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<mods:title id="55931933FFE19798C399C01F0D7B0D3F">A taxonomic revision of the ecologically important Ochna holstii (Ochnaceae) complex using molecular and morphological data</mods:title>
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<mods:namePart id="29463FEA0F184E1150BE747FD3120999">Shah, Toral</mods:namePart>
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<mods:namePart id="C0C6388C41D812FD6B3743626189F5CD">Savolainen, Vincent</mods:namePart>
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1. 
<taxonomicName id="3C3972F0548F752A3DE9235BA2716DC0" ID-CoL="48D5H" LSID="AEB6397B-6DEC-577A-9E58-3B26222BB40A" authority="Engl. (Engler 1894: 69)" authorityName="Engl. (Engler" authorityYear="1894" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna holstii" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">Ochna holstii Engl. (Engler 1894: 69)</taxonomicName>
</paragraph>
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<paragraph id="F64594776468CA6790AF369176CBBFA2" pageId="0" pageNumber="174">
<figureCitation id="209E88CD67A3442C4847A3CA3D10AB33" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Ochna, habit and leaf margin. A. Ochna stolzii (from Richards 22529). B. Ochna oxyphylla (from Carmichael 354). C - D. Ochna holstii (C from Maas Geesteranus 4971, D from Pierlot 1731). Drawn by Andrew Brown." figureDoi="10.5091/plecevo.85589.figure2" httpUri="https://binary.pensoft.net/fig/843991" pageId="0" pageNumber="174">Figs 2C-D</figureCitation>
<figureCitation id="E2EF8A870CF5D21D5C75261F201ADF45" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dissection of rehydrated drupelets shows that there are no distinguishing morphological characters in the embryo of any of the mentioned taxa, particularly separating Ochna holstii I and II. A. Ochna holstii II. B. Ochna holstii I. C. Ochna stolzii. D. Ochna oxyphylla." figureDoi="10.5091/plecevo.85589.figure4" httpUri="https://binary.pensoft.net/fig/843993" pageId="0" pageNumber="174">, 4A-B</figureCitation>
<figureCitation id="AF00C71F7DDC7D76E03EE9AA2B0B8649" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution map of Ochna holstii from georeferenced herbarium specimens in Africa." figureDoi="10.5091/plecevo.85589.figure6" httpUri="https://binary.pensoft.net/fig/843995" pageId="0" pageNumber="174">, 6</figureCitation>
</paragraph>
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<paragraph id="9A6CC881D812CA12243D2BD40E5FB772" pageId="0" pageNumber="174">
<taxonomicName id="5915C43F67C20A1B97BA10A625CD11B6" class="Magnoliopsida" family="Ochnaceae" genus="Diporidium" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Diporidium holstii" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">Diporidium holstii</taxonomicName>
(Engl.) Tiegh. (
<bibRefCitation id="E31041544F88CC36833171B32DA9378D" author="Tieghem, P van" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="174" refId="B72" refString="Tieghem, P van, 1902. Sur les Ochnaceae. Annales des Sciences Naturelles, Botanique, Serie 8 16: 161-416." title="Sur les Ochnaceae. Annales des Sciences Naturelles, Botanique, Serie 8 16: 161 - 416." year="1902">Tieghem 1902</bibRefCitation>
: 356) - Type: same as for 
<taxonomicName id="DD80126D8FFB1267404E5C8922E62B4F" authorityName="Engl. (Engler" authorityYear="1894" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna holstii" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">Ochna holstii</taxonomicName>
.
</paragraph>
<paragraph id="F3B2C54327C6119E8124EAFBD29C5AE6" pageId="0" pageNumber="174">
<taxonomicName id="86DC52B26984C1CBCB948AFD2DF24F3D" class="Magnoliopsida" family="Ochnaceae" genus="Biramella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Biramella holstii" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">Biramella holstii</taxonomicName>
(Engl.) Tiegh. (
<bibRefCitation id="03F05FAB572F18484A194DAE53639A6E" author="Tieghem, P van" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="174" refId="B73" refString="Tieghem, P van, 1903. Nouvelles observations sur les Ochnacees. Annales des Sciences Naturelles, Botanique, Serie 8 18: 1-60." title="Nouvelles observations sur les Ochnacees. Annales des Sciences Naturelles, Botanique, Serie 8 18: 1 - 60." year="1903">Tieghem 1903</bibRefCitation>
: 97) - Type: same as for 
<taxonomicName id="30088204D5D8992410381C85A7B1E47C" authorityName="Engl. (Engler" authorityYear="1894" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna holstii" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">Ochna holstii</taxonomicName>
.
</paragraph>
<paragraph id="163C26FDD8FB5E73CA9483999DF10C87" pageId="0" pageNumber="174">
<taxonomicName id="6EB94BA730762BEDC4F4D58BCCAFBD01" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna prunifolia" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="prunifolia">Ochna prunifolia</taxonomicName>
Engl. (
<bibRefCitation id="5D41737F166618EEC92252ABC259F5F7" author="Engler, A" journalOrPublisher="Abhandlungen der Koeniglichen Akademie der Wissenschaften zu Berlin" pageId="0" pageNumber="174" refId="B28" refString="Engler, A, 1895. Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete. Teil C. Geographische Verlagshandlung D. Reimer, Berlin, 1-433." title="Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete. Teil C. Geographische Verlagshandlung D. Reimer, Berlin, 1 - 433." year="1895">Engler 1895</bibRefCitation>
: 273; 
<bibRefCitation id="831C9D28EC5D95C8884E57FC77D119E3" author="Gilg, E" journalOrPublisher="Botanische Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="0" pageNumber="174" pagination="231 - 275" refId="B34" refString="Gilg, E, 1903. Ochnaceae africanae. Botanische Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 33: 231 - 275" title="Ochnaceae africanae." volume="33" year="1903">Gilg 1903</bibRefCitation>
: 236; 
<bibRefCitation id="5A03195989DEC835A39F5A82D909BB17" author="Brenan, JPM" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="174" refId="B11" refString="Brenan, JPM, Greenway, PJ, 1949. Check-lists of the Forest Trees and Shrubs of the British Empire. No. 5. Tanganyika Territory. Part II. Imperial Forestry Institute, Oxford, 1-664." title="Check-lists of the Forest Trees and Shrubs of the British Empire. No. 5. Tanganyika Territory. Part II. Imperial Forestry Institute, Oxford, 1 - 664." year="1949">Brenan and Greenway 1949</bibRefCitation>
: 384) - Type: TANZANIA • Rombo Mku; 1894; 
<emphasis id="4950E09475E7B8641183E9F356A3BD22" italics="true" pageId="0" pageNumber="174">Volkens 1960</emphasis>
; lectotype (
<emphasis id="FA3FFB05D0E6D659E8D43A510E1F17CB" bold="true" pageId="0" pageNumber="174">designated here</emphasis>
): E! [E00200459]; syntype: B† • KENYA • Kitui; 
<emphasis id="C077BF58D2747D06F15D91E295E80C84" italics="true" pageId="0" pageNumber="174">Hildebrandt 2825</emphasis>
; syntype: B† • TANZANIA • Mkulia; 
<emphasis id="47DD361AA6ADEE0498DBF6258ACA408A" italics="true" pageId="0" pageNumber="174">Volkens s.n.</emphasis>
; syntype: B†.
</paragraph>
<paragraph id="EF9CE8DE1AD940CF106E58A857A7D915" pageId="0" pageNumber="174">
<taxonomicName id="947751A50D588F7B14C605D238883A45" class="Magnoliopsida" family="Ochnaceae" genus="Diporidium" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Diporidium prunifolium" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="prunifolium">Diporidium prunifolium</taxonomicName>
(Engl.) Tiegh. (
<bibRefCitation id="14C2C29D19C3E3300D0B4532BB5D6354" author="Tieghem, P van" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="174" refId="B72" refString="Tieghem, P van, 1902. Sur les Ochnaceae. Annales des Sciences Naturelles, Botanique, Serie 8 16: 161-416." title="Sur les Ochnaceae. Annales des Sciences Naturelles, Botanique, Serie 8 16: 161 - 416." year="1902">Tieghem 1902</bibRefCitation>
: 356) - Type: same as for 
<taxonomicName id="49814F7A8037A47FD49F93A21FEC74C4" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna prunifolia" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="prunifolia">Ochna prunifolia</taxonomicName>
.
</paragraph>
<paragraph id="829DE7DF6F1E9801333A8C910993E069" pageId="0" pageNumber="174">
<taxonomicName id="76B8DB55FACC474A0A1451C6980BB2AF" authorityName="Baker" authorityYear="1897" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna longipes" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="longipes">Ochna longipes</taxonomicName>
Baker (
<bibRefCitation id="353914BD13C079AD40F1005F1A668D9C" author="Baker, JG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="174" refId="B3" refString="Baker, JG, 1897. Ochna longipes. In: [Various authors] Diagnoses Africanae, X. Bulletin of Miscellaneous Information, Royal Gardens, Kew 1897(128-129): 247." title="Ochna longipes. In: [Various authors] Diagnoses Africanae, X. Bulletin of Miscellaneous Information, Royal Gardens, Kew 1897 (128 - 129): 247." year="1897">Baker 1897</bibRefCitation>
: 247; 
<bibRefCitation id="DB989A7E595DD2BB9769EF23C95BF560" author="Brenan, JPM" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="174" pagination="191 - 256" refId="B10" refString="Brenan, JPM, 1953. Plants collected by the Vernay Nyasaland expedition of 1946. Memoirs of the New York Botanical Garden 8: 191 - 256" title="Plants collected by the Vernay Nyasaland expedition of 1946." volume="8" year="1953">Brenan 1953</bibRefCitation>
: 234; 
<bibRefCitation id="0500B405FF1BC6B17686BA3BDB8B4BDB" author="White, F" journalOrPublisher="Opera Botanica" pageId="0" pageNumber="174" refId="B81" refString="White, F, 1962. Forest Flora of Northern Rhodesia. Oxford University Press, Oxford, 1-455." title="Forest Flora of Northern Rhodesia. Oxford University Press, Oxford, 1 - 455." year="1962">White 1962</bibRefCitation>
: 251) - Type: MALAWI • Mt Malosa near Zomba; 1896: 
<emphasis id="28ECF3C2E6848FD1163E1ED9F38087D9" italics="true" pageId="0" pageNumber="174">Whyte 429</emphasis>
; holotype: K! [K000431165].
</paragraph>
<paragraph id="13F3A6DC3FCAB19F6F7E03E3564FD49F" pageId="0" pageNumber="174">
<taxonomicName id="951E056B72648EFD6EF4C1E2A15ED309" authorityName="Baker" authorityYear="1897" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna shirensis" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="shirensis">Ochna shirensis</taxonomicName>
Baker (
<bibRefCitation id="E39F41DF0EA0B24A25CCBE326734DC37" author="Baker, JG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="174" refId="B3" refString="Baker, JG, 1897. Ochna longipes. In: [Various authors] Diagnoses Africanae, X. Bulletin of Miscellaneous Information, Royal Gardens, Kew 1897(128-129): 247." title="Ochna longipes. In: [Various authors] Diagnoses Africanae, X. Bulletin of Miscellaneous Information, Royal Gardens, Kew 1897 (128 - 129): 247." year="1897">Baker 1897</bibRefCitation>
: 247) - Type: MALAWI • Mt Zomba &amp; Mt Malosa; 1896: 
<emphasis id="722F617A86BF62CB8B9C62C4D3F0B2A6" italics="true" pageId="0" pageNumber="174">Whyte 430</emphasis>
; holotype: K! [K000431163].
</paragraph>
<paragraph id="88611F59F052CC972042C7278AA5D380" pageId="0" pageNumber="174">
<taxonomicName id="08BEFA5130CE61656BE498B35B7B23A6" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna acutifolia" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="acutifolia">Ochna acutifolia</taxonomicName>
Engl. (
<bibRefCitation id="777F6218F0345FD4241427BDD1C6A8A4" author="Engler, A" journalOrPublisher="Botanische Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="0" pageNumber="174" pagination="385 - 510" refId="B29" refString="Engler, A, 1900. Berichte ueber die botanischen Ergebnisse der Nyassa-See und Kinga-Gebirgs Expedition der Hermann und Elise geb. Heckmann-Wentzel-Stiftung. III. Die von W. Goetze und Dr. Stuhlmann im Ulugurugebirge, sowie die von W. Goetze in der Kisaki- und Khutu-Steppe und in Uhehe gesammelten Pflanzen. Botanische Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 28: 385 - 510" title="Berichte ueber die botanischen Ergebnisse der Nyassa-See und Kinga-Gebirgs Expedition der Hermann und Elise geb. Heckmann-Wentzel-Stiftung. III. Die von W. Goetze und Dr. Stuhlmann im Ulugurugebirge, sowie die von W. Goetze in der Kisaki- und Khutu-Steppe und in Uhehe gesammelten Pflanzen." volume="28" year="1900">Engler 1900</bibRefCitation>
: 433) - Type: TANZANIA • West Usambaras; 
<emphasis id="4124B2B8EF306FC3101D128AA73114E9" italics="true" pageId="0" pageNumber="174">Buchwald 350</emphasis>
; lectotype (
<emphasis id="F82B3D09F119BBF555E006143434A892" bold="true" pageId="0" pageNumber="174">designated here</emphasis>
): K! [K000431145]; isolectotype: BM; syntype: B† • TANZANIA • Uluguru, near Nghweme; 
<emphasis id="B541205CE104A793DCD6D60DF6F1B9FC" italics="true" pageId="0" pageNumber="174">Stuhlmann 8852</emphasis>
; syntype: B†.
</paragraph>
<paragraph id="DF52B89D46331698EF7BCE30CE1E5707" pageId="0" pageNumber="174">
<taxonomicName id="F8F09ED7C85183C7770965037983A30F" class="Magnoliopsida" family="Ochnaceae" genus="Diporidium" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Diporidium acutifolium" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="acutifolium">Diporidium acutifolium</taxonomicName>
(Engl.) Tiegh. (
<bibRefCitation id="A0803E04FF904AB315B88F9AF379C81A" author="Tieghem, P van" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="174" refId="B72" refString="Tieghem, P van, 1902. Sur les Ochnaceae. Annales des Sciences Naturelles, Botanique, Serie 8 16: 161-416." title="Sur les Ochnaceae. Annales des Sciences Naturelles, Botanique, Serie 8 16: 161 - 416." year="1902">Tieghem 1902</bibRefCitation>
: 356) - Type: same as for 
<taxonomicName id="F61BE9026E664805BD86258B3764E749" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna acutifolia" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="acutifolia">Ochna acutifolia</taxonomicName>
.
</paragraph>
<paragraph id="C41FF56F86ED4504B92CA23152437C41" pageId="0" pageNumber="174">
<taxonomicName id="019E861A3B47804EB2AB3F223C77240B" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna densicoma" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="densicoma">Ochna densicoma</taxonomicName>
Engl. &amp; Gilg (
<bibRefCitation id="17529A02CDBD4F0EFF2232E6A61DEAF9" author="Gilg, E" journalOrPublisher="Botanische Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="0" pageNumber="174" pagination="231 - 275" refId="B34" refString="Gilg, E, 1903. Ochnaceae africanae. Botanische Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 33: 231 - 275" title="Ochnaceae africanae." volume="33" year="1903">Gilg 1903</bibRefCitation>
: 241; 
<bibRefCitation id="954E4AC85626028041D1E12D69DC4075" author="Brenan, JPM" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="174" refId="B11" refString="Brenan, JPM, Greenway, PJ, 1949. Check-lists of the Forest Trees and Shrubs of the British Empire. No. 5. Tanganyika Territory. Part II. Imperial Forestry Institute, Oxford, 1-664." title="Check-lists of the Forest Trees and Shrubs of the British Empire. No. 5. Tanganyika Territory. Part II. Imperial Forestry Institute, Oxford, 1 - 664." year="1949">Brenan and Greenway 1949</bibRefCitation>
: 382) - Type: TANZANIA • East Usambaras, Nderema; 1901; 
<emphasis id="CA844DA488F7402BBAB43C9DE0D51DC9" italics="true" pageId="0" pageNumber="174">Scheffler 169</emphasis>
; holotype: B†; lectotype (
<emphasis id="09E7719E6B7908241C6ED53F658471AD" bold="true" pageId="0" pageNumber="174">designated here)</emphasis>
: EA! [EA000002025]; isolectotypes: BM [BM000798536], BR [BR0000006266356], E! [E00200461], K! [K000431144].
</paragraph>
<paragraph id="3FF0F3DA16B951798C8BD000156A4003" pageId="0" pageNumber="174">
<taxonomicName id="22C725B69A39DC49058EE355860640BE" class="Magnoliopsida" family="Ochnaceae" genus="Ochnella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochnella densicoma" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="densicoma">Ochnella densicoma</taxonomicName>
(Engl. &amp; Gilg) Tiegh. (
<bibRefCitation id="CA21320BE14C7FA38BC228B321E5F58F" author="Tieghem, P van" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="174" refId="B73" refString="Tieghem, P van, 1903. Nouvelles observations sur les Ochnacees. Annales des Sciences Naturelles, Botanique, Serie 8 18: 1-60." title="Nouvelles observations sur les Ochnacees. Annales des Sciences Naturelles, Botanique, Serie 8 18: 1 - 60." year="1903">Tieghem 1903</bibRefCitation>
: 40) - Type: same as for 
<taxonomicName id="7346FEC8BE7525F883BC1E60ED8A3B9E" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna densicoma" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="densicoma">Ochna densicoma</taxonomicName>
.
</paragraph>
<paragraph id="AF06C997760F9025BB4EA1BB4A31CB64" pageId="0" pageNumber="174">
<taxonomicName id="A08EBB5A3E39023F927A3A0A14F04CB7" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna chirindica" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="chirindica">Ochna chirindica</taxonomicName>
Baker f. (
<bibRefCitation id="D45459689431026005A43EF5DC6B9B0D" DOI="https://doi.org/10.1111/j.1095-8339.1911.tb00869.x" author="Rendle, AB" journalOrPublisher="The Journal of the Linnean Society, Botany" pageId="0" pageNumber="174" pagination="1 - 245" refId="B62" refString="Rendle, AB, Baker, EG, Moore, S, Gepp, A, 1911. A contribution to our knowledge of the flora of Gazaland: being an account of the collections made by C. F. M. Swynnerton Esq., F.L.S. The Journal of the Linnean Society, Botany 40: 1 - 245, DOI: https://doi.org/10.1111/j.1095-8339.1911.tb00869.x" title="A contribution to our knowledge of the flora of Gazaland: being an account of the collections made by C. F. M. Swynnerton Esq., F. L. S." url="https://doi.org/10.1111/j.1095-8339.1911.tb00869.x" volume="40" year="1911">Rendle et al. 1911</bibRefCitation>
: 37) - Type: ZIMBABWE • Melsetter District, Chirinda Forest; 1905; 
<emphasis id="26CCE77583044B05662D6FDF5B8B0400" italics="true" pageId="0" pageNumber="174">Swynnerton 106</emphasis>
; holotype: K! [K000431166]; isotype BM [BM000798543].
</paragraph>
<paragraph id="6143A494DDA47733FCB3989894963838" pageId="0" pageNumber="174">
<taxonomicName id="4957C39BA81DE224D4EB547E07BF1630" authorityName="Sleumer" authorityYear="1934" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Ochna keniensis" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="keniensis">Ochna keniensis</taxonomicName>
Sleumer (
<bibRefCitation id="1BF88FE4834D603FDAA0BC539F41981A" author="Sleumer, H" journalOrPublisher="Botanischen Gartens und Museums zu Berlin" pageId="0" pageNumber="174" pagination="65 - 70" refId="B70" refString="Sleumer, H, 1934. Vermischte Diagnosen. Notizblatt des Koenigl. Botanischen Gartens und Museums zu Berlin 12: 65 - 70" title="Vermischte Diagnosen. Notizblatt des Koenigl." volume="12" year="1934">Sleumer 1934</bibRefCitation>
: 69; 
<bibRefCitation id="B82C09330C16B65F15D9C4112CEA2999" author="Dale, IR" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="174" refId="B15" refString="Dale, IR, Greenway, PJ, 1961. Kenya Trees and Shrubs. Buchanan's Kenya Estate Ltd., Nairobi in assoc. with Hatchards, London, 1-654." title="Kenya Trees and Shrubs. Buchanan's Kenya Estate Ltd., Nairobi in assoc. with Hatchards, London, 1 - 654." year="1961">Dale and Greenway 1961</bibRefCitation>
: 338; 
<bibRefCitation id="E2CD29EBB204F1A2BDB3DDC0CAF112C8" author="Beentje, H" journalOrPublisher="Journal of Computational Biology" pageId="0" pageNumber="174" refId="B7" refString="Beentje, H, 1994. Kenya trees, shrubs, and lianas. National Museums of Kenya, Nairobi, 1-762." title="Kenya trees, shrubs, and lianas. National Museums of Kenya, Nairobi, 1 - 762." year="1994">Beentje 1994</bibRefCitation>
: 121) - Type: KENYA • Mt Rupingazi River; 1922; 
<emphasis id="352E7A376972E55BF86E2D6D8171F221" italics="true" pageId="0" pageNumber="174">Fries &amp; Fries 2031</emphasis>
; holotype: UPS [fide 
<bibRefCitation id="B0664AD5A2B3C7032B4AC9A1B060785B" author="Verdcourt, B" editor="Beentje, HJ" journalOrPublisher="Royal Botanic Gardens, Kew" pageId="0" pageNumber="174" pagination="1 - 64" refId="B78" refString="Verdcourt, B, 2005. Ochnaceae. In: Beentje, HJ, Ghazanfar, SA, Eds., Flora of Tropical East Africa. Royal Botanic Gardens, Kew: 1 - 64" title="Ochnaceae." volumeTitle="Flora of Tropical East Africa." year="2005">Verdcourt 2005</bibRefCitation>
]; isotype: S [S-G-4319].
</paragraph>
</subSubSection>
<subSubSection id="B95C0C983C3F89CCB18EF29590A1A0A5" pageId="0" pageNumber="174" type="materials_examined">
<paragraph id="510DC3C9681B8127AFCDDF21E5FE7A32" pageId="0" pageNumber="174">
<typeStatus id="E769D656B0408DFF1E968980FC12589D">Type</typeStatus>
.
</paragraph>
<paragraph id="A0A4336889EE07C7B33840FC8CA24F49" pageId="0" pageNumber="174">
<materialsCitation id="4723F12C4E80D65AE97B82522FF90AEC" ID-GBIF-Occurrence="4098977308" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="TANZANIA" location="West Usambaras" specimenCount="1" typeStatus="holotype">
<collectingCountry id="41A0BE8D519C6F012509C4855DA6D50B" name="Tanzania">TANZANIA</collectingCountry>
• 
<location id="7DEC4A7B35CBAA22F0F6710F9B81365B" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:7DEC4A7B35CBAA22F0F6710F9B81365B" country="TANZANIA" name="West Usambaras">West Usambaras</location>
, Mbalu; 1893; 
<emphasis id="836021808F754FE642F17B258CEC9128" italics="true" pageId="0" pageNumber="174">Holst 2601</emphasis>
; 
<typeStatus id="7CE976B51FD5FAF476A1985EC333B81E">holotype</typeStatus>
: B†; 
<typeStatus id="06FEC69DCC52DA4C4C468FBD258AE6F5">lectotype</typeStatus>
(
<emphasis id="52464310094E90E487A546952896B384" bold="true" pageId="0" pageNumber="174">designated here</emphasis>
): K! [K000431146]; isolectotypes: BR [BR0000006265694], HBG [HBG509293]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D826F33756437D75B3F72A8AE4A00D76" pageId="0" pageNumber="174" type="description">
<paragraph id="FE7AD6AB87884D00E671BF17EB7AE915" pageId="0" pageNumber="174">Description.</paragraph>
<paragraph id="760DF1AF40ACA9E6D40729B84347260D" pageId="0" pageNumber="174">Small to large tree 4-27 m tall, usually over 10 m tall, rarely with shrubby forms 1.5-3 m tall. Bark grey-brown. Stems dark grey, sometimes peeling, with white lenticels, these sometimes sparse; young stems dark brown, with white lenticels, glabrous. Stipules brown, linear, sometimes persistent to deciduous, 2-15 mm long. Leaves green, often drying brown or rarely blue-black, chartaceous, glabrous, lanceolate to oblanceolate, 4-12(-16.5) cm long, 1-5(-5.5) cm wide; leaf base attenuate-cuneate (rarely rounded), leaf apex attenuate, margins serrate; lateral veins 20-28, appearing at right angles to midrib, tertiary vernation reticulate; petiole 0.5-3.5 mm long; buds brown, large, up to 7.5(-9) mm long. Flowers in racemes with 6-16 flowers; rachis 0.5-2 cm long, glabrous; pedicels 1.5-4 cm long, articulated at or near the base, or up to 3(-10) mm above the base, glabrous throughout. Sepals green to brown in flower, oblong to elliptic, 8-10 mm long, 4-6 mm wide in flower, turning red in fruit, then 11-14 mm long, 4-8(-10) mm wide. Petals yellow, often pale, obovate, 8.5-10.5 mm long, 3.5-4 mm wide. Anthers dehiscing by longitudinal slits, 1.5-2 mm long; filaments 2-3.5 mm long. Carpels 5; styles united to apex, 4-7 mm long Drupelets black, ellipsoid, 8-13 mm long, 4-6 cm wide, attached at the base.</paragraph>
</subSubSection>
<subSubSection id="588583F9BA1785C0A869A5213C4E36FB" pageId="0" pageNumber="174" type="distribution">
<paragraph id="82443351395B2BE52FAB2E892BE8CC5F" pageId="0" pageNumber="174">Distribution.</paragraph>
<paragraph id="2E23AADEC6C7E9BA8AFCA423007A3358" pageId="0" pageNumber="174">
Widespread in montane regions of eastern Africa: South Sudan, Ethiopia, Democratic Republic of the Congo (DRC), Uganda, Kenya, Tanzania, Burundi, Rwanda, Mozambique, Malawi, Zambia, Zimbabwe, South Africa (Fig. 
<figureCitation id="2F05DA52A3DF4B9C6879B9847B53499E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution map of Ochna holstii from georeferenced herbarium specimens in Africa." figureDoi="10.5091/plecevo.85589.figure6" httpUri="https://binary.pensoft.net/fig/843995" pageId="0" pageNumber="174">6</figureCitation>
).
</paragraph>
<caption id="B0F6E9F90A8E4CD47FDA6B6BE4CCEA74" doi="10.5091/plecevo.85589.figure6" httpUri="https://binary.pensoft.net/fig/843995" pageId="0" pageNumber="174" start="Figure 6" startId="F6">
<paragraph id="FD5C6D7D0ABE3C61F455282AE57CEB4A" pageId="0" pageNumber="174">
<emphasis id="E9C41DE6E97D0796290E2872AED7F3B9" bold="true" pageId="0" pageNumber="174">Figure 6.</emphasis>
Distribution map of 
<taxonomicName id="B20FAE51ACEF23F9A49709DA214C8817" authorityName="Engl. (Engler" authorityYear="1894" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochna holstii" order="Gentianales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">
<emphasis id="9C762FF950F37D48A3F11C1E37803E24" italics="true" pageId="0" pageNumber="174">Ochna holstii</emphasis>
</taxonomicName>
from georeferenced herbarium specimens in Africa.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2F4436521F212FDFCF11E7A186E27343" pageId="0" pageNumber="174" type="habitat">
<paragraph id="E54CB468C90735393C9544EFF2BA0555" pageId="0" pageNumber="174">Habitat.</paragraph>
<paragraph id="A80B172F24EC1EE9785FAA5A4D302CFF" pageId="0" pageNumber="174">Usually found in upland and montane (Afromontane) moist forests as a medium- to large-sized tree; occasionally recorded in more open habitats including montane grassland and amongst rocky boulders. Altitude: 1000-4600 m.</paragraph>
</subSubSection>
<subSubSection id="D15D09AF78335192F643D3DE73E921F6" pageId="0" pageNumber="174" type="phenology">
<paragraph id="4F2A1EA11AE1210F2BBF82C5A83CA0A8" pageId="0" pageNumber="174">Phenology.</paragraph>
<paragraph id="198840B6004FCD96646A6A1515E1A135" pageId="0" pageNumber="174">Most flowering and fruiting specimens are collected from October to March. This corresponds to the shorter rainfall season in East Africa (October to December) and the longer rainfall season in southern Africa (November to March).</paragraph>
</subSubSection>
<subSubSection id="0931C722EAC81DB1E9CD50210D3F64B7" pageId="0" pageNumber="174" type="preliminary iucn conservation assessment">
<paragraph id="7C196881958E20B1E47D04E841E852EB" pageId="0" pageNumber="174">Preliminary IUCN conservation assessment.</paragraph>
<paragraph id="87E1CB3A5CE5CC50925F65FF1E1B7DB1" pageId="0" pageNumber="174">
<taxonomicName id="9AB187626605AC4F1C6B9872A82E3178" authorityName="Engl. (Engler" authorityYear="1894" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochna holstii" order="Gentianales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="holstii">
<emphasis id="D4645FE27027F7972C6B2A3DC44F2308" italics="true" pageId="0" pageNumber="174">Ochna holstii</emphasis>
</taxonomicName>
is a medium- to large-sized tree (rarely in shrubby form), widespread across montane forests in East and Southern Africa. The estimated extent of occurrence (EOO) is 3,888.294 km2, and the area of occupancy (AOO) is 548 km2. Forest loss due to agricultural expansion and wood harvesting, particularly at its lower altitudinal limit, may have an impact on some subpopulations. Although the species falls within the range of values for the vulnerable category due to its AOO for criteria B2, the species is known from substantially more than 10 threat-defined locations, it is assessed as Least Concern (LC).
</paragraph>
</subSubSection>
<subSubSection id="E70F4806363BA01D39A915FD0A55DB2F" pageId="0" pageNumber="174" type="materials_examined">
<paragraph id="028BA04E83D8DE438BE90D55DE01A0F7" pageId="0" pageNumber="174">Additional material examined.</paragraph>
<paragraph id="2B4D47FA98A9B12049128E26BDB08A27" pageId="0" pageNumber="174">
<collectingCountry id="DEAF548381A13442533F8DF9DAB951CA" name="South Sudan">SOUTH SUDAN</collectingCountry>
- 
<emphasis id="444D67BE842B34EE7E1658B373DABB96" bold="true" pageId="0" pageNumber="174">Equatoria Province</emphasis>
• Mt Lotuke, Didina Mts; 
<quantity id="EC8F62B9D54D83CDFC8DC3291DABF7C6" metricMagnitude="3" metricUnit="m" metricValue="1.9812" unit="ft" value="6500.0">6500 ft</quantity>
; 
<date id="0ADCC8315ACEE36C04F34E749468EB0B" value="1939-04-20">20 Apr. 1939</date>
; fl.; 
<emphasis id="8FA79B64F5B738AC5B7111E52D5AE002" italics="true" pageId="0" pageNumber="174">Myers 10959</emphasis>
; K [K001383230] • Imatong Mountains, Bushbuck Hill; 
<quantity id="C76413E012CADBB356B3A7FBD3D8E839" metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">2300 m</quantity>
; 
<date id="54929A9912F685A2C9D2DD042D9450AC" value="1982-02-21">21 Feb. 1982</date>
; fl.; 
<emphasis id="ACECB9E7D769C84EBB00B12D0B1222A4" italics="true" pageId="0" pageNumber="174">Friis &amp; Vollesen 982</emphasis>
; K [K001383227].
</paragraph>
<paragraph id="8898FD15344A0F4D9BA91FD2821C3199" pageId="0" pageNumber="174">
<materialsCitation id="30BEA8152C15D121560F332175ACE24D" ID-GBIF-Occurrence="4098977375" collectingDate="1976-06" country="ETHIOPIA" elevation="1700" latitude="5.5" location="Mogada" longLatPrecision="1305" longitude="38.333332" specimenCount="1" stateProvince="Oromia">
<collectingCountry id="8CD2FE9A9E2C26FE210347756D712645" name="Ethiopia">ETHIOPIA</collectingCountry>
- 
<emphasis id="478836EAB4594D6F29CE219A8E4E949D" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="5FBA97FCA9ABD11D648B15A92FF862A5" country="Ethiopia" name="Oromiya">Oromia</collectingRegion>
</emphasis>
• 
<location id="299AE96E3174EDAF893ACD6ECABF1CDF" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:299AE96E3174EDAF893ACD6ECABF1CDF" country="ETHIOPIA" latitude="5.5" longLatPrecision="1305" longitude="38.333332" name="Mogada" stateProvince="Oromia">Mogada</location>
; 
<geoCoordinate id="32C96DA368CC6E6D20233CEE53F21F5C" degrees="5" direction="north" minutes="30" orientation="latitude" precision="925" value="5.5">5°30'N</geoCoordinate>
, 
<geoCoordinate id="538D4E78DFFE3514D4BDE4F491FDDA18" degrees="38" direction="east" minutes="20" orientation="longitude" precision="925" value="38.333332">38°20'E</geoCoordinate>
; 
<quantity id="857DE920606476A2CF2E4CE88052C8D1" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="0A7FF121EF6B013356D8B4A39F5E9733" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</elevation>
</quantity>
; 
<collectingDate id="2B6F5BAAB862DB7F597215DEE8CE8D38" value="1976-06">Jun. 1976</collectingDate>
; fr.; 
<emphasis id="70D580C87442EEF8402FB951F79B14D5" italics="true" pageId="0" pageNumber="174">Haile 696</emphasis>
; K [K001271514]
</materialsCitation>
.
</paragraph>
<paragraph id="0CC950A2CCB0104140E5AA6840D5DA49" pageId="0" pageNumber="174">
<materialsCitation id="B69D40DD0C4C010B08AECB35BC078B4A" ID-GBIF-Occurrence="4098977385" collectingDate="1900-05" collectorName="Whyte" country="KENYA" elevation="2286" location="Mau" municipality="Eldama Ravine" specimenCount="1" stateProvince="Baringo">
<collectingCountry id="5ADAFA47286708D15BA7B2175D19B3DD" name="Kenya">KENYA</collectingCountry>
• 
<emphasis id="CBE272D195865358663BBAA33193C7F3" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="07884E4B9127B382DDFC5EC5E164854F" country="Kenya" name="Baringo">Baringo</collectingRegion>
</emphasis>
• 
<collectingMunicipality id="6FF91DDB31EAC83C2AC4E21C876F1087">Eldama Ravine</collectingMunicipality>
and 
<location id="6E92D13BA6030C15F4931EAE5E476EAD" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:6E92D13BA6030C15F4931EAE5E476EAD" country="KENYA" municipality="Eldama Ravine" name="Mau" stateProvince="Baringo">Mau</location>
; 
<quantity id="D4F1166723771DBD27897F11952F5A9F" metricMagnitude="3" metricUnit="m" metricValue="2.286" unit="ft" value="7500.0">
<elevation id="E630F97CED95AFC4951925E52C85B244" metricMagnitude="3" metricUnit="m" metricValue="2.286" unit="ft" value="7500.0">7500 ft</elevation>
</quantity>
; 
<collectingDate id="0B8861AEEEF5F5CF154DF9194526DB89" value="1900-05">May 1900</collectingDate>
; fr.; 
<emphasis id="73077D08AF2D5FF24D191D711A591F5C" italics="true" pageId="0" pageNumber="174">
<collectorName id="AA061E8703D930A7D9E41FB3316E0328">Whyte</collectorName>
1898
</emphasis>
; K
</materialsCitation>
. - 
<materialsCitation id="F857C5B885D4C023F3B9DD3341ED6D45" ID-GBIF-Occurrence="4098977355" collectingDate="1961-01" collectorName="van Someren EA" country="Kenya" location="Lebetero Hills" municipality="Nguruman Range" specimenCount="1" stateProvince="Narok">
<emphasis id="2DBA5022E9C8C21F5B7F9DA7D8502EEB" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="1F30590F88B28E1AD81909A8370F12CF" country="Kenya" name="Narok">Narok</collectingRegion>
</emphasis>
• 
<collectingMunicipality id="90973EF87DEAAF72733517A2CA008A41">Nguruman Range</collectingMunicipality>
, 
<location id="2619848C5D9D1D4E9736A22BAEF8F1B4" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:2619848C5D9D1D4E9736A22BAEF8F1B4" country="Kenya" municipality="Nguruman Range" name="Lebetero Hills" stateProvince="Narok">Lebetero Hills</location>
; 
<collectingDate id="604870A1D618ECCFAE54CFE91964BAF1" value="1961-01">Jan 1961</collectingDate>
; fr.; 
<emphasis id="9B6E450D1B0DE17003D73C47D8309349" italics="true" pageId="0" pageNumber="174">
<collectorName id="3217E9E81275EE7CA68E6004CCBB1A7F">van Someren EA</collectorName>
12286
</emphasis>
; K [K001082543]
</materialsCitation>
. - 
<materialsCitation id="D5611E013E7F9BF53B9248B17B3C099F" ID-GBIF-Occurrence="4098977344" collectingDate="1949-07-01" collectorName="Maas Geesteranus" country="Kenya" county="Londiani District" elevation="2300" latitude="-0.083333336" location="Camp" longLatPrecision="21" longitude="35.358334" municipality="Tinderet Forest Reserve" specimenCount="1" stateProvince="Kericho">
<emphasis id="00F5D63ED8999D932E25A463531C4561" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="6593BB4E829FFB82A32F3F0BE918924C" country="Kenya" name="Kericho">Kericho</collectingRegion>
</emphasis>
• 
<collectingCounty id="E89BCC73539462CA36061D011DA9D519">Londiani District</collectingCounty>
, 
<collectingMunicipality id="F92D4893DC6329D9601F7832C5A45A1C">Tinderet Forest Reserve</collectingMunicipality>
, between 
<location id="767EAD24A834ABD7F59BE95412FE4E14" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:767EAD24A834ABD7F59BE95412FE4E14" country="Kenya" county="Londiani District" latitude="-0.083333336" longLatPrecision="21" longitude="35.358334" municipality="Tinderet Forest Reserve" name="Camp" stateProvince="Kericho">Camp</location>
4 and 
<location id="FDB548C553F881F367FF2E91F2F1A0D5" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:FDB548C553F881F367FF2E91F2F1A0D5" country="Kenya" county="Londiani District" latitude="-0.083333336" longLatPrecision="21" longitude="35.358334" municipality="Tinderet Forest Reserve" name="Tinderet Triangulation Point" stateProvince="Kericho">Tinderet Triangulation Point</location>
; 
<geoCoordinate id="5A5529696E4535599AA388279E5BD3B2" degrees="0" direction="south" minutes="05" orientation="latitude" precision="15" seconds="00" value="-0.083333336">
0°05 
<normalizedToken id="4B0DB365173B7A17A0B8E074D21BAA3B" originalValue="’00”">'00&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate id="072EA73800095831E6C8633495536FC4" degrees="35" direction="east" minutes="21" orientation="longitude" precision="15" seconds="30" value="35.358334">
35°21 
<normalizedToken id="0924B421C6133FBA32CC35B917185114" originalValue="’30”">'30&quot;</normalizedToken>
E
</geoCoordinate>
; 
<quantity id="DD18BB5765C9EF15BB409C7ED5BEDCF7" metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">
<elevation id="1CEE820227A871DF7D2F207FCA8EEFD7" metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">2300 m</elevation>
</quantity>
; 
<collectingDate id="82BD08942DE00DC8FA4D61809BD6B98A" value="1949-07-01">1 Jul. 1949</collectingDate>
; fr.; 
<emphasis id="E22460B4DFA4A57A9545FD3CFFEB4883" italics="true" pageId="0" pageNumber="174">
<collectorName id="2A009403623AA79DDFFE0747BD16DCA6">Maas Geesteranus</collectorName>
5311
</emphasis>
; K [K001082539]
</materialsCitation>
. - 
<materialsCitation id="7426D476BAFF0294AEE1F2635133F18E" ID-GBIF-Occurrence="4098977326" collectingDate="1967-03-26" collectingDateMax="1968-04-23" collectingDateMin="1967-03-26" collectorName="Gillett, Elgon, Hamilton" country="Kenya" elevation="2170" latitude="1.0666667" location="Lorogi forest" longLatPrecision="1308" longitude="36.866665" specimenCount="1" stateProvince="Samburu">
<emphasis id="3EBEC5D4B01A9F5417F2B8646397FF38" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="0948EA9B22320864A1C90221E22F1BCE" country="Kenya" name="Samburu">Samburu</collectingRegion>
</emphasis>
• 
<location id="2D86C481923B8A96A78BDC89FA3C13AD" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:2D86C481923B8A96A78BDC89FA3C13AD" country="Kenya" latitude="1.0666667" longLatPrecision="1308" longitude="36.866665" name="Lorogi forest" stateProvince="Samburu">Lorogi forest</location>
; 
<geoCoordinate id="7CB27C0EB009B28E85D2041B5327A71B" degrees="1" direction="north" minutes="04" orientation="latitude" precision="925" value="1.0666667">1°04'N</geoCoordinate>
, 
<geoCoordinate id="69A0657E64B190F10986AEC6B3F4D83A" degrees="36" direction="east" minutes="52" orientation="longitude" precision="925" value="36.866665">36°52'E</geoCoordinate>
; 
<quantity id="6121FFA42160027560700B1A8FBA31B1" metricMagnitude="3" metricUnit="m" metricValue="2.17" unit="m" value="2170.0">
<elevation id="4766D9F6F7D342E5C75FD1B2681677E0" metricMagnitude="3" metricUnit="m" metricValue="2.17" unit="m" value="2170.0">2170 m</elevation>
</quantity>
; 
<collectingDate id="947A2EC399260F30A065EA1505E34457" value="1967-03-26">26 Mar. 1967</collectingDate>
; fr.; 
<emphasis id="4DAC4CD2281D4067D0EAD1658B6D18ED" italics="true" pageId="0" pageNumber="174">
<collectorName id="70641ADCD5711E4755EDCD566903CB7B">Gillett</collectorName>
18095
</emphasis>
; EA, K [K001082519] • 
<collectorName id="FDF58DC224688440D6182CEA63A649C6">Elgon</collectorName>
; 
<quantity id="BADB83E6283D7B78C8426F7F819CC699" metricMagnitude="3" metricUnit="m" metricValue="2.286" unit="ft" value="7500.0">
<elevation id="6260150FC88F4073E3A70B472E7DBFB5" metricMagnitude="3" metricUnit="m" metricValue="2.286" unit="ft" value="7500.0">7500 ft</elevation>
</quantity>
; 
<collectingDate id="6CE152E4B5116EE778C79FC13AE2FC42" value="1968-04-23">23 Apr. 1968</collectingDate>
; fr.; 
<emphasis id="BB54FCC5D33A0629E5E6D7DFC1C3FD99" italics="true" pageId="0" pageNumber="174">
<collectorName id="E1DF6FAD7CB55242307C5AAAC72779BF">Hamilton</collectorName>
809
</emphasis>
; K [K001082523]
</materialsCitation>
. - 
<materialsCitation id="C2B7810153AB04414CFDA875593A2F16" ID-GBIF-Occurrence="4098977369" collectingDate="1973-02-21" collectorName="Lawton" country="Kenya" location="Naro-Moro" municipality="Mount Kenya Forest Reserve" specimenCount="1" stateProvince="Meru">
<emphasis id="C978A0C82DD5D23DB712E9251E5C93A7" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="E03A63A569CA0C779E1F1247B73A55F5" country="Kenya" name="Meru">Meru</collectingRegion>
</emphasis>
• 
<collectingMunicipality id="ED2C47B30B43D758CE49D9AD158B8F29">Mount Kenya Forest Reserve</collectingMunicipality>
between 
<location id="3BCEF590EB884401ACA59203BAC76562" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:3BCEF590EB884401ACA59203BAC76562" country="Kenya" municipality="Mount Kenya Forest Reserve" name="Naro-Moro" stateProvince="Meru">Naro-Moro</location>
and 
<collectingRegion id="D2E4B0F7D66D423A0C8D72D32DC37D0E" country="Kenya" name="Nyeri">Nyeri</collectingRegion>
; 
<collectingDate id="39C4FBA93C441364080F6B5D08778ECD" value="1973-02-21">21 Feb. 1973</collectingDate>
; fr.; 
<emphasis id="56B64FDBFC6DEFB8370BD72517DAC9B6" italics="true" pageId="0" pageNumber="174">
<collectorName id="83951616AF68987B5E539F763378498C">Lawton</collectorName>
1751
</emphasis>
; K [K001082534]
</materialsCitation>
.
</paragraph>
<paragraph id="5B18F8E37FF558B2D4EAE48436EBAB19" pageId="0" pageNumber="174">
<collectingCountry id="7599DE08779443EA1CF74553CE4616CE" name="Uganda">UGANDA</collectingCountry>
- 
<emphasis id="9DE63921761CFD3EE6939CBB0559AFB6" bold="true" pageId="0" pageNumber="174">Eastern</emphasis>
• N.E. Mt Elgon; 
<quantity id="4C523DE30C8E397105D2F02E67A3B327" metricMagnitude="3" metricUnit="m" metricValue="1.9812" unit="ft" value="6500.0">6500 ft</quantity>
; 
<date id="4FDBB6C61CD9392C7BD925F8FD8F1560" value="1933-05">May 1933</date>
; fr.; 
<emphasis id="11C4A44042291811E87A083A1789D588" italics="true" pageId="0" pageNumber="174">Dale 3121</emphasis>
; K [K001082516; K001082517] • Imatong Mountains; 
<quantity id="7921FC8BC9E81CFA38930B08096E74FB" metricMagnitude="3" metricUnit="m" metricValue="2.37744" unit="ft" value="7800.0">7800 ft</quantity>
; 
<date id="0E7029D2AD9C3FBF5EF946EDADA570F8" value="1938-04">Apr. 1938</date>
; fr.; 
<emphasis id="5F016520CE2BABA676DE1D64C938413B" italics="true" pageId="0" pageNumber="174">Eggeling E3539</emphasis>
; K [K001082514]. - 
<emphasis id="7E2E0BE8387511D5A2C1D07ECE94A683" bold="true" pageId="0" pageNumber="174">Northern</emphasis>
• Dodoth, Kidepo National Park; 
<quantity id="B29C5D1794984A203577A4204A27D8BA" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</quantity>
; 
<date id="B5AFD773559546225B4A136768AEADF3" value="1972-05-16">16 May 1972</date>
; fr.; 
<emphasis id="A76EB30E2D7AFF438B128DF20AF0641B" italics="true" pageId="0" pageNumber="174">Synnott 984</emphasis>
; EA, K [K001082512].
</paragraph>
<paragraph id="45176258E2A37B20F0D73B7A36B21294" pageId="0" pageNumber="174">
<materialsCitation id="2640F147DF7E6F706AF40B86E477F9AF" ID-GBIF-Occurrence="4098977384" collectingDate="1947-01-07" collectorName="Hughes" country="TANZANIA" county="Shume" location="Shume valley" municipality="Korogwe" specimenCount="1" stateProvince="Korogwe District">
<collectingCountry id="F57DBE6621B7DB65456E660A6AEB22F4" name="Tanzania">TANZANIA</collectingCountry>
- 
<emphasis id="0A85BC0502B11301D3601609BE504066" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="AD564BCBE3C3F28645EE7CB9D5C459D5" inferredBySuffix="true">Korogwe District</collectingRegion>
</emphasis>
• 
<collectingCounty id="FB2234843F86456BC1DF37E2BE468BD7">Shume</collectingCounty>
, 
<collectingMunicipality id="08CE2667C7E789360AA28E20EECEFAA8">Korogwe</collectingMunicipality>
, in the 
<location id="50861E55DD28C73DA7B1B100BB3643AD" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:50861E55DD28C73DA7B1B100BB3643AD" country="TANZANIA" county="Shume" municipality="Korogwe" name="Shume valley" stateProvince="Korogwe District">Shume valley</location>
bottom, about 
<quantity id="A433F00B2026E58F705AFE522DA0E83C" metricMagnitude="2" metricUnit="m" metricValue="8.04672" unit="mi" value="0.5">0.5 miles</quantity>
from the 
<location id="B329E71E788280AC56F6DFBF7BAE5F49" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:B329E71E788280AC56F6DFBF7BAE5F49" country="TANZANIA" county="Shume" municipality="Korogwe" name="Forest House" stateProvince="Korogwe District">Forest House</location>
; 
<collectingDate id="3CBA6DEAB5A1E4BB0374541812774ACC" value="1947-01-07">7 Jan. 1947</collectingDate>
; fr.; 
<emphasis id="CA4E8DA580AD01ECFFA4E39CEC6038DF" italics="true" pageId="0" pageNumber="174">
<collectorName id="FFD9C13D9475419FA0339379A715355E">Hughes</collectorName>
16
</emphasis>
; EA, K [K001082587]
</materialsCitation>
. - 
<materialsCitation id="621B8029610ED9C34C4B2D031FD51679" ID-GBIF-Occurrence="4098977303" collectingDate="1932-12-29" collectingDateMax="1962-02-20" collectingDateMin="1932-12-29" collectorName="W. Usambaras, Parry, Near Lushoto, Mgaza" country="Tanzania" elevation="975" location="Derema" municipality="East Usambaras" specimenCount="1" stateProvince="Lushoto District">
<emphasis id="2922D34C329218CBE47BF936E117FEE6" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="B50AC466A836EF53FF3F9C0D18C1002E" inferredBySuffix="true">Lushoto District</collectingRegion>
</emphasis>
<collectingMunicipality id="809D2232939B9CF7DBE7CC42F8222108">East Usambaras</collectingMunicipality>
, 
<location id="B9FD6DCCA2FDC3A544C5DCBC4D48F8DA" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:B9FD6DCCA2FDC3A544C5DCBC4D48F8DA" country="Tanzania" municipality="East Usambaras" name="Derema" stateProvince="Lushoto District">Derema</location>
; 
<quantity id="17B4AA15E28F8A2E556C3F1AFCF22A37" metricMagnitude="2" metricUnit="m" metricValue="9.7536" unit="ft" value="3200.0">
<elevation id="952A06F5D0CC4D4AE2EAF1A0A7AECFB8" metricMagnitude="2" metricUnit="m" metricValue="9.7536" unit="ft" value="3200.0">3200 ft</elevation>
</quantity>
; 
<collectingDate id="3E1F5DAD31B3E64B520D4CA6F7697009" value="1932-12-29">29 Dec. 1932</collectingDate>
; fl.; 
<emphasis id="D512192A4928F5E7EB4586B611CB17B8" italics="true" pageId="0" pageNumber="174">Greenway 3308</emphasis>
; EA, K[K001082578] • 
<collectorName id="5D4B4481E195F8E407E97D28FF3FFEBF">W. Usambaras</collectorName>
; 
<quantity id="BB40820855D91C0CB8A5F8A2B5651AC8" metricMagnitude="3" metricUnit="m" metricValue="1.8897599999999997" unit="ft" value="6200.0">
<elevation id="544F3D1AE82AFFC624BFFD503D7E3CBF" metricMagnitude="3" metricUnit="m" metricValue="1.8897599999999997" unit="ft" value="6200.0">6200 ft</elevation>
</quantity>
; 
<collectingDate id="2EB1148AEA0AC91DDAEA5DCD01E3470E" value="1952-01-16">16 Jan. 1952</collectingDate>
; fr.; 
<emphasis id="7B13ECB336C180220B2A73570B455D08" italics="true" pageId="0" pageNumber="174">
<collectorName id="D50BD4ED0F764893A0DA753E542BA226">Parry</collectorName>
105
</emphasis>
; EA, K [K001082589] • 
<collectorName id="374F9E67379FCD4D2F363A0B38F3A8C8">Near Lushoto</collectorName>
water supply; 
<collectingDate id="C28D860E3D9933FEC5D03EF7F6F3BD77" value="1962-02-20">20 Feb. 1962</collectingDate>
; fl.; 
<emphasis id="4ED73BD75ABFAA094B13D6F2F9EFD12C" italics="true" pageId="0" pageNumber="174">
<collectorName id="DC4CE3C8E40ADD050B97DD3A8352DD15">Mgaza</collectorName>
452
</emphasis>
; EA, K [K001082591]
</materialsCitation>
. - 
<materialsCitation id="31BA57E08997AC18C2EAE5A092C04AFE" ID-GBIF-Occurrence="4098977378" collectingDate="2020-01-26" collectorName="Shah TS" country="Tanzania" elevation="1823" latitude="-9.2368" location="Ruhuji Waterfalls" longLatPrecision="1" longitude="34.76235" specimenCount="1" stateProvince="Njombe District">
<emphasis id="9B741609D42637F46CFEC5610B425B43" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="6AF873EBA5B2C536DC2CCCE274165959" country="Tanzania" name="Njombe">Njombe District</collectingRegion>
</emphasis>
• 
<location id="6ECB549592D17F39C1195A2157B743F7" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:6ECB549592D17F39C1195A2157B743F7" country="Tanzania" latitude="-9.2368" longLatPrecision="1" longitude="34.76235" name="Ruhuji Waterfalls" stateProvince="Njombe District">Ruhuji Waterfalls</location>
; 
<geoCoordinate id="C11C63EF96B96224724DF65B841D0CD8" degrees="9" direction="south" minutes="14" orientation="latitude" precision="1" seconds="12.48" value="-9.2368">
9°14 
<normalizedToken id="656B26060147C1FDA1999007713F1EA0" originalValue="’12.48”">'12.48&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate id="0C0E4254F19A0DB1E79793CA26C73F95" degrees="34" direction="east" minutes="45" orientation="longitude" precision="1" seconds="44.46" value="34.76235">
34°45 
<normalizedToken id="55A3AB55FDABFA8ECB991F761EAB8F51" originalValue="’44.46”">'44.46&quot;</normalizedToken>
E
</geoCoordinate>
; 
<quantity id="8D314A5AFDF8AC97FF85F614B4BF878E" metricMagnitude="3" metricUnit="m" metricValue="1.823" unit="m" value="1823.0">
<elevation id="74C9E4E932B105BFA3C62CC9F347B612" metricMagnitude="3" metricUnit="m" metricValue="1.823" unit="m" value="1823.0">1823 m</elevation>
</quantity>
; 
<collectingDate id="DEB8614B0DC175A788A7A098B7705445" value="2020-01-26">26 Jan. 2020</collectingDate>
; fr.; 
<emphasis id="98E9709D51B94093072F2F6906C08AEE" italics="true" pageId="0" pageNumber="174">
<collectorName id="06FA6B9C5C02E87D46E0427576FA20FD">Shah TS</collectorName>
98
</emphasis>
; K [K001271562]
</materialsCitation>
.
</paragraph>
<paragraph id="5987D9F0AA0B6D7A455F1FBCAFD80197" pageId="0" pageNumber="174">
<materialsCitation id="A8EE86D6E4D7C17072B2FECF17BBBD4D" ID-GBIF-Occurrence="4098977351" collectingDate="1957-12-10" collectorName="de la Tshinganda" country="DEMOCRATIC REPUBLIC OF THE CONGO" elevation="1950" location="South Kivu" specimenCount="1" stateProvince="South Kivu">
<collectingCountry id="0F4C2776DEC9122CDE34140E346C500F" name="Democratic Republic of the Congo">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
- 
<emphasis id="0ABBD7DF58EC02F75CA0D514D95BC08E" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="FD24FB0C9D9B37D98EE8656D50A4A78D" country="Democratic Republic of the Congo" name="Sud-Kivu">South Kivu</collectingRegion>
</emphasis>
• Kalehe, Mikouzi, 
<normalizedToken id="9478C97DF94F0660F6BFCF4430451533" originalValue="vallée">vallee</normalizedToken>
<collectorName id="B6D619B8D1EC21F0428B344E895A6FB0">de la Tshinganda</collectorName>
; 
<quantity id="543069FA053EDB732F982F85264F653B" metricMagnitude="3" metricUnit="m" metricValue="1.95" unit="m" value="1950.0">
<elevation id="DBBA41FDC25FFA8B87C7BEDED28518BB" metricMagnitude="3" metricUnit="m" metricValue="1.95" unit="m" value="1950.0">1950 m</elevation>
</quantity>
; 
<collectingDate id="6578AC2774B59424392F2BA93C3EDD15" value="1957-12-10">10 Dec. 1957</collectingDate>
; fr.; 
<emphasis id="E2C339ADD8B41F3B07E4A88A75515B73" italics="true" pageId="0" pageNumber="174">Pierlot 1731</emphasis>
; K [K001391607]. - 
<emphasis id="EBDCA203E884221DE3FCC94BAA0B2F92" bold="true" pageId="0" pageNumber="174">
Haut 
<materialsCitation id="366CF51A554ED1D1AD3742A89B9BB4C8" ID-GBIF-Occurrence="4098977339" collectingDate="1968-11-28" country="Democratic Republic of the Congo" elevation="1960" location="Plateau des Kundelungu" specimenCount="1" stateProvince="Katanga">
<collectingRegion id="4102562BC5BBE537C4F0E04530D3CAEF" country="Democratic Republic of the Congo" name="Katanga">Katanga</collectingRegion>
• 
<location id="F58B36DD99900F4B6CFC282EA273EDA2" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:F58B36DD99900F4B6CFC282EA273EDA2" country="Democratic Republic of the Congo" name="Plateau des Kundelungu" stateProvince="Katanga">Plateau des Kundelungu</location>
; 
<quantity id="3438156495BB7E6F1A8A5C1F0C31C7C8" metricMagnitude="3" metricUnit="m" metricValue="1.96" unit="m" value="1960.0">
<elevation id="C576ADB6720293C06D22E8E90B8467AE" metricMagnitude="3" metricUnit="m" metricValue="1.96" unit="m" value="1960.0">1960 m</elevation>
</quantity>
; 
<collectingDate id="BAA1779EF5754D27C38E0F4391B04D0C" value="1968-11-28">28 Nov. 1968</collectingDate>
; fr.; 
<emphasis id="91E3E5D04263F6193F2AF5B2A5FD9E6D" italics="true" pageId="0" pageNumber="174">Malaisse 6122</emphasis>
; K [K001391606]
</materialsCitation>
</emphasis>
</materialsCitation>
.
</paragraph>
<paragraph id="7CCF27CD0301BBB08703BE4DF0503486" pageId="0" pageNumber="174">
<materialsCitation id="A5FD91B691671E266DC0A7150BF3CA9F" ID-GBIF-Occurrence="4098977390" collectingDate="1946-09-20" collectorName="Brass" country="MALAWI" elevation="1400" location="Cholo Mt" specimenCount="1" stateProvince="Thyolo District">
<collectingCountry id="1589F893F08434CC2D5DF16D8FCD58B3" name="Malawi">MALAWI</collectingCountry>
- 
<emphasis id="4A96AB8D3B6C8BCA2F78C36B83D5EF99" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="96665127EFDFA0F91F933088F120043F" country="Malawi" name="Thyolo">Thyolo District</collectingRegion>
</emphasis>
• 
<location id="10E7C85CB7004F816840EBA1B1FD637A" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:10E7C85CB7004F816840EBA1B1FD637A" country="MALAWI" name="Cholo Mt" stateProvince="Thyolo District">Cholo Mt</location>
; 
<quantity id="235F0E30B247627376CD1261E0B8E1B5" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation id="6D495C031B1E24B3B823D9B6ACEBCB18" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
; 
<collectingDate id="9F1E95B75895BFA7F92B96CBC6745868" value="1946-09-20">20 Sep. 1946</collectingDate>
; fl.; 
<emphasis id="AD720061018B4CD1C3F51712622052BD" italics="true" pageId="0" pageNumber="174">
<collectorName id="9C5D12F5350374267A62C7AB7B7881E0">Brass</collectorName>
17668
</emphasis>
; BM, K [K001387307]
</materialsCitation>
. - 
<materialsCitation id="3AFE8D81CFCC1A9ED515B8768236F630" ID-GBIF-Occurrence="4098977319" collectingDate="1952-05-08" collectingDateMax="1957-10-08" collectingDateMin="1952-05-08" collectorName="White, Chapman, Nyika Plateau, Salubeni" country="Malawi" county="Lundazi" elevation="2134" location="Nyika Plateau" municipality="Kangampande Mt" specimenCount="1" stateProvince="Chitipa District">
<emphasis id="6D92317F783D858E31E88E6874DC99FE" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="5EB68D81F9342580D9D2D90284524AD4" country="Malawi" name="Chitipa">Chitipa District</collectingRegion>
</emphasis>
• 
<collectingCounty id="D2CB97E21098530DC94F0E5EB17D5C4D">Lundazi</collectingCounty>
, upper slopes of 
<collectingMunicipality id="02D82F2473821475D05C66A2022DDE5D">Kangampande Mt</collectingMunicipality>
, 
<location id="7E573F3BB0ED6429EE4D33F7CA470EB5" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:7E573F3BB0ED6429EE4D33F7CA470EB5" country="Malawi" county="Lundazi" municipality="Kangampande Mt" name="Nyika Plateau" stateProvince="Chitipa District">Nyika Plateau</location>
; 
<quantity id="BB82BCA43626A65DD206C780B3FD51FD" metricMagnitude="3" metricUnit="m" metricValue="2.1336" unit="ft" value="7000.0">
<elevation id="A60E32CA33F75FC920656467A2AACE73" metricMagnitude="3" metricUnit="m" metricValue="2.1336" unit="ft" value="7000.0">7000 ft</elevation>
</quantity>
; 
<collectingDate id="CD73D727DD44764999A374393A7D3CD0" value="1952-05-08">8 May 1952</collectingDate>
; 
<emphasis id="CFAE1E31B459B0274223570083A72A6D" italics="true" pageId="0" pageNumber="174">
<collectorName id="DE29F4C182FA9D5ACFC02BEEC9FAAA90">White</collectorName>
2793
</emphasis>
; K [K001271529] • 
<collectingRegion id="7961DC4627EBDB9B4B89171EBE5FB469" country="Malawi" name="Mulanje">Mulanje</collectingRegion>
<location id="D40B6B90F9DD10DBA9835611255D820C" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:D40B6B90F9DD10DBA9835611255D820C" country="Malawi" county="Lundazi" municipality="Kangampande Mt" name="Mt forest" stateProvince="Chitipa District">Mt forest</location>
; 
<collectingDate id="93005BD92245315E2DCB8362FAE2FBE4" value="1957-10-08">8 Oct. 1957</collectingDate>
; fl.; 
<emphasis id="EC13F9F817F3CEABA4E99592BD1B4ECD" italics="true" pageId="0" pageNumber="174">
<collectorName id="4F291A5D53C1B7DD064184B7AD84A4B5">Chapman</collectorName>
454
</emphasis>
; BM, K [K001387304] • 
<collectorName id="EE8A702A1BF19C47BBDBFF2DF47BCD76">Nyika Plateau</collectorName>
, 
<location id="77447B98DE59B6933214902951B91E28" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:77447B98DE59B6933214902951B91E28" country="Malawi" county="Lundazi" municipality="Kangampande Mt" name="Luselo Evergreen Forest" stateProvince="Chitipa District">Luselo Evergreen Forest</location>
patch; 
<emphasis id="940A39F4B9AAEA11EA9F2CFAD1B96C3A" italics="true" pageId="0" pageNumber="174">
<collectorName id="4606A043BB99C720D38E53D1DE6A33CA">Salubeni</collectorName>
356
</emphasis>
; K [K001387298]
</materialsCitation>
. - 
<materialsCitation id="E6627DDB18326E02066E085A299ED1DD" ID-GBIF-Occurrence="4098977311" collectingDate="1986-11-03" collectorName="Chapman" country="Malawi" elevation="1840" location="Boma" municipality="Mt" specimenCount="1" stateProvince="Mulanje District">
<emphasis id="7DC3E1EAB1E5B3A241650DA5E7DF1D8A" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="342FAAE1AC285805ECC134B60C1D64FB" country="Malawi" name="Mulanje">Mulanje District</collectingRegion>
</emphasis>
• 
<collectingMunicipality id="5916E815F4E777944106C33DEA2CD727">Mt</collectingMunicipality>
<collectingRegion id="1108EFBD6D0139BE3FD2FC768A8BAAD9" country="Malawi" name="Mulanje">Mulanje</collectingRegion>
, close to the site of the old 
<location id="C0D61B9CAC0D40962EEA21EA86DECD00" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:C0D61B9CAC0D40962EEA21EA86DECD00" country="Malawi" municipality="Mt" name="Boma" stateProvince="Mulanje District">Boma</location>
hut; 
<quantity id="A83EABECCE798825F0D5486059E893AA" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" unit="m" value="1840.0">
<elevation id="B6B9F763327B6DAB9E9AE2E413D9451A" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" unit="m" value="1840.0">1840 m</elevation>
</quantity>
; 
<collectingDate id="7C40EED62EC01E82CB1CCC4CDFA04AA3" value="1986-11-03">3 Nov. 1986</collectingDate>
; 
<emphasis id="912038C1B433B81CD61A96B7C0140F3C" italics="true" pageId="0" pageNumber="174">
<collectorName id="7C44F90112A2EAF94F1582A302849171">Chapman</collectorName>
&amp; 
<collectorName id="825D3AE12BE2445E4065F892002FD460">Chapman</collectorName>
8186
</emphasis>
; E [E01123512], K [K001387295]
</materialsCitation>
.
</paragraph>
<paragraph id="5A0169612F43731C458CA3913936092C" pageId="0" pageNumber="174">
<materialsCitation id="E57AD24CDBD39DECFE92560EEAC393C6" ID-GBIF-Occurrence="4098977387" collectingDate="2006-11-29" collectingDateMax="2007-11-21" collectingDateMin="2006-11-29" collectorName="Patel, Bayliss HP, Naconha Plateau" country="MOZAMBIQUE" county="Chiperone Mountain" elevation="1977" latitude="-15.399278" location="Namuli Mountain" longLatPrecision="1" longitude="37.041637" municipality="Namuli Mountain" specimenCount="1" stateProvince="Zambezia Province">
<collectingCountry id="27B86B325C82E45BBE67F104C185054A" name="Mozambique">MOZAMBIQUE</collectingCountry>
- 
<emphasis id="D6FA7A2C4A20189DA0DBD97E04644E77" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="1B847A42FC53C20D72A4EE50A9B4991B" country="Mozambique" name="Zambezia">Zambezia Province</collectingRegion>
</emphasis>
• 
<collectingCounty id="87E8D17A6E30350F97E1038C9958D691">Chiperone Mountain</collectingCounty>
; 
<collectingDate id="0C4B3B464F38232B9BE373FDF0A7A59B" value="2006-11-29">29 Nov. 2006</collectingDate>
; 
<emphasis id="158E529679AE19812642AABDC0B45F2B" italics="true" pageId="0" pageNumber="174">
<collectorName id="25FE785131495CC0B45E03ED3404F07C">Patel</collectorName>
&amp; 
<collectorName id="F5008D52B818F71669A764FCE0EB8579">Bayliss HP</collectorName>
7183
</emphasis>
; K [K000545146] • 
<collectingMunicipality id="7EB856FDEF1003957A9443515EB06BA3">Namuli Mountain</collectingMunicipality>
, 
<collectorName id="85D773C79AA25D66EEE1BE9040649FA6">Naconha Plateau</collectorName>
; 
<geoCoordinate id="50A424735A6B89D0281A3B8B5841A847" degrees="15" direction="south" minutes="22" orientation="latitude" precision="15" seconds="40" value="-15.377778">
15°22 
<normalizedToken id="6304335013E70554EFD7AF0D0C2C653B" originalValue="’40”">'40&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate id="8703E1350E3473DDB935D5929A63B895" degrees="37" direction="east" minutes="01" orientation="longitude" precision="1" seconds="09.2" value="37.019222">
37°01 
<normalizedToken id="FFD3B09B39361F60A65E4D1E15A141EF" originalValue="’09.2”">'09.2&quot;</normalizedToken>
E
</geoCoordinate>
; 
<quantity id="BAD30F0434EA7BD05DB2779FB5511537" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">
<elevation id="0CBE38F2754FD9997D9A4C90ADD9C3ED" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">1977 m</elevation>
</quantity>
; 
<collectingDate id="625DE58D0EF47EA357211C4B2FC44D0D" value="2007-11-21">21 Nov. 2007</collectingDate>
; fr.; 
<emphasis id="85A1649147D6BC4BB8319415E67E3587" italics="true" pageId="0" pageNumber="174">Timberlake et al. 5267</emphasis>
; K [K000613996] • 
<location id="954978C6EAC9EFA24A98A01FE3990B84" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:954978C6EAC9EFA24A98A01FE3990B84" country="MOZAMBIQUE" county="Chiperone Mountain" latitude="-15.399278" longLatPrecision="1" longitude="37.041637" municipality="Namuli Mountain" name="Namuli Mountain" stateProvince="Zambezia Province">Namuli Mountain</location>
, 
<location id="BE98CD5803F0FD364D180FE5772C3F08" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:BE98CD5803F0FD364D180FE5772C3F08" country="MOZAMBIQUE" county="Chiperone Mountain" latitude="-15.399278" longLatPrecision="1" longitude="37.041637" municipality="Namuli Mountain" name="Muretha Plateau-Mukocha forest" stateProvince="Zambezia Province">Muretha Plateau-Mukocha forest</location>
; 
<geoCoordinate id="949963B9F4D705ACC2F303D9C9433DB8" degrees="15" direction="south" minutes="23" orientation="latitude" precision="1" seconds="57.4" value="-15.399278">
15°23 
<normalizedToken id="BB22F573252D69D5D07082BB32954AFB" originalValue="’57.4”">'57.4&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate id="D1A62B20FA07012AF47555250EBF0F9A" degrees="37" direction="east" minutes="02" orientation="longitude" precision="1" seconds="29.9" value="37.041637">
37°02 
<normalizedToken id="0F7157A65D1EEC92DC8483CDC97BCE81" originalValue="’29.9”">'29.9&quot;</normalizedToken>
E
</geoCoordinate>
; 
<quantity id="7FD6F98EAF4C0AF29EDA35A86CFB591F" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">
<elevation id="4F9011A7AB739D02132BD44B2D386236" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">1730 m</elevation>
</quantity>
; 
<collectingDate id="A22038AB72B04F4CA8FFEC9345266768" value="2007-05-28">28 May 2007</collectingDate>
; 
<emphasis id="C9575E6191246A0A017C8AED8B96B4C0" italics="true" pageId="0" pageNumber="174">Timberlake et al. 5054</emphasis>
; K [K000613434]
</materialsCitation>
.
</paragraph>
<paragraph id="E28316D83E895E4BC1182DC59B368DE0" pageId="0" pageNumber="174">
<collectingCountry id="F7C565245F1B0D2A0906527D57D517ED" name="Zimbabwe">ZIMBABWE</collectingCountry>
- 
<emphasis id="DA9167E77159641A251F54BF166A12C4" bold="true" pageId="0" pageNumber="174">Mutare District</emphasis>
• Umtali; 
<date id="EDFF256ABD6E7625EB251E70B4AD2BCF" value="1948-11-08">8 Nov. 1948</date>
; fl.; 
<emphasis id="7BE6E83631A84298D2F7CF1D36B2C730" italics="true" pageId="0" pageNumber="174">Chase 1256</emphasis>
; BM, K [K001387323]. - 
<emphasis id="472E8D398541771318E8D0B304A963D2" bold="true" pageId="0" pageNumber="174">Chipinge District</emphasis>
• Chirinda Forest; 
<date id="C4C5C61174ABEAE4727772BF323CC18A" value="1948-01-04">4 Jan. 1948</date>
; fr.; 
<emphasis id="2F99525210F77C72CEBBB070DE44E599" italics="true" pageId="0" pageNumber="174">Chase 433</emphasis>
; BM, K [K001387316] • Old Bikita, on edge of forest plateau; 
<quantity id="0CCF64FBDC1B4ADCC99758CF5F83844F" metricMagnitude="3" metricUnit="m" metricValue="1.3106399999999998" unit="ft" value="4300.0">4300 ft</quantity>
; 
<date id="9E3F768F0D55D22E25EE64A363AE7495" value="1953-12-16">16 Dec. 1953</date>
; fr.; 
<emphasis id="F3AE1190095E86E8171CB401DB5CA64D" italics="true" pageId="0" pageNumber="174">Wild 4407</emphasis>
; K [K001387315] • Gwasha, on the first range; 
<quantity id="9E2EEFDA2011AEAEEF6784EAFB95EA5D" metricMagnitude="3" metricUnit="m" metricValue="1.3716" unit="ft" value="4500.0">4500 ft</quantity>
; 
<date id="B4B7D65EA7558EB308A14945DF3CFAB1" value="1960-09-22">22 Sep. 1960</date>
; fr.; 
<emphasis id="149D67410CAA10ABFFC8DB5353369DCB" italics="true" pageId="0" pageNumber="174">Rutherford-Smith 168</emphasis>
; K [K001387335]. - 
<emphasis id="0834A6D6439CA3BAFCBB60278E43AE87" bold="true" pageId="0" pageNumber="174">Chimanimani District</emphasis>
• Chimanimani Mountain, Martin Forest Reserve; 
<date id="0378B69581769540DF33CC8A7D65C7A0" value="1967-11-17">17 Nov. 1967</date>
; fl.; 
<emphasis id="E3CF7CD333B00402C0F146FB3F41CB52" italics="true" pageId="0" pageNumber="174">Mavi 678</emphasis>
; K [K001387313]. - 
<emphasis id="68F2D6BE010C3C62056CEFF2B774D77A" bold="true" pageId="0" pageNumber="174">Mberengwa District</emphasis>
• Bukwa Mt, main ridge; 
<quantity id="882F82399D3F99DB2FBB82E539FC7AB5" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
; 
<date id="A4A30BE65BFF03051F2863F625ABDA82" value="1973-10-30">30 Oct. 1973</date>
; fl.; 
<emphasis id="0E650F567AC6186465900F8EE9AE5DB4" italics="true" pageId="0" pageNumber="174">Pope et al. 1126</emphasis>
; K [K001387336].
</paragraph>
<paragraph id="5AF72A75309A43AB9609D85938449D81" pageId="0" pageNumber="174">
<materialsCitation id="CDC8F9E6412915583749442BD94D3597" ID-GBIF-Occurrence="4098977365" collectingDate="1915-12-18" collectorName="Chilvers" country="SOUTH AFRICA" location="Ingeli Forest Reserve" specimenCount="1" stateProvince="KwaZulu-Natal">
<collectingCountry id="604D552CDD84A6AFF68B9D3715C3A48B" name="South Africa">SOUTH AFRICA</collectingCountry>
- 
<emphasis id="A94E2173FD179F63E1C867CEBA40F9F4" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="BDCA47BD5B16C2B843AF436A1DFD1A71" country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
</emphasis>
• 
<location id="892FBBA0204B5363E319F6845043C56A" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:892FBBA0204B5363E319F6845043C56A" country="SOUTH AFRICA" name="Ingeli Forest Reserve" stateProvince="KwaZulu-Natal">Ingeli Forest Reserve</location>
; 
<collectingDate id="E319797FD45DD4A5E4ED627815BC0166" value="1915-12-18">18 Dec. 1915</collectingDate>
; fr.; 
<emphasis id="365DDDE6E3D45D4506D2BA2E1C6787F7" italics="true" pageId="0" pageNumber="174">
<collectorName id="ACA0753E8E3A615AC8F8EF880541CF52">Chilvers</collectorName>
1946
</emphasis>
; K [K001271523]
</materialsCitation>
. - 
<materialsCitation id="154E6EBC727DBD5E18C453CAF80E5B65" ID-GBIF-Occurrence="4098977380" collectingDate="1957-01-22" collectorName="Story" country="South Africa" location="Barberton" specimenCount="1" stateProvince="Mpumalanga">
<emphasis id="2E6F8618DF5A16E342E9820C82D85596" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="A1A31BFC75AFA812894C3A55BC996481" country="South Africa" name="Mpumalanga">Mpumalanga</collectingRegion>
</emphasis>
• 
<location id="BBC684867140DAAD25EDAA788A59632D" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:BBC684867140DAAD25EDAA788A59632D" country="South Africa" name="Barberton" stateProvince="Mpumalanga">Barberton</location>
; 
<collectingDate id="873EEF0DE09623D4C38A50DF42C84B92" value="1957-01-22">22 Jan. 1957</collectingDate>
; 
<emphasis id="46DE2B913EA567679E4641A6D86506D3" italics="true" pageId="0" pageNumber="174">
<collectorName id="37581D0C2C663AEBCD171011542C2088">Story</collectorName>
5993
</emphasis>
; K [K001271517]
</materialsCitation>
. - 
<materialsCitation id="85EC6AB497B9C1837457CE757A19DC52" ID-GBIF-Occurrence="4098977373" collectingDate="1959-12-03" collectorName="Schiff" country="South Africa" location="Mariepskop" specimenCount="1" stateProvince="Limpopo">
<emphasis id="54D4C17F7E7F84F248F7ED7FC8EC9013" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="1650E3D8B2D4758AED8FEA3629529BF6" country="South Africa" name="Limpopo">Limpopo</collectingRegion>
</emphasis>
• 
<location id="956B67F1A8C3D702F0414826FBEA593A" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:956B67F1A8C3D702F0414826FBEA593A" country="South Africa" name="Mariepskop" stateProvince="Limpopo">Mariepskop</location>
; 
<collectingDate id="1FCB214FF575008ADF431023DD30461F" value="1959-12-03">3 Dec. 1959</collectingDate>
; fr.; 
<emphasis id="3BD07DF71F23F62C2D9630AE3C482933" italics="true" pageId="0" pageNumber="174">
<collectorName id="601D34606725AC29ACFBB0C3777E2D55">Schiff</collectorName>
4754
</emphasis>
; K [K001271518]
</materialsCitation>
. - 
<materialsCitation id="F77E6BFF5407B5E097EB2AD265036F44" ID-GBIF-Occurrence="4098977347" collectingDate="1996-01-18" collectorName="Flanagan" country="South Africa" location="Engcobo Forest" specimenCount="1" stateProvince="Eastern Cape">
<emphasis id="80B06E3ED2D2C7E0B25924E287B10FBE" bold="true" pageId="0" pageNumber="174">
<collectingRegion id="A1B6675D1B0BA519303B4D9251F73E0C" country="South Africa" name="Eastern Cape">Eastern Cape</collectingRegion>
</emphasis>
• 
<location id="6051883D5F8728A6C52956D9CE6EBD7B" LSID="urn:lsid:plazi:treatment:AEB6397B6DEC577A9E583B26222BB40A:6051883D5F8728A6C52956D9CE6EBD7B" country="South Africa" name="Engcobo Forest" stateProvince="Eastern Cape">Engcobo Forest</location>
; 
<collectingDate id="4A065A060EA3521FEAB513D22ECBA9CD" value="1996-01-18">18 Jan. 1996</collectingDate>
; fr.; 
<emphasis id="C880139C86589597FFCBF118B38D51E2" italics="true" pageId="0" pageNumber="174">
<collectorName id="69FF50A0F1E5D42FB11BE988F26D6D46">Flanagan</collectorName>
2696
</emphasis>
; K [K001271522]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8C55537530A94F850AA34B6C2418A71F" pageId="0" pageNumber="174" type="bibliography">
<paragraph id="55F3E2970B11E75B0C814B9FAB634EC7" pageId="0" pageNumber="174">Bibliography.</paragraph>
<paragraph id="EB039909CF9BF4B70D91551D8B8707C5" pageId="0" pageNumber="174">
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: 2).
</paragraph>
</subSubSection>
<subSubSection id="54073BC7A9E5AEAB911BABC4A1552167" pageId="0" pageNumber="174" type="notes">
<paragraph id="9AFAC030FFF6E42CDE2181F10CD2E9E8" pageId="0" pageNumber="174">Notes.</paragraph>
<paragraph id="FF19B106D3794C325021C89D7D0C4F7A" pageId="0" pageNumber="174">
Morphological work on this species has proved challenging. Like taxonomists in the past (
<bibRefCitation id="28170B8C3F98A4EFFFCEB22623FB396A" author="Robson, NK" editor="Exell, AW" journalOrPublisher="2 (1). Flora Zambesiaca Managing Committee, London" pageId="0" pageNumber="174" pagination="225 - 251" refId="B64" refString="Robson, NK, 1963. Ochnaceae. In: Exell, AW, Fernandes, A, Wild, H, Eds., Flora Zambesiaca, vol. 2(1). Flora Zambesiaca Managing Committee, London: 225 - 251" title="Ochnaceae." volumeTitle="Flora Zambesiaca, vol." year="1963">Robson 1963</bibRefCitation>
; 
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), we note significant variation in habit, leaf size and arrangement of leaves along the branches, and inflorescence structure. This widespread species is generally recognizable by being a medium- to large-sized tree, occurring in upland moist forest with leaves variable in size (4-12 cm long and 1-5 cm wide) with a distinct attenuate leaf apex, and 6-16 flowers with long pedicles arranged in a distinct raceme. Difficulty arises with specimens that are sterile, shrubbier in habit, and/or with smaller leaves occurring in more open and drier habitats. These individuals may be confused with 
<taxonomicName id="1972ADC0D03DE1D4805C1992B34EEB8B" class="Magnoliopsida" family="Ochnaceae" kingdom="Plantae" lsidName="O. stolzii" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="stolzii">
<emphasis id="004BB0BD418093544C1AF70E695D6A13" italics="true" pageId="0" pageNumber="174">O. stolzii</emphasis>
</taxonomicName>
and 
<taxonomicName id="F01F0A77B350883CAACC643B04D1DC3A" class="Magnoliopsida" family="Ochnaceae" kingdom="Plantae" lsidName="O. oxyphylla" order="Malpighiales" pageId="0" pageNumber="174" phylum="Tracheophyta" rank="species" species="oxyphylla">
<emphasis id="002533A4FAC1CB02EAAFD8BA0BA16EAD" italics="true" pageId="0" pageNumber="174">O. oxyphylla</emphasis>
</taxonomicName>
; however, they differ by the latter two species generally having smaller leaves (1-4.5 cm long, 0.5-2 cm wide and 2-8 cm long, 1-4 cm wide respectively) without prominent attenuate leaf apices, and fewer flowers per inflorescence, often solitary but sometimes up to six flowers.
</paragraph>
</subSubSection>
</treatment>
</document>