics="true" pageId="6" pageNumber="65">Satsuma longkiauwensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FEB832B97314343FC79FF1BC302C58C" box="[973,1069,151,177]" pageId="6" pageNumber="65" rank="species">
<emphasis id="54D83E5097314343FC79FF1BC302C58C" bold="true" box="[973,1069,151,177]" pageId="6" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from that of the largesized 
<taxonomicName id="A1AC99C197314343FF69FF4CC66EC5E4" box="[221,321,192,217]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="54D83E5097314343FF69FF4CC66EC5E4" box="[221,321,192,217]" italics="true" pageId="6" pageNumber="65">Satsuma</emphasis>
</taxonomicName>
species from 
<collectingCountry id="1EBBA2D297314343FE5FFF4CC56EC5E7" box="[491,577,192,218]" name="Taiwan" pageId="6" pageNumber="65">Taiwan</collectingCountry>
and 
<collectingCountry id="1EBBA2D297314343FDCBFF4CC5EAC5E7" box="[639,709,192,218]" name="Japan" pageId="6" pageNumber="65">Japan</collectingCountry>
, e.g. 
<taxonomicName id="A1AC99C197314343FCB3FF4CC37CC5E7" authority="H. Adams, 1866" authorityName="H. Adams" authorityYear="1866" box="[775,1107,192,218]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="bairdi">
<emphasis id="54D83E5097314343FCB3FF4CC441C5E4" box="[775,878,192,217]" italics="true" pageId="6" pageNumber="65">S. bairdi</emphasis>
(
<bibRefCitation id="023D9FB397314343FC35FF4CC365C5E7" author="Adams" box="[897,1098,192,218]" pageId="6" pageNumber="67" refString="Adams, H. (1866) Descriptions of fifteen new species land and freshwater shells from Formosa, collected by Robert Swinhoe, Esq., consul at Taiwan in that island. Proceedings of the Zoological Society, London, 1866, 316 - 319." type="proceedings paper" year="1866">H. Adams, 1866</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A1AC99C197314343FBD5FF4CC2B2C5E7" authority="Kuroda, 1941" authorityName="Kuroda" authorityYear="1941" box="[1121,1437,192,218]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="arisana">
<emphasis id="54D83E5097314343FBD5FF4CC3F5C5E4" box="[1121,1242,192,217]" italics="true" pageId="6" pageNumber="65">S. arisana</emphasis>
(
<bibRefCitation id="023D9FB397314343FB59FF4CC2BBC5E7" author="Kuroda" box="[1261,1428,192,218]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
ssp., 
<taxonomicName id="A1AC99C197314343FF67FF64C531C43F" authority="Moellendorff, 1888" authorityName="Moellendorff" authorityYear="1888" box="[211,542,232,258]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="nux">
<emphasis id="54D83E5097314343FF67FF64C635C43C" box="[211,282,232,257]" italics="true" pageId="6" pageNumber="65">S. nux</emphasis>
(
<bibRefCitation id="023D9FB397314343FE9EFF64C53BC43F" author="Moellendorff" box="[298,532,232,258]" pageId="6" pageNumber="67" refString="Moellendorff, O. F. (1888) Mittheilungen aus dem Gebiete der Malakozoologie. Diagnoses specierum novarum sinensium. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 20, 38 - 44." type="journal article" year="1888">Moellendorff, 1888</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A1AC99C197314343FD9EFF64C48AC43F" authority="Pfeiffer, 1866" authorityName="Pfeiffer" authorityYear="1866" box="[554,933,232,258]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="sphaeroconus">
<emphasis id="54D83E5097314343FD9EFF64C5C6C43C" box="[554,745,232,257]" italics="true" pageId="6" pageNumber="65">S. sphaeroconus</emphasis>
(
<bibRefCitation id="023D9FB397314343FD4CFF64C4B2C43F" author="Pfeiffer" box="[760,925,232,258]" pageId="6" pageNumber="67" refString="Pfeiffer, L. (1866) Descriptions of thirteen new species of land-shells from Formosa, in the collection of the late Hugh Cuming, collected by Mr. Robert Swinhoe, vice-consul of that island. Proceedings of the Zoological Society, London [1865], 828 - 831." type="proceedings paper" year="1866">Pfeiffer, 1866</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="A1AC99C197314343FC6BFF64C212C43F" authority="Pfeiffer, 1845" authorityName="Pfeiffer" authorityYear="1845" box="[991,1341,232,258]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="mercatoria">
<emphasis id="54D83E5097314343FC6BFF64C351C43C" box="[991,1150,232,257]" italics="true" pageId="6" pageNumber="65">S. mercatoria</emphasis>
(
<bibRefCitation id="023D9FB397314343FB39FF64C21BC43F" author="Pfeiffer" box="[1165,1332,232,258]" pageId="6" pageNumber="67" refString="Pfeiffer, L. (1845) Diagnosen einiger neuer Heliceen. Zeitschrift fur Malakozoologie, 2, 152 - 158." type="journal article" year="1845">Pfeiffer, 1845</bibRefCitation>
)
</taxonomicName>
, in having a weak expansion instead of an elongated penial caecum externally and a hemispherical verge instead of elongated and thickened pilasters internally (
<bibRefCitation id="023D9FB397314343FD14FEB4C413C46F" author="Minato" box="[672,828,312,338]" pageId="6" pageNumber="67" refString="Minato, H. (1975) Systematic position of &quot; Coniglobus &quot; sphaeroconus (Pfeiffer, 1865) from Formosa (Pulmonata: Camaenidae). Venus, 34, 99 - 104." type="journal article" year="1975">Minato 1975</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FCF3FEB4C309C46F" author="Wu" box="[839,1062,312,338]" pageId="6" pageNumber="68" refString="Wu, W. K. &amp; Chang, K. M. (1975) Anatomy of the land snail Ganesella (Coniglobus) nux (Moellendorff). Bulletin of Malacology, 2, 107 - 111." type="journal article" year="1975">Wu &amp; Chang 1975</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FB86FEB4C26BC46F" author="Azuma" box="[1074,1348,312,338]" pageId="6" pageNumber="67" refString="Azuma, M. &amp; Azuma, Y. (1987) Land molluscan fauna of Miyako Islands, the southwestern Okinawa, Japan, with descriptions of new taxa - I. Venus, 46, 159 - 165." type="journal article" year="1987">Azuma &amp; Azuma 1987</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FAE4FEB4C7FBC447" author="Chang" pageId="6" pageNumber="67" refString="Chang, K. M. (1994) Anatomy of Satsuma bairdi (H. Adams, 1866) from Taiwan (Pulmonata). Venus, 53, 319 - 325." type="journal article" year="1994">Chang 1994</bibRefCitation>
, 
<bibRefCitation id="023D9FB397314343FF55FEECC60FC447" author="Chang" box="[225,288,352,378]" pageId="6" pageNumber="67" refString="Chang, K. M. (1997) Anatomy of Coniglobus arisanus (Kuroda) from Taiwan (Pulmonata: Camaenidae). Bulletin of Malacology, 21, 41 - 48." type="journal article" year="1997">1997</bibRefCitation>
). The reproductive system of the new species is similar to that of 
<taxonomicName id="A1AC99C197314343FB9FFEECC2B7C447" authority="H. Adams, 1866" authorityName="H. Adams" authorityYear="1866" box="[1067,1432,352,378]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="succincta">
<emphasis id="54D83E5097314343FB9FFEECC397C444" box="[1067,1208,352,377]" italics="true" pageId="6" pageNumber="65">S. succincta</emphasis>
(
<bibRefCitation id="023D9FB397314343FB7DFEECC2A1C447" author="Adams" box="[1225,1422,352,378]" pageId="6" pageNumber="67" refString="Adams, H. (1866) Descriptions of fifteen new species land and freshwater shells from Formosa, collected by Robert Swinhoe, Esq., consul at Taiwan in that island. Proceedings of the Zoological Society, London, 1866, 316 - 319." type="proceedings paper" year="1866">H. Adams, 1866</bibRefCitation>
)
</taxonomicName>
, 
<emphasis id="54D83E5097314343FF23FE04C66AC49C" box="[151,325,392,417]" italics="true" pageId="6" pageNumber="65">Pancala bacca</emphasis>
(
<bibRefCitation id="023D9FB397314343FEE0FE04C6D7C49F" author="Pfeiffer" box="[340,504,392,418]" pageId="6" pageNumber="67" refString="Pfeiffer, L. (1866) Descriptions of thirteen new species of land-shells from Formosa, in the collection of the late Hugh Cuming, collected by Mr. Robert Swinhoe, vice-consul of that island. Proceedings of the Zoological Society, London [1865], 828 - 831." type="proceedings paper" year="1866">Pfeiffer, 1866</bibRefCitation>
) and 
<emphasis id="54D83E5097314343FD8DFE04C40EC49C" box="[569,801,392,417]" italics="true" pageId="6" pageNumber="65">P. batanica pancala</emphasis>
(
<bibRefCitation id="023D9FB397314343FC84FE04C3A7C49F" author="Schmacker" box="[816,1160,392,418]" pageId="6" pageNumber="68" refString="Schmacker, B. &amp; Boettger, O. (1891) Neue Materialien zur Charakteristik und geographischen Verbreitung chinesischer und japanischer Binnenmollusken II. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 23, 145 - 194." type="journal article" year="1891">Schmacker &amp; Boettger, 1891</bibRefCitation>
) (
<bibRefCitation id="023D9FB397314343FB2AFE04C231C49F" author="Tabe" box="[1182,1310,392,418]" pageId="6" pageNumber="68" refString="Tabe, M. (1937) Morphological study of Coniglobus batanicus pancalus (Schmacker &amp; Boettger) and C. b. boteltobagoensis Kuroda and on the systematic position of the genus Coniglobus. Zoological Magazine, 49, 12 - 20." type="journal article" year="1937">Tabe 1937</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FA9EFE04C7FAC4F7" author="Sinagawa" pageId="6" pageNumber="68" refString="Sinagawa, K. (1979) Anatomy of Camaena cicatricosa and Pancala bacca and Pancala batanica pancala. Kainakama, 13, 4 - 7." type="journal article" year="1979">Sinagawa 1979</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FF56FE3CC65BC4F7" author="Chang" box="[226,372,432,458]" pageId="6" pageNumber="67" refString="Chang, K. M. (1992) Dwarf type of Camaena batanica pancala (Schmacker and Boettger) from South Cape, Taiwan and its allied species (Pulmonata: Camaenidae). Bulletin of Malacology, 17, 11 - 20." type="journal article" year="1992">Chang 1992</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FE35FE3CC533C4F7" author="Hwang" box="[385,540,432,458]" pageId="6" pageNumber="67" refString="Hwang, C. C. (1995) Analysis of mitochondrial DNA phylogeny of Camaena (Pulmonata: Camaenidae). Unpublished Master thesis, National Sun Yat-sen University, Kaohsiung, Taiwan, 101 pp." type="book" year="1995">Hwang 1995</bibRefCitation>
), but differs from 
<emphasis id="54D83E5097314343FD40FE3CC477C4F4" box="[756,856,432,457]" italics="true" pageId="6" pageNumber="65">P. bacca</emphasis>
and 
<emphasis id="54D83E5097314343FC27FE3CC352C4F4" box="[915,1149,432,457]" italics="true" pageId="6" pageNumber="65">P. batanica pancala</emphasis>
in having a larger shell, roundly angulated peripheries adjacent to the peristome, an angled and less swollen junction between epiphallus and penis, robust and non-twisted flagellum, a more dilated penis at the proximal part and a swollen distal vagina.
</paragraph>
<paragraph id="6613E24297314343FF72FDDCC417C787" blockId="6.[151,1437,151,1418]" pageId="6" pageNumber="65">
Shell morphology of the new species differs from other camaenid species distributed in Hengchun Peninsula, as 
<emphasis id="54D83E5097314343FF41FDF4C676C7AC" box="[245,345,632,657]" italics="true" pageId="6" pageNumber="65">P. bacca</emphasis>
, 
<emphasis id="54D83E5097314343FEDCFDF4C57DC7AC" box="[360,594,632,657]" italics="true" pageId="6" pageNumber="65">P. batanica pancala</emphasis>
, 
<taxonomicName id="A1AC99C197314343FDD5FDF4C48DC7AF" authority="Rolle, 1911" authorityName="Rolle" authorityYear="1911" box="[609,930,632,658]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="pekanensis">
<emphasis id="54D83E5097314343FDD5FDF4C42FC7AC" box="[609,768,632,657]" italics="true" pageId="6" pageNumber="65">S. pekanensis</emphasis>
(
<bibRefCitation id="023D9FB397314343FCA4FDF4C4B4C7AF" author="Rolle" box="[784,923,632,658]" pageId="6" pageNumber="68" refString="Rolle, H. (1911) Diagnosen neuer Arten. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 43, 30 - 32." type="journal article" year="1911">Rolle, 1911</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="A1AC99C197314343FC69FDF4C2B8C7AF" authority="Pilsbry &amp; Hirase, 1909" authorityName="Pilsbry &amp; Hirase" authorityYear="1909" box="[989,1431,632,658]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="contraria">
<emphasis id="54D83E5097314343FC69FDF4C345C7AC" box="[989,1130,632,657]" italics="true" pageId="6" pageNumber="65">S. contraria</emphasis>
(
<bibRefCitation id="023D9FB397314343FBCDFDF4C2A1C7AF" author="Pilsbry" box="[1145,1422,632,658]" pageId="6" pageNumber="67" refString="Pilsbry, H. A. &amp; Hirase, Y. (1909) New land Mollusca of the Japanese empire. Proceedings of the Academy of Natural Sciences of Philadelphia, 60, 586 - 599, [1908]." type="journal article" year="1909">Pilsbry &amp; Hirase, 1909</bibRefCitation>
)
</taxonomicName>
, in having a larger and/or dextral shell (
<bibRefCitation id="023D9FB397314343FDE9FD2CC404C787" author="Hsieh" box="[605,811,672,698]" pageId="6" pageNumber="67" refString="Hsieh, B. C., Hwang, C. C. &amp; Wu, S. P. (2006) Landsnails of Taiwan. Forestry Bureau Council of Agriculture, Executive Yuan, Taipei, Taiwan, 277 pp." type="book" year="2006">
Hsieh 
<emphasis id="54D83E5097314343FD1EFD2CC5CAC784" box="[682,741,672,697]" italics="true" pageId="6" pageNumber="65">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997314343FF73FD44C5F1C1FF" pageId="6" pageNumber="65" type="reference_group">
<paragraph id="6613E24297314343FF73FD44C40BC6BF" blockId="6.[151,1437,151,1418]" pageId="6" pageNumber="65">
<taxonomicName id="A1AC99C197314343FF73FD44C5AAC7DF" authority="Kuroda, 1941" authorityName="Kuroda" authorityYear="1941" box="[199,645,712,738]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="arisana" subSpecies="tani">
<emphasis id="54D83E5097314343FF73FD44C6EBC7DC" box="[199,452,712,737]" italics="true" pageId="6" pageNumber="65">Satsuma arisana tani</emphasis>
(
<bibRefCitation id="023D9FB397314343FE60FD44C553C7DF" author="Kuroda" box="[468,636,712,738]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
, which is distributed in southeastern 
<collectingCountry id="1EBBA2D297314343FBF6FD44C3B4C7DF" box="[1090,1179,712,738]" name="Taiwan" pageId="6" pageNumber="65">Taiwan</collectingCountry>
, is also similar to the new species in shell dimensions (
<typeStatus id="B9175CE097314343FD9FFD7CC5B7C637" box="[555,664,752,778]" pageId="6" pageNumber="65" type="holotype">holotype</typeStatus>
: height 30.0 mm, width 41.0 mm) and shape. Unfortunately the 
<typeStatus id="B9175CE097314343FF23FC94C7E5C60F" box="[151,202,792,818]" pageId="6" pageNumber="65">type</typeStatus>
specimens were not traceable currently. However, it can be deduced from the original description and photographs of the 
<typeStatus id="B9175CE097314343FECBFCCCC6C9C667" box="[383,486,832,858]" pageId="6" pageNumber="65" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="023D9FB397314343FE42FCCCC5BAC667" author="Kuroda" box="[502,661,832,858]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda 1941</bibRefCitation>
) that the umbilicus of 
<taxonomicName id="A1AC99C197314343FC12FCCCC379C664" box="[934,1110,832,857]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="arisana" subSpecies="tani">
<emphasis id="54D83E5097314343FC12FCCCC379C664" box="[934,1110,832,857]" italics="true" pageId="6" pageNumber="65">S. arisana tani</emphasis>
</taxonomicName>
is perfectly covered by the reflected superior columellar lip and completely closed.
</paragraph>
<paragraph id="6613E24297314343FF73FC1CC5F1C1FF" blockId="6.[151,1437,151,1418]" pageId="6" pageNumber="65">
Among camaenid species from 
<collectingCountry id="1EBBA2D297314343FDF6FC1CC5B4C697" box="[578,667,912,938]" name="Taiwan" pageId="6" pageNumber="65">Taiwan</collectingCountry>
, 
<taxonomicName id="A1AC99C197314343FD1CFC1CC4ABC694" box="[680,900,912,937]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="succincta">
<emphasis id="54D83E5097314343FD1CFC1CC4ABC694" box="[680,900,912,937]" italics="true" pageId="6" pageNumber="65">Satsuma succincta</emphasis>
</taxonomicName>
shows a high similarity in shell morphology and banding patterns. Nevertheless, this species is smaller (shell height 
<quantity id="A1544FA797314343FC57FC34C3BEC6EF" box="[995,1169,952,978]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.28" metricValueMin="1.6199999999999999" pageId="6" pageNumber="65" unit="mm" value="19.5" valueMax="22.8" valueMin="16.2">16.2–22.8 mm</quantity>
, shell width 23.0– 
<quantity id="A1544FA797314343FADDFC34C7E7C6C7" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="6" pageNumber="65" unit="mm" value="31.2">31.2 mm</quantity>
, 6 
<typeStatus id="B9175CE097314343FF5DFC6CC67CC6C7" box="[233,339,992,1018]" pageId="6" pageNumber="65" type="syntype">syntypes</typeStatus>
, BMNH 1866.5.9.9) in shell dimensions than the new species. Both species are ground-living in the low land forests of south 
<collectingCountry id="1EBBA2D297314343FDB9FB84C549C11F" box="[525,614,1032,1058]" name="Taiwan" pageId="6" pageNumber="65">Taiwan</collectingCountry>
, but not sympatric. The smoothly rounded junction between the outer and inferior lips, yellowish coloration of shell, the robust and smoothly curved flagellum, and the swollen distal vagina of 
<taxonomicName id="A1AC99C197314343FE9BFBD4C52EC14C" box="[303,513,1112,1137]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="longkiauwensis" status="sp. nov.">
<emphasis id="54D83E5097314343FE9BFBD4C52EC14C" box="[303,513,1112,1137]" italics="true" pageId="6" pageNumber="65">S. longkiauwensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FEB832B97314343FDB3FBDBC54BC14C" box="[519,612,1111,1137]" pageId="6" pageNumber="65" rank="species">
<emphasis id="54D83E5097314343FDB3FBDBC54BC14C" bold="true" box="[519,612,1111,1137]" pageId="6" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
distinguish it further from 
<taxonomicName id="A1AC99C197314343FC14FBD4C304C14C" box="[928,1067,1112,1137]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="succincta">
<emphasis id="54D83E5097314343FC14FBD4C304C14C" box="[928,1067,1112,1137]" italics="true" pageId="6" pageNumber="65">S. succincta</emphasis>
</taxonomicName>
. 
<taxonomicName id="A1AC99C197314343FB8EFBD4C22BC14C" box="[1082,1284,1112,1137]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="succinct">
<emphasis id="54D83E5097314343FB8EFBD4C22BC14C" box="[1082,1284,1112,1137]" italics="true" pageId="6" pageNumber="65">Satsuma succinct</emphasis>
</taxonomicName>
and its allies show an angulated junction at peristome, reddish coloration of shell mostly, slender and wiggly flagellum and slender distal vagina (
<bibRefCitation id="023D9FB397314343FE23FB24C505C1FF" author="Chang" box="[407,554,1192,1218]" pageId="6" pageNumber="67" refString="Chang, K. M. (1989) Anatomy of Coniglobus nux paiwanis (Kuroda) and Coniglobus pekanensis (Rolle) from south Taiwan (Pulmonata: Camaenidae). Bulletin of Malacology, 14, 1 - 8." type="journal article" year="1989">Chang 1989</bibRefCitation>
; 
<bibRefCitation id="023D9FB397314343FD83FB24C5FEC1FF" author="Hwang" box="[567,721,1192,1218]" pageId="6" pageNumber="67" refString="Hwang, C. C. (1995) Analysis of mitochondrial DNA phylogeny of Camaena (Pulmonata: Camaenidae). Unpublished Master thesis, National Sun Yat-sen University, Kaohsiung, Taiwan, 101 pp." type="book" year="1995">Hwang 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2EB6B1C997314343FF72FB5CC564C0B7" pageId="6" pageNumber="65" type="materials_examined">
<paragraph id="6613E24297314343FF72FB5CC564C0B7" blockId="6.[151,1437,151,1418]" pageId="6" pageNumber="65">
All the 26 valid (sub-)species of 
<taxonomicName id="A1AC99C197314343FDE2FB5CC5C8C1D7" box="[598,743,1232,1258]" class="Gastropoda" family="Camaenidae" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="family">Camaenidae</taxonomicName>
from 
<collectingCountry id="1EBBA2D297314343FC80FB5CC4A5C1D7" box="[820,906,1232,1258]" name="Taiwan" pageId="6" pageNumber="65">Taiwan</collectingCountry>
can be separated using the key below. This identification key is based on shell characters of 
<typeStatus id="B9175CE097314343FD6FFB74C422C02F" box="[731,781,1272,1298]" pageId="6" pageNumber="65">type</typeStatus>
specimens examined except for 
<taxonomicName id="A1AC99C197314343FB26FB74C6BBC007" authority="Kuroda, 1941" authorityName="Kuroda" authorityYear="1941" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="arisana" subSpecies="takahasii">
<emphasis id="54D83E5097314343FB26FB74C7FFC004" italics="true" pageId="6" pageNumber="65">Satsuma arisana takahasii</emphasis>
(
<bibRefCitation id="023D9FB397314343FF56FAACC6A4C007" author="Kuroda" box="[226,395,1312,1338]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A1AC99C197314343FE17FAACC435C007" authority="Kuroda, 1941" authorityName="Kuroda" authorityYear="1941" box="[419,794,1312,1338]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="arisana" subSpecies="tani">
<emphasis id="54D83E5097314343FE17FAACC57AC004" box="[419,597,1312,1337]" italics="true" pageId="6" pageNumber="65">S. arisana tani</emphasis>
(
<bibRefCitation id="023D9FB397314343FDD3FAACC43EC007" author="Kuroda" box="[615,785,1312,1338]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A1AC99C197314343FC9DFAACC385C007" authority="Kuroda, 1941" authorityName="Kuroda" authorityYear="1941" box="[809,1194,1312,1338]" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="nux" subSpecies="paiwanis">
<emphasis id="54D83E5097314343FC9DFAACC4C8C004" box="[809,999,1312,1337]" italics="true" pageId="6" pageNumber="65">S. nux paiwanis</emphasis>
(
<bibRefCitation id="023D9FB397314343FC4EFAACC38EC007" author="Kuroda" box="[1018,1185,1312,1338]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="A1AC99C197314343FB5CFAACC660C05F" authority="Kuroda, 1941" authorityName="Kuroda" authorityYear="1941" class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="nux" subSpecies="sericata">
<emphasis id="54D83E5097314343FB5CFAACC2B4C004" box="[1256,1435,1312,1337]" italics="true" pageId="6" pageNumber="65">S. nux sericata</emphasis>
(
<bibRefCitation id="023D9FB397314343FF2BFAC4C669C05F" author="Kuroda" box="[159,326,1352,1378]" pageId="6" pageNumber="67" refString="Kuroda, T. (1941) A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 22, 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
. The 
<typeStatus id="B9175CE097314343FE3AFAC4C6EEC05F" box="[398,449,1352,1378]" pageId="6" pageNumber="65">type</typeStatus>
material of these four taxa could not be located. Anatomical characters are not used, as those are not available for all taxa.
</paragraph>
</subSubSection>
</treatment>
</document>