treatments-xml/data/17/8A/53/178A5399B3695D3E8563F03BC9FD6D3F.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.945.39044" ID-GBIF-Dataset="b7273c8d-e5fe-4153-8210-9c966e935fa0" ID-PMC="PMC7351798" ID-Pensoft-Pub="1313-2970-945-17" ID-Pensoft-UUID="5720B610AE6F502EBE10B09F42EDF080" ID-PubMed="32714005" ID-ZooBank="A91BFE95C9534B86871074871CDFAC94" ModsDocID="1313-2970-945-17" checkinTime="1593797452737" checkinUser="pensoft" docAuthor="Pitriana, Pipit, Valente, Luis, von Rintelen, Thomas, Jones, Diana S., Prabowo, Romanus E. &amp; von Rintelen, Kristina" docDate="2020" docId="178A5399B3695D3E8563F03BC9FD6D3F" docLanguage="en" docName="ZooKeys 945: 17-83" docOrigin="ZooKeys 945" docSource="http://dx.doi.org/10.3897/zookeys.945.39044" docTitle="Amphibalanus variegatus" docType="treatment" docVersion="4" id="5720B610AE6F502EBE10B09F42EDF080" lastPageNumber="17" masterDocId="5720B610AE6F502EBE10B09F42EDF080" masterDocTitle="An annotated checklist and integrative biodiversity discovery of barnacles (Crustacea, Cirripedia) from the Moluccas, East Indonesia" masterLastPageNumber="83" masterPageNumber="17" pageNumber="17" updateTime="1668169051581" updateUser="ExternalLinkService">
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<mods:title>An annotated checklist and integrative biodiversity discovery of barnacles (Crustacea, Cirripedia) from the Moluccas, East Indonesia</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pitriana, Pipit</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde- Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany &amp; Research Centre for Deep-sea, Indonesian Institute of Science (LIPI), Jl. Y. Syaranamual, Poka, Tlk. Ambon, Kota Ambon, Maluku, Indonesia &amp; Institute of Geological Sciences, Freie Universitaet Berlin, Malteserstrasse 74 - 100 Building C and D, 12249 Berlin, Germany</mods:affiliation>
<mods:nameIdentifier type="email">pipit.pitriana@mfn.berlin</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Valente, Luis</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde- Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany &amp; Naturalis Biodiversity Center, Understanding Evolution Group, Postbus 9517, 2300 RA Leiden, the Netherlands</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>von Rintelen, Thomas</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde- Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6253-3078</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jones, Diana S.</mods:namePart>
<mods:affiliation>The Western Australian Museum, 49 Kew Street, Welshpool WA 6106, Locked Bag 49, Welshpool DC WA 6986, Australia</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Prabowo, Romanus E.</mods:namePart>
<mods:affiliation>Faculty of Biology, Universitas Jenderal Soedirman, Purwokerto, 53122, Indonesia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0632-7461</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>von Rintelen, Kristina</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde- Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>945</mods:number>
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<mods:start>17</mods:start>
<mods:end>83</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.945.39044</mods:url>
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<mods:classification>journal article</mods:classification>
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<treatment ID-GBIF-Taxon="164910951" LSID="urn:lsid:plazi:treatment:178A5399B3695D3E8563F03BC9FD6D3F" httpUri="http://treatment.plazi.org/id/178A5399B3695D3E8563F03BC9FD6D3F" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection pageId="0" pageNumber="17" type="nomenclature">
<paragraph pageId="0" pageNumber="17">
<taxonomicName LSID="178A5399-B369-5D3E-8563-F03BC9FD6D3F" authority="(Darwin, 1854)" baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">Amphibalanus variegatus (Darwin, 1854)</taxonomicName>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">Figure 23a-p</figureCitation>
, 
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Annotated checklist tabulation of barnacle species from the Moluccas, Eastern Indonesia." pageId="0" pageNumber="17">Table 1: species no. 89</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="reference_group">
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="amphitrite">Balanus amphitrite</taxonomicName>
var. (8) 
<taxonomicName authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="variety" species="amphitrite" variety="variegatus">variegatus</taxonomicName>
Darwin, 1854: 241.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite subsp. var. var. stutsburi" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="variety" species="amphitrite" subSpecies="var." variety="stutsburi">Balanus amphitrite var. stutsburi</taxonomicName>
<normalizedToken originalValue="Krüger">Krueger</normalizedToken>
, 1914: 437.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus concavus subsp. sinensis" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="concavus" subSpecies="sinensis">Balanus concavus sinensis</taxonomicName>
Broch, 1931: 63, fig. 23.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite subsp. rafflesia" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="amphitrite" subSpecies="rafflesia">Balanus amphitrite rafflesia</taxonomicName>
Nilsson-Cantell, 1934a: 64.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite subsp. var. var. cirratus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="variety" species="amphitrite" subSpecies="var." variety="cirratus">Balanus amphitrite var. cirratus</taxonomicName>
: Pope 1945: 362, pl. 28 fig. 6, pl. 29 fig. 6, pl. 30 figs 13, 14.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite subsp. cirratus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="amphitrite" subSpecies="cirratus">Balanus amphitrite cirratus</taxonomicName>
: Skerman 1960: 610, figs 1, 3 (
<emphasis bold="true" pageId="0" pageNumber="17">non</emphasis>
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite subsp. cirratus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="amphitrite" subSpecies="cirratus">Balanus amphitrite cirratus</taxonomicName>
Darwin, 1854).
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">Balanus variegatus</taxonomicName>
: Harding 1962: 291, pl. 10 figs a-k; Zevina et al. 1992: 92, fig. 64.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus variegatus subsp. var. var. cirratus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="variety" species="variegatus" subSpecies="var." variety="cirratus">Balanus variegatus var. cirratus</taxonomicName>
: Pope 1966: 179.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="amphitrite">Balanus amphitrite</taxonomicName>
: Foster 1967: 83 (part.).287.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus kondakovi" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="kondakovi">Balanus kondakovi</taxonomicName>
: Henry &amp; McLaughlin 1975: 78 (part., New Zealand specimens; 
<emphasis bold="true" pageId="0" pageNumber="17">non</emphasis>
<taxonomicName family="Balanidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="B. kondakovi" order="Sessilia" pageId="0" pageNumber="17" rank="species" species="kondakovi">B. kondakovi</taxonomicName>
Tarasov &amp; Zevina, 1957).
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName authority="variegatus" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus variegatus subsp. variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="variegatus" subSpecies="variegatus">Balanus variegatus variegatus</taxonomicName>
: Foster 1979: 111, fig. 67, pl. 14b.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName authorityName="Ren &amp; Liu" authorityYear="1978" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus cirratus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="cirratus">Balanus cirratus</taxonomicName>
: Ren &amp; Liu, 1978: 145, figs 14, 15 (1-13), pl. 4 figs 15-20, pl. 5 figs 1-6.non 
<taxonomicName authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite subsp. variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="amphitrite" subSpecies="variegatus">Balanus amphitrite variegatus</taxonomicName>
: Nilsson-Cantell 1934a: 60.
</paragraph>
<paragraph pageId="0" pageNumber="17">
non 
<taxonomicName authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">Balanus variegatus</taxonomicName>
: Henry &amp; McLaughlin 1975: 78, fig. 17, pls. 6, 7; Utinomi 1968b: 171 (= 
<taxonomicName family="Balanidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="B. cirratus" order="Sessilia" pageId="0" pageNumber="17" rank="species" species="cirratus">B. cirratus</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">Amphibalanus variegatus</taxonomicName>
: Pitombo 2004: 274; Horikoshi and Okamoto 2005: 49, fig.3.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="materials_examined">
<paragraph pageId="0" pageNumber="17">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" italics="true" pageId="0" pageNumber="17">Ambon Island</emphasis>
: 15 specimens, MZB Cru Cir 014, Waitatiri, 
<geoCoordinate degrees="3" direction="south" minutes="37" orientation="latitude" precision="1" seconds="04.0" value="-3.6177778">3°37'04.0&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="128" direction="east" minutes="16" orientation="longitude" precision="1" seconds="20.3" value="128.27231">128°16'20.3&quot;E</geoCoordinate>
, coll. P. Pitriana &amp; D. Tala, 19 Sep 2017. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="17">Saparua Island</emphasis>
: 10 specimens, MZB Cru Cir 015, Teluk Saparua, 
<geoCoordinate degrees="3" direction="south" minutes="34" orientation="latitude" precision="1" seconds="25.7" value="-3.5738056">3°34'25.7&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="128" direction="east" minutes="39" orientation="longitude" precision="1" seconds="25.8" value="128.65717">128°39'25.8&quot;E</geoCoordinate>
, coll. P. Pitriana &amp; D. Tala, 22 Sep 2016.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="genbank accession numbers">
<paragraph pageId="0" pageNumber="17">GenBank accession numbers.</paragraph>
<paragraph pageId="0" pageNumber="17">COI gene (MK995342-MK995345), 18S (MK981355).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="diagnosis">
<paragraph pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="17">Primary parietal tubes with transverse septa; exterior of shell with longitudinal and horizontal striations; anterior margin of cirrus III without conical denticles, erect hooks below posterior angles of distal articles of rami absent.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="description">
<paragraph pageId="0" pageNumber="17">Description.</paragraph>
<paragraph pageId="0" pageNumber="17">
Shell steeply conical, tubular in crowded populations; six parietal plates, smooth, thin, brownish purple externally with longitudinal stripes crosshatched by transverse bands, single row of internal tubes (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23b</figureCitation>
); carina forming a spout-like projection; radii wide, summits oblique, pink-purple; alae with summits oblique; orifice toothed (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23a-c</figureCitation>
); sheath purple with white bands, vesicular; basis calcareous, porous; scutum externally with growth lines prominent, internally with articular ridge high, adductor ridge moderately long; tergum with spur furrow, externally purple, margins white, spur pointed, basal margin deeply excavated on either side of spur, depressor muscle crests prominent(Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23d, e</figureCitation>
); cirrus III without conical denticles on anterior margin (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23h</figureCitation>
); maxilla without notch (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23l</figureCitation>
), mandible with four teeth (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23n</figureCitation>
), labrum notched, denticulate (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23o</figureCitation>
); penis with basidorsal point, with two apical setae (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphibalanus variegatus (Darwin, 1854) (MZB Cru Cir 014 - 1) a upper view b lower view c side view d external view of scutum and tergum e internal view of scutum and tergum f cirrus I g cirrus II h cirrus III i cirrus IV j cirrus V k cirrus VI l maxilla m maxillule n mandible o labrum p penis. Scale bars: 3 mm (a-c); 1 mm (d-h); 2 mm (i-k); 2 mm (p); 0.5 mm (l-o)." figureDoi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17">23p</figureCitation>
). Basal length (8.3-11.8 mm, basal width 6.9-10.4 mm, height 4.3-8.4 mm; orifice length 4.8-11.8 mm; orifice width 3.4-5.3 mm (measurements for ten specimens are presented in Suppl. material 1: Table S22).
</paragraph>
<caption doi="10.3897/zookeys.945.39044.figure23" httpUri="https://binary.pensoft.net/fig/428609" pageId="0" pageNumber="17" start="Figure 23" startId="F23">
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Figure 23.</emphasis>
<taxonomicName baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis italics="true" pageId="0" pageNumber="17">Amphibalanus variegatus</emphasis>
</taxonomicName>
(Darwin, 1854) (MZB Cru Cir 014-1) 
<emphasis bold="true" pageId="0" pageNumber="17">a</emphasis>
upper view 
<emphasis bold="true" pageId="0" pageNumber="17">b</emphasis>
lower view 
<emphasis bold="true" pageId="0" pageNumber="17">c</emphasis>
side view 
<emphasis bold="true" pageId="0" pageNumber="17">d</emphasis>
external view of scutum and tergum 
<emphasis bold="true" pageId="0" pageNumber="17">e</emphasis>
internal view of scutum and tergum 
<emphasis bold="true" pageId="0" pageNumber="17">f</emphasis>
cirrus I 
<emphasis bold="true" pageId="0" pageNumber="17">g</emphasis>
cirrus II 
<emphasis bold="true" pageId="0" pageNumber="17">h</emphasis>
cirrus III 
<emphasis bold="true" pageId="0" pageNumber="17">i</emphasis>
cirrus IV 
<emphasis bold="true" pageId="0" pageNumber="17">j</emphasis>
cirrus V 
<emphasis bold="true" pageId="0" pageNumber="17">k</emphasis>
cirrus VI 
<emphasis bold="true" pageId="0" pageNumber="17">l</emphasis>
maxilla 
<emphasis bold="true" pageId="0" pageNumber="17">m</emphasis>
maxillule 
<emphasis bold="true" pageId="0" pageNumber="17">n</emphasis>
mandible 
<emphasis bold="true" pageId="0" pageNumber="17">o</emphasis>
labrum 
<emphasis bold="true" pageId="0" pageNumber="17">p</emphasis>
penis. Scale bars: 3 mm (
<emphasis bold="true" pageId="0" pageNumber="17">a-c</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="17">d-h</emphasis>
); 2 mm (
<emphasis bold="true" pageId="0" pageNumber="17">i-k</emphasis>
); 2 mm (
<emphasis bold="true" pageId="0" pageNumber="17">p</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="17">l-o</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="distribution">
<paragraph pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis italics="true" pageId="0" pageNumber="17">Amphibalanus variegatus</emphasis>
</taxonomicName>
has been reported from the Indo-west Pacific: Bay of Bengal; Sumatra; New Zealand, Australia; Indonesia; Singapore; Vietnam; Gulf of Siam; Hong Kong; W Kyushu; Vladivostok; and is a common fouling species (
<bibRefCitation author="Henry, DP" journalOrPublisher="Zoologische Verhandelingen" pageId="0" pageNumber="17" pagination="1 - 254" publicationUrl="http://www.repository.naturalis.nl/record/317846" refId="B26" refString="Henry, DP, McLaughlin, PA, 1975. The barnacles of the Balanus amphitrite complex (Cirripedia, Thoracica). Zoologische Verhandelingen 141 (1): 1 - 254, http://www.repository.naturalis.nl/record/317846" title="The barnacles of the Balanus amphitrite complex (Cirripedia, Thoracica)." url="http://www.repository.naturalis.nl/record/317846" volume="141" year="1975">Henry and McLaughlin 1975</bibRefCitation>
; 
<bibRefCitation author="Jones, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="17" pagination="241 - 311" refId="B36" refString="Jones, DS, Hosie, AM, 2016. A checklist of the barnacles (Cirripedia: Thoracica) of Singapore and neighbouring waters. Raffles Bulletin of Zoology 34: 241 - 311" title="A checklist of the barnacles (Cirripedia: Thoracica) of Singapore and neighbouring waters." volume="34" year="2016">Jones and Hosie 2016</bibRefCitation>
). In this study, 
<taxonomicName lsidName="A. variegatus" pageId="0" pageNumber="17" rank="species" species="variegatus">
<emphasis italics="true" pageId="0" pageNumber="17">A. variegatus</emphasis>
</taxonomicName>
was found on the islands of Ambon (at Waitatiri) and Saparua (at Teluk Saparua) on stones and a plastic bag (a map with the occurrence of 
<taxonomicName baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis italics="true" pageId="0" pageNumber="17">Amphibalanus variegatus</emphasis>
</taxonomicName>
in the Moluccas is shown in Suppl. material 1: Fig. S4).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="remarks">
<paragraph pageId="0" pageNumber="17">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis italics="true" pageId="0" pageNumber="17">Amphibalanus variegatus</emphasis>
</taxonomicName>
is a member of the 
<taxonomicName class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balanus amphitrite" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="amphitrite">
<emphasis italics="true" pageId="0" pageNumber="17">Balanus amphitrite</emphasis>
</taxonomicName>
complex, whose members can be difficult to distinguish morphologically. 
<taxonomicName baseAuthorityName="Darwin" baseAuthorityYear="1854" class="Maxillopoda" family="Balanidae" genus="Amphibalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphibalanus variegatus" order="Sessilia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis italics="true" pageId="0" pageNumber="17">Amphibalanus variegatus</emphasis>
</taxonomicName>
can be differentiated by its vesicular sheath, and from 
<taxonomicName lsidName="A. reticulatus" pageId="0" pageNumber="17" rank="species" species="reticulatus">
<emphasis italics="true" pageId="0" pageNumber="17">A. reticulatus</emphasis>
</taxonomicName>
by features of the tergum, armature of cirrus II and the lack of erect teeth below the posterior distal angles of cirri III-VI (
<bibRefCitation author="Henry, DP" journalOrPublisher="Zoologische Verhandelingen" pageId="0" pageNumber="17" pagination="1 - 254" publicationUrl="http://www.repository.naturalis.nl/record/317846" refId="B26" refString="Henry, DP, McLaughlin, PA, 1975. The barnacles of the Balanus amphitrite complex (Cirripedia, Thoracica). Zoologische Verhandelingen 141 (1): 1 - 254, http://www.repository.naturalis.nl/record/317846" title="The barnacles of the Balanus amphitrite complex (Cirripedia, Thoracica)." url="http://www.repository.naturalis.nl/record/317846" volume="141" year="1975">Henry and McLaughlin 1975</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>