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269 lines
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<mods:titleInfo id="BD99ED8F4323BB454489F8B9D9353ADE">
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<mods:title id="14227869EBECCD3FCD68BA5AC128470E">A revision of Chrysopidae (Neuroptera) from the late Eocene Florissant Formation Colorado, with description of new species</mods:title>
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<mods:name id="8547F4C4586D270BD27A95FE85D36D09" type="personal">
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<mods:namePart id="FE257D2E9C58E29E5CA658A90762CC2C">Makarkin, Vladimir N.</mods:namePart>
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<mods:affiliation id="410853F9B27100CC43F610C8241B8D7E">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia</mods:affiliation>
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<mods:namePart id="496D0DAFEBC805A5C2F20C2DAB238A87">Antell, Gwen S.</mods:namePart>
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<mods:affiliation id="C99D1C4DB1CCF87A663438047705AC39">Department of Earth Sciences, University of Oxford, Oxford OX 1 3 AN, United Kingdom</mods:affiliation>
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<mods:namePart id="F25AF4835DBAC20077FFEB763C0EC592">Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier id="FBD18C1CD5AFE0E152AC2D14313EC9F3" type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation id="442B6AE173565BDD5DAF5D35AD4A956A">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada sba 48 @ sfu. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier id="6BE420F805179C123AC199F42503BC0A" type="email">sba48@sfu.ca</mods:nameIdentifier>
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<mods:title id="88D1847B00731968376A9347D3F4E1CE">Zootaxa</mods:title>
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<mods:part id="89CD94D1C53B7FDD440579DE6AD911B8">
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<mods:date id="924C5B7EE38553F3FDB5808287A03585">2022</mods:date>
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<mods:number id="611AA5154C8FE92EBE85E10AA4B9BFCD">2022-05-05</mods:number>
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<mods:identifier id="8E589BEF79AD29FF9ED4F9A6DFF39BC3" type="GBIF-Dataset">dd1981ca-6b0b-42cf-839b-f1d3ba251c73</mods:identifier>
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<treatment id="AD76878EFFF5FFFEE292A6CCCCDE752B" ID-DOI="http://doi.org/10.5281/zenodo.6521451" ID-GBIF-Taxon="195192045" ID-Zenodo-Dep="6521451" LSID="urn:lsid:plazi:treatment:AD76878EFFF5FFFEE292A6CCCCDE752B" httpUri="http://treatment.plazi.org/id/AD76878EFFF5FFFEE292A6CCCCDE752B" lastPageId="28" lastPageNumber="328" pageId="27" pageNumber="328">
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<subSubSection id="6DC56513FFF5FFF9E292A6CCCCD570AE" box="[151,661,1194,1221]" pageId="27" pageNumber="328" type="nomenclature">
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<paragraph id="25603698FFF5FFF9E292A6CCCCD570AE" blockId="27.[151,661,1194,1257]" box="[151,661,1194,1221]" pageId="27" pageNumber="328">
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<heading id="7E2881F4FFF5FFF9E292A6CCCCD570AE" bold="true" box="[151,661,1194,1221]" fontSize="11" level="1" pageId="27" pageNumber="328" reason="1">
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<taxonomicName id="E2DF4D1BFFF5FFF9E292A6CCCCD570AE" ID-CoL="6QKYP" authority="(Cockerell, 1909)" baseAuthorityName="Cockerell" baseAuthorityYear="1909" box="[151,661,1194,1221]" class="Insecta" family="Chrysopidae" genus="Lithochrysa" kingdom="Animalia" order="Neuroptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="concinnula">
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<emphasis id="17ABEA8AFFF5FFF9E292A6CCCCD570AE" bold="true" box="[151,661,1194,1221]" pageId="27" pageNumber="328">
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<emphasis id="17ABEA8AFFF5FFF9E292A6CCCFF970AF" bold="true" box="[151,441,1194,1221]" italics="true" pageId="27" pageNumber="328">Lithochrysa concinnula</emphasis>
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(Cockerell, 1909)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6DC56513FFF5FFF9E292A6A9CEA77083" box="[151,231,1231,1257]" pageId="27" pageNumber="328" type="description">
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<paragraph id="25603698FFF5FFF9E292A6A9CEA77083" blockId="27.[151,661,1194,1257]" box="[151,231,1231,1257]" pageId="27" pageNumber="328">
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<figureCitation id="BDE42A1DFFF5FFF9E292A6A9CEA77083" box="[151,231,1231,1257]" captionStart="FIGURE 20" captionStartId="28.[151,250,1876,1900]" captionTargetBox="[151,1436,345,1845]" captionTargetId="figure-97@28.[151,1436,345,1853]" captionTargetPageId="28" captionText="FIGURE 20. Lithochrysa concinnula (Cockerell, 1909), holotype UCM 4220. A, specimen in current condition, obscured by Canada balsam; B, forewing venation; C, hind wing venation (redrawn from Cockerell 1909b: Fig. 5). Scale bar = 3 mm (B, C not to scale, no scale on original)." figureDoi="http://doi.org/10.5281/zenodo.6522503" httpUri="https://zenodo.org/record/6522503/files/figure.png" pageId="27" pageNumber="328">Fig. 20</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6DC56513FFF5FFF9E292A775CA5C71CC" pageId="27" pageNumber="328" type="reference_group">
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<paragraph id="25603698FFF5FFF9E292A775CAD47103" blockId="27.[151,1437,1299,1446]" pageId="27" pageNumber="328">
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<treatmentCitationGroup id="05CF11B6FFF5FFF9E292A775CAD47103" pageId="27" pageNumber="328">
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<treatmentCitation id="A47E1089FFF5FFF9E292A775CCCE7141" author="Cockerell, T. D. A." box="[151,654,1299,1323]" page="218" pageId="27" pageNumber="328" year="1909">
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<taxonomicName id="E2DF4D1BFFF5FFF9E292A775CCCE7141" ID-CoL="75KFB" authority="Cockerell, 1909 b: 218" authorityName="Cockerell" authorityPageNumber="218" authorityYear="1909" box="[151,654,1299,1323]" class="Insecta" family="Chrysopidae" genus="Palaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="concinnula">
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<emphasis id="17ABEA8AFFF5FFF9E292A775CFDC7141" box="[151,412,1299,1323]" italics="true" pageId="27" pageNumber="328">Palaeochrysa concinnula</emphasis>
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<bibRefCitation id="414E4B69FFF5FFF9E3A0A775CCCE7141" author="Cockerell, T. D. A." box="[421,654,1299,1323]" pageId="27" pageNumber="328" pagination="218 - 219" refId="ref23172" refString="Cockerell, T. D. A. (1909 b) Two fossil Chrysopidae. The Canadian Entomologist, 41, 218 - 219. https: // doi. org / 10.4039 / Ent 41218 - 7" type="journal article" year="1909">Cockerell, 1909b: 218</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, 219,
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<figureCitation id="BDE42A1DFFF5FFF9E0D4A775CD517141" box="[721,785,1299,1323]" captionStart="FIGURE 5" captionStartId="8.[151,250,1442,1466]" captionTargetBox="[151,1436,597,1419]" captionTargetId="figure-207@8.[151,1436,597,1419]" captionTargetPageId="8" captionText="FIGURE 5. Archaeochrysa sp., specimen NHM.I.26921 (counterpart). NHMUK PI I.26921 image © The Trustees of the Natural History Museum. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6522463" httpUri="https://zenodo.org/record/6522463/files/figure.png" pageId="27" pageNumber="328">Fig. 5</figureCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E11AA775CDB67141" author="Cockerell, T. D. A." box="[799,1014,1299,1323]" page="717" pageId="27" pageNumber="328" year="1914">
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<bibRefCitation id="414E4B69FFF5FFF9E11AA775CDB67141" author="Cockerell, T. D. A." box="[799,1014,1299,1323]" pageId="27" pageNumber="328" pagination="714 - 724" refId="ref23258" refString="Cockerell, T. D. A. (1914) New and little-known insects from the Miocene of Florissant, Colorado. Journal of Geology, 22, 714 - 724. https: // doi. org / 10.1086 / 622186" type="journal article" year="1914">Cockerell 1914: 717</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E606A775CA9B7141" author="Carpenter, F. M." box="[1027,1243,1299,1323]" page="262" pageId="27" pageNumber="328" year="1935">
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<bibRefCitation id="414E4B69FFF5FFF9E606A775CA9B7141" author="Carpenter, F. M." box="[1027,1243,1299,1323]" pageId="27" pageNumber="328" pagination="259 - 271" refId="ref22778" refString="Carpenter, F. M. (1935) Tertiary insects of the family Chrysopidae. Journal of Paleontology, 9, 259 - 271." type="journal article" year="1935">Carpenter 1935: 262</bibRefCitation>
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</treatmentCitation>
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; Schlüter 1984: 4;
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<treatmentCitation id="A47E1089FFF5FFF9E2C2A754CC797120" author="Martins-Neto, R. G. & Vulcano, M. A." box="[199,569,1330,1354]" page="189" pageId="27" pageNumber="328" year="1989">
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<bibRefCitation id="414E4B69FFF5FFF9E2C2A754CC797120" author="Martins-Neto, R. G. & Vulcano, M. A." box="[199,569,1330,1354]" pageId="27" pageNumber="328" pagination="189 - 201" refId="ref24601" refString="Martins-Neto, R. G. & Vulcano, M. A. (1989) Neuropteros (Insecta: Planipennia) da Formacao Santana (Cretaceo Inferior), Bacia do Araripe, Nordeste do Brasil. I - Familia Chrysopidae. Anais da Academia Brasileira de Ciencias, 60, 189 - 201. [for 1988]" type="journal article" year="1989">Martins-Neto & Vulcano 1989: 189</bibRefCitation>
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</treatmentCitation>
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, 190;
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<treatmentCitation id="A47E1089FFF5FFF9E07EA754CD3D7120" author="Semeria, Y. & Nel, A." box="[635,893,1330,1354]" page="30" pageId="27" pageNumber="328" year="1990">
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<bibRefCitation id="414E4B69FFF5FFF9E07EA754CD3D7120" author="Semeria, Y. & Nel, A." box="[635,893,1330,1354]" pageId="27" pageNumber="328" pagination="27 - 32" refId="ref25886" refString="Semeria, Y. & Nel, A. (1990) Paleochrysopa monteilsensis gen. et sp. nov., a new fossil of Chrysopidae from the Upper Eocene Formation of Monteils (France), with a review of the known chrysopid fossils (Insecta: Neuroptera). In: Mansell, M. W. & Aspock, H. (Eds.), Advances in Neuropterology. Proceedings of the Third International Symposium on Neuropterology (Berg in Dal, Kruger National Park, 1988). South African Department of Agricultural Development, Pretoria, pp. 27 - 32." type="book chapter" year="1990">Séméria & Nel 1990: 30</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E18CA754CA7E7120" author="Meyer, H. W." box="[905,1086,1330,1354]" page="208" pageId="27" pageNumber="328" year="2003">
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<bibRefCitation id="414E4B69FFF5FFF9E18CA754CA7E7120" author="Meyer, H. W." box="[905,1086,1330,1354]" pageId="27" pageNumber="328" refId="ref24711" refString="Meyer, H. W. (2003) The Fossils of Florissant. Smithsonian Books, Washington, 258 pp." type="book" year="2003">Meyer 2003: 208</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E64FA754CB517120" author="Nel, A. & Delclos, X. & Hutin, A." box="[1098,1297,1330,1354]" page="67" pageId="27" pageNumber="328" year="2005">
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<bibRefCitation id="414E4B69FFF5FFF9E64FA754CB517120" author="Nel, A. & Delclos, X. & Hutin, A." box="[1098,1297,1330,1354]" pageId="27" pageNumber="328" pagination="29 - 68" refId="ref24759" refString="Nel, A., Delclos, X. & Hutin, A. (2005) Mesozoic chrysopid-like Planippenia: a phylogenetic approach (Insecta: Neuroptera). Annales de la Societe Entomologique de France, 41, 29 - 68. https: // doi. org / 10.1080 / 00379271.2005.10697440" type="journal article" year="2005">
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Nel
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<emphasis id="17ABEA8AFFF5FFF9E672A755CAE57120" box="[1143,1189,1330,1354]" italics="true" pageId="27" pageNumber="328">et al</emphasis>
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. 2005: 67
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E71BA754CFDC7103" author="Makarkin, V. N. & Archibald, S. B." page="143" pageId="27" pageNumber="328" year="2013">
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<bibRefCitation id="414E4B69FFF5FFF9E71BA754CFDC7103" author="Makarkin, V. N. & Archibald, S. B." pageId="27" pageNumber="328" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald 2013: 143</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E3A0A736CCBC7102" author="Archibald, S. B. & Makarkin, V. N." box="[421,764,1360,1385]" page="361" pageId="27" pageNumber="328" year="2015">
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<bibRefCitation id="414E4B69FFF5FFF9E3A0A736CCBC7102" author="Archibald, S. B. & Makarkin, V. N." box="[421,764,1360,1385]" pageId="27" pageNumber="328" pagination="359 - 369" refId="ref22418" refString="Archibald, S. B. & Makarkin, V. N. (2015) A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the Early Eocene of Driftwood Canyon, British Columbia, Canada. Canadian Entomologist, 147, 359 - 369. https: // doi. org / 10.4039 / tce. 2014.53" type="journal article" year="2015">Archibald & Makarkin 2015: 361</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFF5FFF9E100A736CA1C7103" author="Archibald, S. B. & Makarkin, V. N." box="[773,1116,1360,1385]" page="399" pageId="27" pageNumber="328" year="2017">
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|
|
<bibRefCitation id="414E4B69FFF5FFF9E100A736CA1C7103" author="Archibald, S. B. & Makarkin, V. N." box="[773,1116,1360,1385]" pageId="27" pageNumber="328" pagination="397 - 400" refId="ref22479" refString="Archibald, S. B. & Makarkin, V. N. (2017) A new fossil green lacewing (Neuroptera: Chrysopidae) from the early Eocene Driftwood Canyon, Canada. Zootaxa, 4324 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4324.2.13" type="journal article" year="2017">Archibald & Makarkin 2017: 399</bibRefCitation>
|
||
|
|
</treatmentCitation>
|
||
|
|
, 400.
|
||
|
|
</treatmentCitationGroup>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="25603698FFF5FFF9E292A709CA6A71ED" blockId="27.[151,1437,1299,1446]" box="[151,1066,1391,1415]" pageId="27" pageNumber="328">
|
||
|
|
<treatmentCitationGroup id="05CF11B6FFF5FFF9E292A709CA6A71ED" box="[151,1066,1391,1415]" pageId="27" pageNumber="328">
|
||
|
|
<treatmentCitation id="A47E1089FFF5FFF9E292A709CC2771ED" author="Carpenter, F. M." box="[151,615,1391,1415]" page="266" pageId="27" pageNumber="328" year="1935">
|
||
|
|
<taxonomicName id="E2DF4D1BFFF5FFF9E292A709CC2771ED" ID-CoL="6QKYP" authority=": Carpenter 1935: 266" authorityName="Carpenter" authorityPageNumber="266" authorityYear="1935" box="[151,615,1391,1415]" class="Insecta" family="Chrysopidae" genus="Lithochrysa" kingdom="Animalia" order="Neuroptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="concinnula">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E292A709CFC871ED" box="[151,392,1391,1415]" italics="true" pageId="27" pageNumber="328">Lithochrysa concinnula</emphasis>
|
||
|
|
:
|
||
|
|
<bibRefCitation id="414E4B69FFF5FFF9E391A709CC2771ED" author="Carpenter, F. M." box="[404,615,1391,1415]" pageId="27" pageNumber="328" pagination="259 - 271" refId="ref22778" refString="Carpenter, F. M. (1935) Tertiary insects of the family Chrysopidae. Journal of Paleontology, 9, 259 - 271." type="journal article" year="1935">Carpenter 1935: 266</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
</treatmentCitation>
|
||
|
|
,
|
||
|
|
<figureCitation id="BDE42A1DFFF5FFF9E074A709CCEF71ED" box="[625,687,1391,1415]" captionStart="FIGURE 5" captionStartId="8.[151,250,1442,1466]" captionTargetBox="[151,1436,597,1419]" captionTargetId="figure-207@8.[151,1436,597,1419]" captionTargetPageId="8" captionText="FIGURE 5. Archaeochrysa sp., specimen NHM.I.26921 (counterpart). NHMUK PI I.26921 image © The Trustees of the Natural History Museum. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6522463" httpUri="https://zenodo.org/record/6522463/files/figure.png" pageId="27" pageNumber="328">Fig. 5</figureCitation>
|
||
|
|
;
|
||
|
|
<treatmentCitation id="A47E1089FFF5FFF9E0BFA709CA6671ED" author="Martins-Neto, R. G. & Vulcano, M. A." box="[698,1062,1391,1415]" page="190" pageId="27" pageNumber="328" year="1989">
|
||
|
|
<bibRefCitation id="414E4B69FFF5FFF9E0BFA709CA6671ED" author="Martins-Neto, R. G. & Vulcano, M. A." box="[698,1062,1391,1415]" pageId="27" pageNumber="328" pagination="189 - 201" refId="ref24601" refString="Martins-Neto, R. G. & Vulcano, M. A. (1989) Neuropteros (Insecta: Planipennia) da Formacao Santana (Cretaceo Inferior), Bacia do Araripe, Nordeste do Brasil. I - Familia Chrysopidae. Anais da Academia Brasileira de Ciencias, 60, 189 - 201. [for 1988]" type="journal article" year="1989">Martins-Neto & Vulcano 1989: 190</bibRefCitation>
|
||
|
|
</treatmentCitation>
|
||
|
|
.
|
||
|
|
</treatmentCitationGroup>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="25603698FFF5FFF9E292A7E8CA5C71CC" blockId="27.[151,1437,1299,1446]" box="[151,1052,1422,1446]" pageId="27" pageNumber="328">
|
||
|
|
<treatmentCitationGroup id="05CF11B6FFF5FFF9E292A7E8CA5C71CC" box="[151,1052,1422,1446]" pageId="27" pageNumber="328">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E292A7E8CF5671CC" box="[151,278,1422,1446]" italics="true" pageId="27" pageNumber="328">Paleochrysa</emphasis>
|
||
|
|
<treatmentCitation id="A47E1089FFF5FFF9E319A7E8CCCA71CC" author="Rodeck, H. G." box="[284,650,1422,1446]" page="290" pageId="27" pageNumber="328" year="1938">
|
||
|
|
<taxonomicName id="E2DF4D1BFFF5FFF9E319A7E8CCCA71CC" ID-CoL="6QKYP" authority=": Rodeck 1938: 290" authorityName="Rodeck" authorityPageNumber="290" authorityYear="1938" box="[284,650,1422,1446]" class="Insecta" family="Chrysopidae" genus="Lithochrysa" isUncertain="true" kingdom="Animalia" order="Neuroptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="concinnula">
|
||
|
|
[
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E320A7E8CF0171CF" box="[293,321,1422,1445]" italics="true" pageId="27" pageNumber="328">sic</emphasis>
|
||
|
|
]
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E34AA7E8CF8071CC" box="[335,448,1422,1446]" italics="true" pageId="27" pageNumber="328">concinnula</emphasis>
|
||
|
|
:
|
||
|
|
<bibRefCitation id="414E4B69FFF5FFF9E3C8A7E8CCCA71CC" author="Rodeck, H. G." box="[461,650,1422,1446]" pageId="27" pageNumber="328" pagination="281 - 304" refId="ref25354" refString="Rodeck, H. G. (1938) Type specimens of fossils in the University of Colorado Museum. The University of Colorado Studies, Series A, 25, 281 - 304." type="journal article" year="1938">Rodeck 1938: 290</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
</treatmentCitation>
|
||
|
|
;
|
||
|
|
<treatmentCitation id="A47E1089FFF5FFF9E096A7E8CD0A71CC" author="Adams, P. A." box="[659,842,1422,1446]" page="231" pageId="27" pageNumber="328" year="1967">
|
||
|
|
<bibRefCitation id="414E4B69FFF5FFF9E096A7E8CD0A71CC" author="Adams, P. A." box="[659,842,1422,1446]" pageId="27" pageNumber="328" pagination="215 - 238" refId="ref21987" refString="Adams, P. A. (1967) A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). Bulletin of the Museum of Comparative Zoology, 135, 215 - 238." type="journal article" year="1967">Adams 1967: 231</bibRefCitation>
|
||
|
|
</treatmentCitation>
|
||
|
|
;
|
||
|
|
<treatmentCitation id="A47E1089FFF5FFF9E150A7E8CA5871CC" author="Schluter, T." box="[853,1048,1422,1446]" page="261" pageId="27" pageNumber="328" year="1982">
|
||
|
|
<bibRefCitation id="414E4B69FFF5FFF9E150A7E8CA5871CC" author="Schluter, T." box="[853,1048,1422,1446]" pageId="27" pageNumber="328" pagination="257 - 264" refId="ref25389" refString="Schluter, T. (1982) Cimbrochrysa moleriensis n. g. n. sp. und Hypochrysa hercyniensis n. sp., zwei fossile Chrysopidae-Arten (Insecta: Planipennia) aus dem europaischen Tertiar. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 5, 257 - 264. https: // doi. org / 10.1127 / njgpm / 1982 / 1982 / 257" type="journal article" year="1982">Schlüter 1982: 261</bibRefCitation>
|
||
|
|
</treatmentCitation>
|
||
|
|
.
|
||
|
|
</treatmentCitationGroup>
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="6DC56513FFF5FFF9E292A7B2CDAD7230" pageId="27" pageNumber="328" type="materials_examined">
|
||
|
|
<paragraph id="25603698FFF5FFF9E292A7B2CA6F7278" blockId="27.[151,1437,1492,2023]" pageId="27" pageNumber="328">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E292A7B2CF057184" bold="true" box="[151,325,1492,1518]" pageId="27" pageNumber="328">Type material.</emphasis>
|
||
|
|
<materialsCitation id="95B73CC5FFF5FFF9E349A7B2CA6F7278" ID-GBIF-Occurrence="3763085304" collectorName="S. A. Rohwer" country="Canada" location="Canada" pageId="27" pageNumber="328" specimenCount="1" typeStatus="holotype">
|
||
|
|
<typeStatus id="FA64883AFFF5FFF9E349A7B2CFF47184" box="[332,436,1492,1518]" pageId="27" pageNumber="328" type="holotype">Holotype</typeStatus>
|
||
|
|
UCM 4220, deposited in UCM, collected by
|
||
|
|
<collectorName id="882A534EFFF5FFF9E1B3A7B2CA0B7184" box="[950,1099,1492,1518]" pageId="27" pageNumber="328">S.A. Rohwer</collectorName>
|
||
|
|
; at least a forewing and a part of a hind wing according to the original description, covered with
|
||
|
|
<collectingCountry id="5DC87608FFF5FFF9E178A79ECD917278" box="[893,977,1528,1554]" name="Canada" pageId="27" pageNumber="328">Canada</collectingCountry>
|
||
|
|
balsam.
|
||
|
|
</materialsCitation>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="25603698FFF5FFF9E2C2A47ACDAD7230" blockId="27.[151,1437,1492,2023]" pageId="27" pageNumber="328">
|
||
|
|
<materialsCitation id="95B73CC5FFF5FFF9E2C2A47ACDA87230" ID-GBIF-Occurrence="3763085315" country="United States of America" county="Teller County" location="Teller County" pageId="27" pageNumber="328" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E2C2A47ACC45725C" bold="true" box="[199,517,1564,1590]" pageId="27" pageNumber="328">
|
||
|
|
<typeStatus id="FA64883AFFF5FFF9E2C2A47ACF42725C" box="[199,258,1564,1590]" pageId="27" pageNumber="328">Type</typeStatus>
|
||
|
|
locality and horizon.
|
||
|
|
</emphasis>
|
||
|
|
<collectingCountry id="5DC87608FFF5FFF9E00BA47ACC0A725C" box="[526,586,1564,1590]" name="United States of America" pageId="27" pageNumber="328">USA</collectingCountry>
|
||
|
|
:
|
||
|
|
<collectingRegion id="E71BF87AFFF5FFF9E052A47ACC84725C" box="[599,708,1564,1590]" country="United States of America" name="Colorado" pageId="27" pageNumber="328">Colorado</collectingRegion>
|
||
|
|
:
|
||
|
|
<collectingCounty id="CC014E14FFF5FFF9E0D5A47ACD31725C" box="[720,881,1564,1590]" pageId="27" pageNumber="328">Teller County</collectingCounty>
|
||
|
|
:
|
||
|
|
<collectorName id="882A534EFFF5FFF9E17BA47ACDAC725C" box="[894,1004,1564,1590]" pageId="27" pageNumber="328">Florissant</collectorName>
|
||
|
|
(
|
||
|
|
<collectorName id="882A534EFFF5FFF9E1F8A47ACA0E725C" box="[1021,1102,1564,1590]" pageId="27" pageNumber="328">Station</collectorName>
|
||
|
|
14, NW SE
|
||
|
|
<collectorName id="882A534EFFF5FFF9E6E0A47BCB31725C" box="[1253,1393,1564,1591]" pageId="27" pageNumber="328">NW Section</collectorName>
|
||
|
|
11,
|
||
|
|
<collectorName id="882A534EFFF5FFF9E292A426CF457230" box="[151,261,1600,1626]" pageId="27" pageNumber="328">Township</collectorName>
|
||
|
|
13S,
|
||
|
|
<collectorName id="882A534EFFF5FFF9E343A427CFCE7231" box="[326,398,1601,1627]" pageId="27" pageNumber="328">Range</collectorName>
|
||
|
|
71W);
|
||
|
|
<collectorName id="882A534EFFF5FFF9E3E6A427CC787231" box="[483,568,1601,1627]" pageId="27" pageNumber="328">Eocene</collectorName>
|
||
|
|
, late
|
||
|
|
<collectorName id="882A534EFFF5FFF9E076A426CCB07230" box="[627,752,1600,1626]" pageId="27" pageNumber="328">Priabonian</collectorName>
|
||
|
|
;
|
||
|
|
<collectorName id="882A534EFFF5FFF9E0FEA426CDA87230" box="[763,1000,1600,1626]" pageId="27" pageNumber="328">Florissant Formation</collectorName>
|
||
|
|
</materialsCitation>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="6DC56513FFF5FFF9E2C2A402CC0372C8" pageId="27" pageNumber="328" type="diagnosis">
|
||
|
|
<paragraph id="25603698FFF5FFF9E2C2A402CC0372C8" blockId="27.[151,1437,1492,2023]" pageId="27" pageNumber="328">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E2C2A402CF027214" bold="true" box="[199,322,1636,1662]" pageId="27" pageNumber="328">Diagnosis.</emphasis>
|
||
|
|
May be distinguished from other species of the genus by the hind wing RP1 located proximad first crossvein ra-rp [other species: distad].
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="6DC56513FFF5FFF9E2C2A4CACF4C73F5" pageId="27" pageNumber="328" type="description">
|
||
|
|
<paragraph id="25603698FFF5FFF9E2C2A4CACEA8733C" blockId="27.[151,1437,1492,2023]" pageId="27" pageNumber="328">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E2C2A4CACF1172AC" bold="true" box="[199,337,1708,1734]" pageId="27" pageNumber="328">Description</emphasis>
|
||
|
|
(emended from
|
||
|
|
<bibRefCitation id="414E4B69FFF5FFF9E016A4CACC9472AD" author="Cockerell, T. D. A." box="[531,724,1708,1735]" pageId="27" pageNumber="328" pagination="218 - 219" refId="ref23172" refString="Cockerell, T. D. A. (1909 b) Two fossil Chrysopidae. The Canadian Entomologist, 41, 218 - 219. https: // doi. org / 10.4039 / Ent 41218 - 7" type="journal article" year="1909">Cockerell 1909b</bibRefCitation>
|
||
|
|
). Forewing
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E15BA4CBCD3872AC" box="[862,888,1709,1734]" italics="true" pageId="27" pageNumber="328">ca</emphasis>
|
||
|
|
.
|
||
|
|
<quantity id="E2279B7DFFF5FFF9E18CA4CACD9E72AD" box="[905,990,1708,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="27" pageNumber="328" unit="mm" value="12.0">12 mm</quantity>
|
||
|
|
. RA space with probably eight or nine crossveins (“nine cells”). Crossvein 2m-cu relatively short, located proximal one-third of
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E672A4B7CAD37280" box="[1143,1171,1745,1770]" italics="true" pageId="27" pageNumber="328">im</emphasis>
|
||
|
|
, only slightly proximad level of 1r-m. Inner gradate series probably in smooth line (“cells between [RP and inner gradate series] “successively shorter from the fourth on, but without any abrupt change of size, as in
|
||
|
|
<taxonomicName id="E2DF4D1BFFF5FFF9E610A57ECADC7358" authorityName="Scudder" authorityYear="1883" box="[1045,1180,1816,1842]" class="Insecta" family="Chrysopidae" genus="Tribochrysa" kingdom="Animalia" order="Neuroptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E610A57ECADC7358" box="[1045,1180,1816,1842]" italics="true" pageId="27" pageNumber="328">Tribochrysa</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
”). Six branches of RP forked.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="25603698FFF5FFF9E2C2A506CF4C73F5" blockId="27.[151,1437,1492,2023]" pageId="27" pageNumber="328">Hind wing. MA basally oblique, then fused with RP from long distance. RP1 originating proximad first crossvein ra-rp.</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="6DC56513FFF5FFFEE2C2A5CECCDE752B" lastPageId="28" lastPageNumber="329" pageId="27" pageNumber="328" type="discussion">
|
||
|
|
<paragraph id="25603698FFF5FFFEE2C2A5CECCDE752B" blockId="27.[151,1437,1492,2023]" lastBlockId="28.[151,1437,151,322]" lastPageId="28" lastPageNumber="329" pageId="27" pageNumber="328">
|
||
|
|
<emphasis id="17ABEA8AFFF5FFF9E2C2A5CECF7B73A8" bold="true" box="[199,315,1960,1986]" pageId="27" pageNumber="328">Remarks.</emphasis>
|
||
|
|
Practically nothing is now discernable in the specimen as it was covered by
|
||
|
|
<collectingCountry id="5DC87608FFF5FFF9E69BA5CECAB273A8" box="[1182,1266,1960,1986]" name="Canada" pageId="27" pageNumber="328">Canada</collectingCountry>
|
||
|
|
balsam, which has lost clarity with age (
|
||
|
|
<figureCitation id="BDE42A1DFFF5FFF9E3BFA5AACC64738C" box="[442,548,1996,2022]" captionStart="FIGURE 20" captionStartId="28.[151,250,1876,1900]" captionTargetBox="[151,1436,345,1845]" captionTargetId="figure-97@28.[151,1436,345,1853]" captionTargetPageId="28" captionText="FIGURE 20. Lithochrysa concinnula (Cockerell, 1909), holotype UCM 4220. A, specimen in current condition, obscured by Canada balsam; B, forewing venation; C, hind wing venation (redrawn from Cockerell 1909b: Fig. 5). Scale bar = 3 mm (B, C not to scale, no scale on original)." figureDoi="http://doi.org/10.5281/zenodo.6522503" httpUri="https://zenodo.org/record/6522503/files/figure.png" pageId="27" pageNumber="328">Fig. 20A</figureCitation>
|
||
|
|
), and so all we know of the species is from the short original description and drawings of fragments of the fore- and hind wing venation reproduced here (
|
||
|
|
<figureCitation id="BDE42A1DFFF2FFFEE609A2F1CAD774DB" box="[1036,1175,151,177]" captionStart="FIGURE 20" captionStartId="28.[151,250,1876,1900]" captionTargetBox="[151,1436,345,1845]" captionTargetId="figure-97@28.[151,1436,345,1853]" captionTargetPageId="28" captionText="FIGURE 20. Lithochrysa concinnula (Cockerell, 1909), holotype UCM 4220. A, specimen in current condition, obscured by Canada balsam; B, forewing venation; C, hind wing venation (redrawn from Cockerell 1909b: Fig. 5). Scale bar = 3 mm (B, C not to scale, no scale on original)." figureDoi="http://doi.org/10.5281/zenodo.6522503" httpUri="https://zenodo.org/record/6522503/files/figure.png" pageId="28" pageNumber="329">Fig. 20B, C</figureCitation>
|
||
|
|
). The original description provides few potentially informative character states (see above), and at least one is diagnostic (see above). Although its generic affinity may be disputed due to the condition of the fossil, the main diagnostic character state of
|
||
|
|
<taxonomicName id="E2DF4D1BFFF2FFFEE2B0A365CF7B7577" box="[181,315,259,285]" class="Insecta" family="Chrysopidae" genus="Lithochrysa" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="17ABEA8AFFF2FFFEE2B0A365CF7B7577" box="[181,315,259,285]" italics="true" pageId="28" pageNumber="329">Lithochrysa</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
distinguishing it from similar genera
|
||
|
|
<taxonomicName id="E2DF4D1BFFF2FFFEE0D8A365CD377577" box="[733,887,259,285]" class="Insecta" family="Chrysopidae" genus="Palaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="17ABEA8AFFF2FFFEE0D8A365CD377577" box="[733,887,259,285]" italics="true" pageId="28" pageNumber="329">Palaeochrysa</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="E2DF4D1BFFF2FFFEE1AEA365CA727577" authorityName="Scudder" authorityYear="1883" box="[939,1074,259,285]" class="Insecta" family="Chrysopidae" genus="Tribochrysa" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="17ABEA8AFFF2FFFEE1AEA365CA727577" box="[939,1074,259,285]" italics="true" pageId="28" pageNumber="329">Tribochrysa</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
(
|
||
|
|
<emphasis id="17ABEA8AFFF2FFFEE647A365CA1E7576" box="[1090,1118,259,284]" italics="true" pageId="28" pageNumber="329">i.e</emphasis>
|
||
|
|
., the smooth arrangement of the inner gradate series) appears to be present.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
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</treatment>
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||
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|
</document>
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