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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.88.12891" ID-PMC="PMC5672133" ID-Pensoft-Pub="1314-2003-88-1" ID-Pensoft-UUID="FF96FFA8FFCD17717013D224FF85FF9F" ID-PubMed="29118645" ID-Zenodo-Dep="1138043" ModsDocID="1314-2003-88-1" checkinTime="1507734418711" checkinUser="pensoft" docAuthor="Wood, John R. I., Munoz-Rodriguez, Pablo, Degen, Rosa &amp; Scotland, Robert W." docDate="2017" docId="684B51783BB45D3AAD8B664FCDEA11AB" docLanguage="en" docName="PhytoKeys 88: 1-38" docOrigin="PhytoKeys 88" docPubDate="2017-10-11" docSource="http://dx.doi.org/10.3897/phytokeys.88.12891" docTitle="Ipomoea cuscoensis J. R. I. Wood &amp; P. Munoz 2017, sp. nov." docType="treatment" docVersion="3" id="FF96FFA8FFCD17717013D224FF85FF9F" lastPageNumber="9" masterDocId="FF96FFA8FFCD17717013D224FF85FF9F" masterDocTitle="New species of Ipomoea (Convolvulaceae) from South America" masterLastPageNumber="38" masterPageNumber="1" pageNumber="5" updateTime="1668142099483" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species of Ipomoea (Convolvulaceae) from South America</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wood, John R. I.</mods:namePart>
<mods:affiliation>Department of Plant Sciences, University of Oxford, South Parks Road, Oxford, OX 1 3 RB, UK &amp; Honorary Research Associate, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AB, UK</mods:affiliation>
<mods:nameIdentifier type="email">jriwood@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Munoz-Rodriguez, Pablo</mods:namePart>
<mods:affiliation>Department of Plant Sciences, University of Oxford, South Parks Road, Oxford, OX 1 3 RB, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Degen, Rosa</mods:namePart>
<mods:affiliation>Facultad de Ciencias Quimicas, Departamento de Botanica, Universidad Nacional de Asuncion, Herbario FCQ, Casilla de Correo Campus UNA, 11001 - 3291, Paraguay</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scotland, Robert W.</mods:namePart>
<mods:affiliation>Department of Plant Sciences, University of Oxford, South Parks Road, Oxford, OX 1 3 RB, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>88</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>38</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.88.12891</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.88.12891</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-88-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF96FFA8FFCD17717013D224FF85FF9F</mods:identifier>
<mods:identifier type="Zenodo-Dep">1138043</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182225739" LSID="urn:lsid:plazi:treatment:684B51783BB45D3AAD8B664FCDEA11AB" httpUri="http://treatment.plazi.org/id/684B51783BB45D3AAD8B664FCDEA11AB" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
<paragraph pageId="4" pageNumber="5">
<taxonomicName LSID="684B5178-3BB4-5D3A-AD8B-664FCDEA11AB" authority="J. R. I. Wood &amp; P. Muñoz" authorityName="J. R. I. Wood &amp; P. Munoz" authorityYear="2017" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ipomoea cuscoensis" order="Solanales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="cuscoensis" status="sp. nov.">
Ipomoea cuscoensis J.R.I.Wood &amp; P.
<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="5">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ipomoea cuscoensis. A habit B habit; C outer sepal; D inner sepal. Drawn by Rosemary Wise from Galiano et al. 5146." figureDoi="10.3897/phytokeys.88.12891.figure3" httpUri="https://binary.pensoft.net/fig/160684" pageId="4" pageNumber="5">Figures 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Image of Vargas 22075 showing the distinctive purple flowers of Ipomoea cuscoensis." figureDoi="10.3897/phytokeys.88.12891.figure4" httpUri="https://binary.pensoft.net/fig/160685" pageId="4" pageNumber="5">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph pageId="5" pageNumber="6">
<pageBreakToken pageId="5" pageNumber="6" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="5" pageNumber="6">
<taxonomicName authorityName="J. R. I. Wood &amp; P. Munoz" authorityYear="2017" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ipomoea cuscoensis" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="cuscoensis">
<emphasis italics="true" pageId="5" pageNumber="6">Ipomoea cuscoensis</emphasis>
</taxonomicName>
is a distinct species because of its glabrous indumentum, 3-5-lobed leaves, large ovate-elliptic scarious-margined sepals and striking purple corolla. It is most likely to be confused with
<taxonomicName lsidName="I. peruviana" pageId="5" pageNumber="6" rank="species" species="peruviana">
<emphasis italics="true" pageId="5" pageNumber="6">I. peruviana</emphasis>
</taxonomicName>
<normalizedToken originalValue="O’Donell">O'Donell</normalizedToken>
and
<taxonomicName lsidName="I. clavata" pageId="5" pageNumber="6" rank="species" species="clavata">
<emphasis italics="true" pageId="5" pageNumber="6">I. clavata</emphasis>
</taxonomicName>
(G. Don) Ooststr. ex J.F. Macbr., but differs from both in the purple corolla, c. 6.5 cm long, and in having all leaves 3-5-lobed.
<taxonomicName lsidName="I. peruviana" pageId="5" pageNumber="6" rank="species" species="peruviana">
<emphasis italics="true" pageId="5" pageNumber="6">I. peruviana</emphasis>
</taxonomicName>
differs additionally in its much larger (10-11 cm long), very pale blue corolla and mostly entire leaves while
<taxonomicName lsidName="I. clavata" pageId="5" pageNumber="6" rank="species" species="clavata">
<emphasis italics="true" pageId="5" pageNumber="6">I. clavata</emphasis>
</taxonomicName>
differs in its solitary blue flowers and pilose stems.
</paragraph>
<caption ID-Zenodo-Dep="1138069" doi="10.3897/phytokeys.88.12891.figure3" httpUri="https://binary.pensoft.net/fig/160684" pageId="5" pageNumber="6" start="Figure 3" startId="F3">
<paragraph pageId="5" pageNumber="6">
<emphasis bold="true" pageId="5" pageNumber="6">Figure 3.</emphasis>
<taxonomicName authorityName="J. R. I. Wood &amp; P. Munoz" authorityYear="2017" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ipomoea cuscoensis" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="cuscoensis">
<emphasis italics="true" pageId="5" pageNumber="6">Ipomoea cuscoensis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="5" pageNumber="6">A</emphasis>
habit
<emphasis bold="true" pageId="5" pageNumber="6">B</emphasis>
habit;
<emphasis bold="true" pageId="5" pageNumber="6">C</emphasis>
outer sepal;
<emphasis bold="true" pageId="5" pageNumber="6">D</emphasis>
inner sepal. Drawn by Rosemary Wise from
<emphasis italics="true" pageId="5" pageNumber="6">Galiano et al.</emphasis>
5146.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1138071" doi="10.3897/phytokeys.88.12891.figure4" httpUri="https://binary.pensoft.net/fig/160685" pageId="5" pageNumber="6" start="Figure 4" startId="F4">
<paragraph pageId="5" pageNumber="6">
<emphasis bold="true" pageId="5" pageNumber="6">Figure 4.</emphasis>
Image of
<taxonomicName genus="Vargas" lsidName="Vargas" pageId="5" pageNumber="6" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="6">Vargas</emphasis>
</taxonomicName>
22075 showing the distinctive purple flowers of
<taxonomicName authorityName="J. R. I. Wood &amp; P. Munoz" authorityYear="2017" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ipomoea cuscoensis" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="cuscoensis">
<emphasis italics="true" pageId="5" pageNumber="6">Ipomoea cuscoensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="materials_examined">
<paragraph pageId="5" pageNumber="6">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="5" pageNumber="6">
<materialsCitation collectingDate="1963-03-15" country="PERU" county="Anta" elevation="2300" location="Limatambo" municipality="Sisal" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
<collectingCountry name="Peru">PERU</collectingCountry>
.
<collectingRegion country="Peru" name="Cusco">Cusco</collectingRegion>
:
<collectingCounty>Anta</collectingCounty>
,
<collectingMunicipality>Sisal</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:684B51783BB45D3AAD8B664FCDEA11AB:97E5389B1FD6A34A96F12E10A930A819" country="PERU" county="Anta" municipality="Sisal" name="Limatambo" stateProvince="Cusco">Limatambo</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<collectingDate value="1963-03-15">15 March 1963</collectingDate>
,
<emphasis italics="true" pageId="5" pageNumber="6">
C.
<taxonomicName genus="Vargas" lsidName="Vargas" pageId="5" pageNumber="6" rank="genus">Vargas</taxonomicName>
</emphasis>
14325 (
<typeStatus>holotype</typeStatus>
CUZ, isotype
</materialsCitation>
<collectingCountry name="United States of America">US</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph pageId="5" pageNumber="6">Description.</paragraph>
<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Twining perennial of unknown height; stems glabrous. Leaves petiolate, 3-6
<normalizedToken originalValue="×">x</normalizedToken>
3-6.5 cm, 3-5-lobed, the terminal lobe larger, lobes elliptic in outline, apex acuminate to an obtuse mucronate tip, base shallowly cordate, margin weakly crenate, both surfaces glabrous, abaxially paler with prominent whitish veins; petioles 1.3-3 cm. Inflorescence of pedunculate axillary cymes with up to 7 flowers; peduncles
<pageBreakToken pageId="6" pageNumber="7" start="start">4</pageBreakToken>
-6 cm, glabrous; bracteoles caducous, not seen; pedicels 8-20 mm, glabrous; calyx narrowly ovoid, sepals somewhat unequal, outer sepals 20-22
<normalizedToken originalValue="×">x</normalizedToken>
10 mm, ovate to ovate-elliptic, shortly mucronate, glabrous, margins scarious; inner sepals 15
<normalizedToken originalValue="×">x</normalizedToken>
8 mm, ellipsoid, mucronate, the scarious margins broad; corolla 6.5 cm long, campanulate, glabrous, deep pink, limb 3-4 cm diameter. Capsule and seeds unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="distribution">
<paragraph pageId="7" pageNumber="8">
<pageBreakToken pageId="7" pageNumber="8" start="start">Distribution</pageBreakToken>
and habitat.
</paragraph>
<paragraph pageId="7" pageNumber="8">
Endemic to dry forest and scrub at 2100-2700 m in Southern PERU. Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of equatorial South America showing the distribution of Ipomoea cuscoensis (●), I. graminifolia (✖), I. uninervis (▲) and I. velutinifolia (■)." figureDoi="10.3897/phytokeys.88.12891.figure5" httpUri="https://binary.pensoft.net/fig/160687" pageId="7" pageNumber="8">5</figureCitation>
.
</paragraph>
<caption ID-Zenodo-Dep="1138073" doi="10.3897/phytokeys.88.12891.figure5" httpUri="https://binary.pensoft.net/fig/160687" pageId="7" pageNumber="8" start="Figure 5" startId="F5">
<paragraph pageId="7" pageNumber="8">
<emphasis bold="true" pageId="7" pageNumber="8">Figure 5.</emphasis>
Map of equatorial South America showing the distribution of
<taxonomicName authorityName="J. R. I. Wood &amp; P. Munoz" authorityYear="2017" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ipomoea cuscoensis" order="Solanales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="cuscoensis">
<emphasis italics="true" pageId="7" pageNumber="8">Ipomoea cuscoensis</emphasis>
</taxonomicName>
(●),
<taxonomicName lsidName="I. graminifolia" pageId="7" pageNumber="8" rank="species" species="graminifolia">
<emphasis italics="true" pageId="7" pageNumber="8">I. graminifolia</emphasis>
</taxonomicName>
(✖),
<taxonomicName lsidName="I. uninervis" pageId="7" pageNumber="8" rank="species" species="uninervis">
<emphasis italics="true" pageId="7" pageNumber="8">I. uninervis</emphasis>
</taxonomicName>
(▲) and
<taxonomicName lsidName="I. velutinifolia" pageId="7" pageNumber="8" rank="species" species="velutinifolia">
<emphasis italics="true" pageId="7" pageNumber="8">I. velutinifolia</emphasis>
</taxonomicName>
(■).
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="additional collections seen">
<paragraph pageId="7" pageNumber="8">Additional collections seen.</paragraph>
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Apurimac: Abancay, Cachora, 2700 m, 17 Jan 1950,
<emphasis italics="true" pageId="7" pageNumber="8">
C.
<taxonomicName genus="Vargas" lsidName="Vargas" pageId="7" pageNumber="8" rank="genus">Vargas</taxonomicName>
</emphasis>
9083 (CUZ); Grau, bajando a Kai Ranka, 2400 m, 9 March 1946,
<emphasis italics="true" pageId="7" pageNumber="8">
C.
<taxonomicName genus="Vargas" lsidName="Vargas" pageId="7" pageNumber="8" rank="genus">Vargas</taxonomicName>
</emphasis>
<pageBreakToken pageId="8" pageNumber="9" start="start">5826</pageBreakToken>
(CUZ). Cusco: Anta,
<normalizedToken originalValue="Río">Rio</normalizedToken>
Blanco, 2100 m, 22 March 1971,
<emphasis italics="true" pageId="8" pageNumber="9">
C.
<taxonomicName genus="Vargas" lsidName="Vargas" pageId="8" pageNumber="9" rank="genus">Vargas</taxonomicName>
</emphasis>
22075 (CUZ); Mollepata, 2710 m, 3 May 2003,
<emphasis italics="true" pageId="8" pageNumber="9">W. Galiano et al</emphasis>
. 5146 (MO).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="conservation status">
<paragraph pageId="8" pageNumber="9">Conservation status.</paragraph>
<paragraph pageId="8" pageNumber="9">
Although we have seen five separate collections of this species, they all come from a restricted area of Peru and the labels provide no information about the
<normalizedToken originalValue="plant’s">plant's</normalizedToken>
frequency. It should probably be treated as a &quot;black star&quot; species within the classification of
<bibRefCitation author="Hawthorne, WD" journalOrPublisher="Revista Brasileira de Botanica" pageId="32" pageNumber="33" publicationUrl="http://herbaria-old.plants.ox.ac.uk/rbs/" refId="B7" refString="Hawthorne, WD, Marshall, CAM, 2016. A Manual for Rapid Botanical Survey (RBS) and measurement of vegetation bioquality. Published online, June 2016. Department of Plant Sciences, University of Oxford, U.K. http://herbaria-old.plants.ox.ac.uk/rbs/" title="A Manual for Rapid Botanical Survey (RBS) and measurement of vegetation bioquality. Published online, June 2016. Department of Plant Sciences, University of Oxford, U. K." url="http://herbaria-old.plants.ox.ac.uk/rbs/" year="2016">Hawthorne and Marshall (2016)</bibRefCitation>
and be classified as Endangered (EN) within
<bibRefCitation author="IUCN" journalOrPublisher="Revista Brasileira de Botanica" pageId="32" pageNumber="33" publicationUrl="http://s3.amazonaws.com/iucnredlist-newcms/staging/public/attachments/3097/redlist_cats_crit_en.pdf" refId="B8" refString="IUCN, 2012. IUCN Red List Categories and Criteria. Ed 2. International Union for the Conservation of Nature, Gland, Switzerland. http://s3.amazonaws.com/iucnredlist-newcms/staging/public/attachments/3097/redlist_cats_crit_en.pdf" title="IUCN Red List Categories and Criteria. Ed 2. International Union for the Conservation of Nature, Gland, Switzerland." url="http://s3.amazonaws.com/iucnredlist-newcms/staging/public/attachments/3097/redlist_cats_crit_en.pdf" year="2012">IUCN (2012)</bibRefCitation>
guidelines because of an area of occupancy of &lt;12, 000 km², based on an analysis using GEOCAT. However, these classifications should be treated as provisional until a full field evaluation is carried out.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="etymology">
<paragraph pageId="8" pageNumber="9">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="9">
The epithet &quot;
<taxonomicName genus="Vargas" lsidName="cuscoensis" pageId="8" pageNumber="9" rank="species" species="cuscoensis">
<emphasis italics="true" pageId="8" pageNumber="9">cuscoensis</emphasis>
</taxonomicName>
&quot; refers to the Cusco region to which this species is endemic.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="notes">
<paragraph pageId="8" pageNumber="9">Notes.</paragraph>
<paragraph pageId="8" pageNumber="9">
Phylogenetic analysis using
<emphasis italics="true" pageId="8" pageNumber="9">ITS</emphasis>
sequences (unpublished data) indicates that this species belongs to a small clade, which includes
<taxonomicName lsidName="I. lindenii" pageId="8" pageNumber="9" rank="species" species="lindenii">
<emphasis italics="true" pageId="8" pageNumber="9">I. lindenii</emphasis>
</taxonomicName>
M Martens &amp; Galeotti,
<taxonomicName lsidName="I. corymbosa" pageId="8" pageNumber="9" rank="species" species="corymbosa">
<emphasis italics="true" pageId="8" pageNumber="9">I. corymbosa</emphasis>
</taxonomicName>
(L.) Roth ex Roem. &amp; Schult. and
<taxonomicName lsidName="I. clavata" pageId="8" pageNumber="9" rank="species" species="clavata">
<emphasis italics="true" pageId="8" pageNumber="9">I. clavata</emphasis>
</taxonomicName>
, all of which are characterised by their relatively large sepals. It has been confused with
<taxonomicName lsidName="I. peruviana" pageId="8" pageNumber="9" rank="species" species="peruviana">
<emphasis italics="true" pageId="8" pageNumber="9">I. peruviana</emphasis>
</taxonomicName>
both in the herbarium and also by
<bibRefCitation DOI="https://doi.org/10.1007/s12225-017-9682-9" author="Wood, JRI" journalOrPublisher="Kew Bulletin" pageId="33" pageNumber="34" pagination="1 - 26" refId="B18" refString="Wood, JRI, Scotland, RW, 2017b. Notes on Ipomoea (Convolvulaceae) from the Amazonian periphery. Kew Bulletin 72 (10): 1 - 26, DOI: https://doi.org/10.1007/s12225-017-9682-9" title="Notes on Ipomoea (Convolvulaceae) from the Amazonian periphery." url="https://doi.org/10.1007/s12225-017-9682-9" volume="72" year="2017 b">Wood and Scotland (2017b)</bibRefCitation>
, partly because
<taxonomicName lsidName="I. peruviana" pageId="8" pageNumber="9" rank="species" species="peruviana">
<emphasis italics="true" pageId="8" pageNumber="9">I. peruviana</emphasis>
</taxonomicName>
is poorly known and partly because images of
<taxonomicName lsidName="I. cuscoensis" pageId="8" pageNumber="9" rank="species" species="cuscoensis">
<emphasis italics="true" pageId="8" pageNumber="9">I. cuscoensis</emphasis>
</taxonomicName>
have only recently become available to us after
<normalizedToken originalValue="Muñoz’s">Munoz's</normalizedToken>
visit to CUZ in April 2017. These make clear the very distinctive flowers of the new species and confirm that it grows at the relatively high altitude of 2100 to 2710 m, at a much higher elevation than the lowland
<taxonomicName lsidName="I. peruviana" pageId="8" pageNumber="9" rank="species" species="peruviana">
<emphasis italics="true" pageId="8" pageNumber="9">I. peruviana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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