316 lines
50 KiB
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316 lines
50 KiB
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<document id="4A2384CB2EF57673D0FD41990DE67886" ID-CLB-Dataset="27502" ID-DOI="10.11646/zootaxa.4550.1.1" ID-GBIF-Dataset="291701d4-652e-440e-adab-f6b0201c0972" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625030" ID-ZooBank="2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554274879827" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2019" docId="CA3B104CFFB4FFB9FF3DFDE9FC46E61F" docLanguage="en" docName="zootaxa.4550.1.1.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wanniyala mudita Huber 2019, sp. n." docType="treatment" docVersion="14" lastPageNumber="35" masterDocId="36026834FF95FF9BFFAAFFA1FFB6E16A" masterDocTitle="The pholcid spiders of Sri Lanka (Araneae: Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="34" updateTime="1698716853299" updateUser="ExternalLinkService">
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<mods:titleInfo id="EA5724A4EBD9AD3D2F46F7EFE6B58060">
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<mods:title id="C63506205D4C95BE3D10E0371421E525">The pholcid spiders of Sri Lanka (Araneae: Pholcidae)</mods:title>
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<mods:name id="81E277B1FB8F6D755917323B9A269299" type="personal">
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<mods:namePart id="FADA60B9383561915D330F7EC8FE09EE">Huber, Bernhard A.</mods:namePart>
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<mods:title id="0962D884B922C93B4CA05645425F6C38">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D66F3ECA7F279CFF3C6D08EA21255F40">
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<mods:date id="1BFBF4221BF9A281F0F58407B6FD0C5C">2019</mods:date>
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<mods:number id="82BA40ECE0765D69704E9D3FC2C65C1B">2019-01-23</mods:number>
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<mods:number id="502E4B6CA52272A25DDFBD06ADAA8BC3">4550</mods:number>
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<mods:identifier id="57C116806492AECA76AFA24A51204DC1" type="CLB-Dataset">27502</mods:identifier>
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<mods:identifier id="25316C873BEC8134B6EB4B8D8372AADA" type="DOI">10.11646/zootaxa.4550.1.1</mods:identifier>
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<treatment id="CA3B104CFFB4FFB9FF3DFDE9FC46E61F" ID-DOI="http://doi.org/10.5281/zenodo.4581683" ID-GBIF-Taxon="154534093" ID-Zenodo-Dep="4581683" LSID="urn:lsid:plazi:treatment:CA3B104CFFB4FFB9FF3DFDE9FC46E61F" httpUri="http://treatment.plazi.org/id/CA3B104CFFB4FFB9FF3DFDE9FC46E61F" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
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<subSubSection id="0A88F2D1FFB4FFBAFF3DFDE9FE0BE308" box="[151,445,584,610]" pageId="33" pageNumber="34" type="nomenclature">
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<paragraph id="422DA15AFFB4FFBAFF3DFDE9FE0BE308" blockId="33.[151,508,584,643]" box="[151,445,584,610]" pageId="33" pageNumber="34">
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<heading id="19651636FFB4FFBAFF3DFDE9FE0BE308" bold="true" box="[151,445,584,610]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
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<emphasis id="70E67D48FFB4FFBAFF3DFDE9FE0BE308" bold="true" box="[151,445,584,610]" pageId="33" pageNumber="34">
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<taxonomicName id="8592DAD9FFB4FFBAFF3DFDE9FEC3E308" ID-CoL="7G64W" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[151,373,584,610]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="mudita" status="sp. nov.">
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<emphasis id="70E67D48FFB4FFBAFF3DFDE9FEC3E308" bold="true" box="[151,373,584,610]" italics="true" pageId="33" pageNumber="34">Wanniyala mudita</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6BD5C033FFB4FFBAFED1FDE9FE0BE308" box="[379,445,584,610]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A88F2D1FFB4FFBAFF3DFDCBFCFBE3A2" pageId="33" pageNumber="34" type="description">
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<paragraph id="422DA15AFFB4FFBAFF3DFDCBFE4AE3E9" blockId="33.[151,508,584,643]" box="[151,508,618,643]" pageId="33" pageNumber="34">
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<figureCitation id="DAA9BDDFFFB4FFBAFF3DFDCBFEE0E3E9" box="[151,342,618,643]" captionStart="FIGURES 126–137" captionStartId="32.[151,264,1429,1452]" captionTargetBox="[151,1436,193,1409]" captionTargetId="figure@32.[151,1436,193,1409]" captionTargetPageId="32" captionText="FIGURES 126–137. Wanniyala, live specimens, part 1. 126–128. W. agrabopath Huber & Benjamin, 2005, male and females with egg-sacs from Hakgala. 129–131. W. mudita sp. n., male and females with egg-sacs from Kandy. 132–134. W. orientalis sp. n., males and female with egg-sac from Inginiyagala. 135–137. W. upekkha sp. n., male from Dematagala and females with and without egg-sac from Ella." httpUri="https://zenodo.org/record/2625074/files/figure.png" pageId="33" pageNumber="34">Figures 129–131</figureCitation>
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,
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<figureCitation id="DAA9BDDFFFB4FFBAFEC8FDCBFE71E3E9" box="[354,455,618,643]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">156–164</figureCitation>
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,
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<figureCitation id="DAA9BDDFFFB4FFBAFE78FDCAFE4AE3E9" box="[466,508,619,643]" captionStart="FIGURES 213–219" captionStartId="49.[151,264,1720,1743]" captionTargetBox="[151,1436,412,1657]" captionTargetId="figure@49.[151,1436,412,1657]" captionTargetPageId="49" captionText="FIGURES 213–219. Wanniyala spp., cleared female genitalia in dorsal views. 213. W. mudita sp. n., from Kandy (ZFMK Ar 20077). 214. W. orientalis sp. n., from Dimbulagala (ZFMK Ar 20079). 215. W. upekkha sp. n., from Dematagala (ZFMK Ar 20086). 216. W. ohiya sp. n., from Ohiya (ZFMK Ar 20089). 217. W. viharekele sp. n., from Viharekele (ZFMK Ar 20092). 218. W. mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 219. W. labugama sp. n., from Labugama (ZFMK Ar 20097). At various scales." httpUri="https://zenodo.org/record/2625096/files/figure.png" pageId="33" pageNumber="34">213</figureCitation>
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</paragraph>
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<paragraph id="422DA15AFFB4FFBAFF3DFD13FCFBE3A2" blockId="33.[151,845,690,712]" box="[151,845,690,712]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="62828674FFB4FFBAFF3DFD13FCFBE3A2" box="[151,845,690,712]" pageId="33" pageNumber="34">
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<taxonomicName id="8592DAD9FFB4FFBAFF3DFD13FEB5E3A2" ID-CoL="87JX" authorityName="Huber & Benjamin" authorityYear="2005" box="[151,259,690,712]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="70E67D48FFB4FFBAFF3DFD13FEB5E3A2" box="[151,259,690,712]" italics="true" pageId="33" pageNumber="34">Wanniyala</emphasis>
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</taxonomicName>
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sp. (from Rattota):
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<treatmentCitation id="C333874BFFB4FFBAFE60FD13FCB0E3A2" author="Huber, B. A. & Benjamin, S." box="[458,774,690,712]" httpUri="http://treatment.plazi.org/id/039387D9FFCAFFF2C17D1F7AFDBB6D14" page="3311" pageId="33" pageNumber="34" year="2005">
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<bibRefCitation id="2603DCABFFB4FFBAFE60FD13FD7EE3A2" author="Huber, B. A. & Benjamin, S." box="[458,712,690,712]" pageId="33" pageNumber="34" pagination="3305 - 3319" refId="ref27050" refString="Huber, B. A. & Benjamin, S. (2005) The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of Wanniyala new genus (Araneae: Pholcidae). Journal of Natural History, 39 (37), 3305 - 3319." type="journal article" year="2005">Huber & Benjamin 2005</bibRefCitation>
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: 3311
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</treatmentCitation>
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, fig. 9.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A88F2D1FFB4FFBAFF3DFD54FAC9E2A8" pageId="33" pageNumber="34" type="diagnosis">
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<paragraph id="422DA15AFFB4FFBAFF3DFD54FAC9E2A8" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
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<emphasis id="70E67D48FFB4FFBAFF3DFD54FEBCE264" bold="true" box="[151,266,757,782]" pageId="33" pageNumber="34">Diagnosis</emphasis>
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. Males are distinguished from congeners by details of palp (retrolateral sclerite of procursus with large rounded dorsal process; distinctive processes of palpal trochanter, and short and weakly curved bulbal apophysis;
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<figureCitation id="DAA9BDDFFFB4FFBAFF3DFC9CFE80E23C" box="[151,310,829,854]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Figs 158–161</figureCitation>
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) and by modification of clypeus (pair of processes connected by protruding arc;
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<figureCitation id="DAA9BDDFFFB4FFBAFB5DFC9CFA21E23C" box="[1271,1431,829,854]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Figs 156–157</figureCitation>
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; similar only in
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<taxonomicName id="8592DAD9FFB4FFBAFEEAFCC3FE79E210" authorityName="Huber" authorityYear="2019" box="[320,463,865,890]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="70E67D48FFB4FFBAFEEAFCC3FE79E210" box="[320,463,865,890]" italics="true" pageId="33" pageNumber="34">W. orientalis</emphasis>
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</taxonomicName>
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). Females are easily distinguished from most known congeners (except
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<taxonomicName id="8592DAD9FFB4FFBAFB5FFCC3FAC5E210" authorityName="in Huber & Benjamin" authorityYear="2005" box="[1269,1395,865,890]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="hakgala">
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<emphasis id="70E67D48FFB4FFBAFB5FFCC3FAC5E210" box="[1269,1395,865,890]" italics="true" pageId="33" pageNumber="34">W. hakgala</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8592DAD9FFB4FFBAFA2AFCC3FEB4E2F4" authorityName="Huber" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="70E67D48FFB4FFBAFA2AFCC3FEB4E2F4" italics="true" pageId="33" pageNumber="34">W. orientalis</emphasis>
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</taxonomicName>
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) by shape of epigynum (strongly projecting and pointed in lateral view;
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<figureCitation id="DAA9BDDFFFB4FFBAFBEDFC24FB1DE2F4" box="[1095,1195,901,926]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Fig. 163</figureCitation>
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); they differ from
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<taxonomicName id="8592DAD9FFB4FFBAFA2AFC27FF47E2A8" authorityName="in Huber & Benjamin" authorityYear="2005" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="hakgala">
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<emphasis id="70E67D48FFB4FFBAFA2AFC27FF47E2A8" italics="true" pageId="33" pageNumber="34">W. hakgala</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8592DAD9FFB4FFBAFE8DFC0BFE03E2A8" authorityName="Huber" authorityYear="2019" box="[295,437,937,962]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="70E67D48FFB4FFBAFE8DFC0BFE03E2A8" box="[295,437,937,962]" italics="true" pageId="33" pageNumber="34">W. orientalis</emphasis>
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</taxonomicName>
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by details of internal genitalia (wide anterior ‘valve’; oval pore plates;
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<figureCitation id="DAA9BDDFFFB4FFBAFB7FFC08FA8EE2A8" box="[1237,1336,937,962]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Figs 164</figureCitation>
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,
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|
<figureCitation id="DAA9BDDFFFB4FFBAFAEEFC0BFAC4E2A8" box="[1348,1394,938,962]" captionStart="FIGURES 213–219" captionStartId="49.[151,264,1720,1743]" captionTargetBox="[151,1436,412,1657]" captionTargetId="figure@49.[151,1436,412,1657]" captionTargetPageId="49" captionText="FIGURES 213–219. Wanniyala spp., cleared female genitalia in dorsal views. 213. W. mudita sp. n., from Kandy (ZFMK Ar 20077). 214. W. orientalis sp. n., from Dimbulagala (ZFMK Ar 20079). 215. W. upekkha sp. n., from Dematagala (ZFMK Ar 20086). 216. W. ohiya sp. n., from Ohiya (ZFMK Ar 20089). 217. W. viharekele sp. n., from Viharekele (ZFMK Ar 20092). 218. W. mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 219. W. labugama sp. n., from Labugama (ZFMK Ar 20097). At various scales." httpUri="https://zenodo.org/record/2625096/files/figure.png" pageId="33" pageNumber="34">213</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="0A88F2D1FFB4FFBAFF6DFC6CFBAAE560" pageId="33" pageNumber="34" type="etymology">
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DFC6CFBAAE560" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
<emphasis id="70E67D48FFB4FFBAFF6DFC6CFEF0E28C" bold="true" box="[199,326,973,998]" pageId="33" pageNumber="34">Etymology</emphasis>
|
|||
|
. The species name is derived from the Pāli word
|
|||
|
<taxonomicName id="8592DAD9FFB4FFBAFC37FC6CFC5DE28C" authorityName="Huber" authorityYear="2019" box="[925,1003,973,998]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="mudita">
|
|||
|
<emphasis id="70E67D48FFB4FFBAFC37FC6CFC5DE28C" box="[925,1003,973,998]" italics="true" pageId="33" pageNumber="34">muditā</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, one of the four Buddhist virtues or brahmavihāras (the feeling of joy because others are happy); noun in apposition.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="0A88F2D1FFB4FFBAFF6DFBB4FDBEE440" pageId="33" pageNumber="34" type="materials_examined">
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DFBB4FC07E538" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
<emphasis id="70E67D48FFB4FFBAFF6DFBB4FEC4E544" bold="true" box="[199,370,1045,1070]" pageId="33" pageNumber="34">Type material</emphasis>
|
|||
|
.
|
|||
|
<materialsCitation id="F2FAAB07FFB4FFBAFE2EFBB7FC1BE538" ID-GBIF-Occurrence="2234226244" collectingDate="2017-03-08" collectionCode="ZFMK" collectorName="B. A. Huber" country="Sri Lanka" county="Ar" elevation="640" latitude="7.282" location="Dunumadallawa Forest" longLatPrecision="78" longitude="80.643" municipality="Kandy" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="holotype">
|
|||
|
<collectingCountry id="3A85E1CAFFB4FFBAFE2EFBB7FD95E544" box="[388,547,1046,1070]" name="Sri Lanka" pageId="33" pageNumber="34">SRI LANKA</collectingCountry>
|
|||
|
:
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFD9BFBB7FDF0E547" box="[561,582,1046,1069]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
|
|||
|
<typeStatus id="9D291FF8FFB4FFBAFDF8FBB4FD01E544" box="[594,695,1045,1070]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
|||
|
,
|
|||
|
<collectionCode id="2483399FFFB4FFBAFD6CFBB7FCA2E547" box="[710,788,1046,1069]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="33" pageNumber="34" type="Museum">ZFMK</collectionCode>
|
|||
|
(
|
|||
|
<collectingCounty id="AB4CD9D6FFB4FFBAFC8DFBB7FCF0E544" box="[807,838,1046,1070]" pageId="33" pageNumber="34">Ar</collectingCounty>
|
|||
|
20076),
|
|||
|
<collectingRegion id="80566FB8FFB4FFBAFC1FFBB4FBCFE544" box="[949,1145,1045,1070]" country="Sri Lanka" name="Central" pageId="33" pageNumber="34">Central Province</collectingRegion>
|
|||
|
,
|
|||
|
<collectingMunicipality id="A2493B20FFB4FFBAFB23FBB4FB63E544" box="[1161,1237,1045,1070]" pageId="33" pageNumber="34">Kandy</collectingMunicipality>
|
|||
|
,
|
|||
|
<location id="474DF781FFB4FFBAFB4EFBB4FF6BE53B" LSID="urn:lsid:plazi:treatment:CA3B104CFFB4FFB9FF3DFDE9FC46E61F:474DF781FFB4FFBAFB4EFBB4FF6BE53B" country="Sri Lanka" county="Ar" latitude="7.282" longLatPrecision="78" longitude="80.643" municipality="Kandy" name="Dunumadallawa Forest" pageId="33" pageNumber="34" stateProvince="Central">Dunumadallawa Forest</location>
|
|||
|
(
|
|||
|
<geoCoordinate id="27A6C79DFFB4FFBAFF46FB9BFEF8E538" box="[236,334,1082,1106]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="55" value="7.282">7.282°N</geoCoordinate>
|
|||
|
,
|
|||
|
<geoCoordinate id="27A6C79DFFB4FFBAFEF0FB98FE71E538" box="[346,455,1081,1106]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="55" value="80.643">80.643°E</geoCoordinate>
|
|||
|
),
|
|||
|
<elevation id="C9BF4669FFB4FFBAFE70FB98FD20E538" box="[474,662,1081,1106]" metricMagnitude="2" metricUnit="m" metricValue="6.4" metricValueMax="6.8" metricValueMin="6.0" pageId="33" pageNumber="34" unit="m" value="640.0" valueMax="680.0" valueMin="600.0">
|
|||
|
<quantity id="856A0CBFFFB4FFBAFE70FB98FDEFE53B" box="[474,601,1081,1106]" metricMagnitude="2" metricUnit="m" metricValue="6.4" metricValueMax="6.8" metricValueMin="6.0" pageId="33" pageNumber="34" unit="m" value="640.0" valueMax="680.0" valueMin="600.0">600–680 m</quantity>
|
|||
|
a.s.l.
|
|||
|
</elevation>
|
|||
|
,
|
|||
|
<date id="362C879AFFB4FFBAFD08FB98FCBBE538" box="[674,781,1081,1106]" pageId="33" pageNumber="34" value="2017-03-08">
|
|||
|
<collectingDate id="26687E72FFB4FFBAFD08FB98FCBBE538" box="[674,781,1081,1106]" pageId="33" pageNumber="34" value="2017-03-08">8.iii.2017</collectingDate>
|
|||
|
</date>
|
|||
|
(
|
|||
|
<collectorName id="EF67C48CFFB4FFBAFCB7FB9BFC13E538" box="[797,933,1081,1106]" pageId="33" pageNumber="34">B.A. Huber</collectorName>
|
|||
|
)
|
|||
|
</materialsCitation>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DFBFCFDBEE440" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
<emphasis id="70E67D48FFB4FFBAFF6DFBFCFE4EE51C" bold="true" box="[199,504,1117,1142]" pageId="33" pageNumber="34">Other material examined</emphasis>
|
|||
|
.
|
|||
|
<collectingCountry id="3A85E1CAFFB4FFBAFDA2FBFFFD12E51C" box="[520,676,1118,1142]" name="Sri Lanka" pageId="33" pageNumber="34">SRI LANKA</collectingCountry>
|
|||
|
:
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFD1AFBFFFD65E51C" box="[688,723,1118,1142]" pageId="33" pageNumber="34" type="male">4♂</specimenCount>
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFD76FBFFFD48E51C" box="[732,766,1118,1142]" pageId="33" pageNumber="34" type="female">3♀</specimenCount>
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFCADFBFFFCFDE51C" box="[775,843,1117,1142]" pageId="33" pageNumber="34" type="generic">1 juv.</specimenCount>
|
|||
|
, ZFMK (Ar 20077), and
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFBC5FBFFFB24E51C" box="[1135,1170,1118,1142]" pageId="33" pageNumber="34" type="female">3♀</specimenCount>
|
|||
|
4 juvs in pure ethanol, ZFMK (SL109), same data as
|
|||
|
<typeStatus id="9D291FF8FFB4FFBAFE51FB20FDD7E5F3" box="[507,609,1153,1177]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
|||
|
.
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFDC4FB20FD25E5F3" box="[622,659,1153,1177]" pageId="33" pageNumber="34" type="female">4♀</specimenCount>
|
|||
|
, RMNH, Kandy (
|
|||
|
<geoCoordinate id="27A6C79DFFB4FFBAFCCFFB20FC0CE5F3" box="[869,954,1153,1177]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="7.3">7°18’N</geoCoordinate>
|
|||
|
,
|
|||
|
<geoCoordinate id="27A6C79DFFB4FFBAFC63FB20FB9FE5F3" box="[969,1065,1153,1177]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="80.63333">80°38’E</geoCoordinate>
|
|||
|
; unprecise coordinates!),
|
|||
|
<quantity id="856A0CBFFFB4FFBAFAF8FB20FA2DE5F3" box="[1362,1435,1153,1177]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="34" unit="m" value="600.0">600 m</quantity>
|
|||
|
a.s.l. (not
|
|||
|
<quantity id="856A0CBFFFB4FFBAFEAFFB04FEEFE5D7" box="[261,345,1189,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="33" pageNumber="34" unit="m" value="1600.0">1600 m</quantity>
|
|||
|
as on label and in
|
|||
|
<bibRefCitation id="2603DCABFFB4FFBAFD8DFB04FC89E5D4" author="Huber, B. A. & Benjamin, S." box="[551,831,1189,1214]" pageId="33" pageNumber="34" pagination="3305 - 3319" refId="ref27050" refString="Huber, B. A. & Benjamin, S. (2005) The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of Wanniyala new genus (Araneae: Pholcidae). Journal of Natural History, 39 (37), 3305 - 3319." type="journal article" year="2005">Huber & Benjamin 2005</bibRefCitation>
|
|||
|
), “forest and waterworks, in umbrella web under large tangle”,
|
|||
|
<date id="362C879AFFB4FFBAFF53FB68FE2DE588" box="[249,411,1225,1250]" pageId="33" pageNumber="34" value="1981-08-07" valueMax="1981-08-11" valueMin="1981-08-07">7–11.viii.1981</date>
|
|||
|
(collector not given) (misidentified as
|
|||
|
<taxonomicName id="8592DAD9FFB4FFBAFCC8FB6BFA98E58B" authority="in Huber & Benjamin 2005" authorityName="in Huber & Benjamin" authorityYear="2005" box="[866,1326,1225,1250]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="hakgala">
|
|||
|
<emphasis id="70E67D48FFB4FFBAFCC8FB6BFC57E588" box="[866,993,1225,1250]" italics="true" pageId="33" pageNumber="34">W. hakgala</emphasis>
|
|||
|
in
|
|||
|
<bibRefCitation id="2603DCABFFB4FFBAFBA1FB68FA98E58B" author="Huber, B. A. & Benjamin, S." box="[1035,1326,1225,1249]" pageId="33" pageNumber="34" pagination="3305 - 3319" refId="ref27050" refString="Huber, B. A. & Benjamin, S. (2005) The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of Wanniyala new genus (Araneae: Pholcidae). Journal of Natural History, 39 (37), 3305 - 3319." type="journal article" year="2005">Huber & Benjamin 2005</bibRefCitation>
|
|||
|
</taxonomicName>
|
|||
|
);
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFAEFFB68FADEE58B" box="[1349,1384,1225,1249]" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFAD8FB68FA23E58B" box="[1394,1429,1225,1249]" pageId="33" pageNumber="34" type="female">1♀</specimenCount>
|
|||
|
, RMNH, same data.
|
|||
|
<specimenCount id="54946AD3FFB4FFBAFE2EFB4FFE11E46C" box="[388,423,1262,1286]" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
|
|||
|
4 juvs, ZMUT (AA 3634), Matale District, Rattota [
|
|||
|
<geoCoordinate id="27A6C79DFFB4FFBAFBB7FB4CFB37E46C" box="[1053,1153,1261,1286]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="55" value="7.515">7.515°N</geoCoordinate>
|
|||
|
,
|
|||
|
<geoCoordinate id="27A6C79DFFB4FFBAFB3BFB4CFB49E46C" box="[1169,1279,1261,1286]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="55" value="80.673">80.673°E</geoCoordinate>
|
|||
|
], “in litter of djungle”,
|
|||
|
<date id="362C879AFFB4FFBAFEAFFAB0FEDFE440" box="[261,361,1297,1322]" pageId="33" pageNumber="34" value="1969-02-02">2.ii.1969</date>
|
|||
|
(P. Lehtinen).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="0A88F2D1FFB4FFB9FF6DFA94FEB8E63B" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DFA94FB16E4FC" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
<emphasis id="70E67D48FFB4FFBAFF6DFA94FE28E424" bold="true" box="[199,414,1333,1358]" pageId="33" pageNumber="34">Description. Male</emphasis>
|
|||
|
(
|
|||
|
<typeStatus id="9D291FF8FFB4FFBAFE04FA94FDA0E424" box="[430,534,1333,1358]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
|||
|
). MEASUREMENTS. Total body length 1.9, carapace width 0.80. Distance PME- PME 125 µm, diameter PME 100 µm, distance PME-ALE 30 µm; AME absent. Sternum width/length: 0.62/0.46. Leg 1: 13.8 (3.4 + 0.3 + 3.4 + 5.1 + 1.6), tibia 2: 2.0, tibia 3: 1.5, tibia 4: 2.1; tibia 1 L/d: 49.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DFA03FB16E74F" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">COLOR (in ethanol). Carapace ochre yellow with median and lateral dark bands; ocular area posteriorly dark; clypeus with pair of dark bands below eye triads; sternum mostly black, with indistinct small triangular light mark anteriorly; legs ochre-yellow, with dark rings on femora (subdistally) and tibiae (proximally and subdistally); abdomen ochre-gray with dark marks dorsally and laterally, ventrally also with dark pattern.</paragraph>
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DF993FB60E704" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
BODY. Habitus as in
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFE1DF990FDACE720" box="[439,538,1585,1610]" captionStart="FIGURES 126–137" captionStartId="32.[151,264,1429,1452]" captionTargetBox="[151,1436,193,1409]" captionTargetId="figure@32.[151,1436,193,1409]" captionTargetPageId="32" captionText="FIGURES 126–137. Wanniyala, live specimens, part 1. 126–128. W. agrabopath Huber & Benjamin, 2005, male and females with egg-sacs from Hakgala. 129–131. W. mudita sp. n., male and females with egg-sacs from Kandy. 132–134. W. orientalis sp. n., males and female with egg-sac from Inginiyagala. 135–137. W. upekkha sp. n., male from Dematagala and females with and without egg-sac from Ella." httpUri="https://zenodo.org/record/2625074/files/figure.png" pageId="33" pageNumber="34">Fig. 129</figureCitation>
|
|||
|
; ocular area slightly raised; carapace with shallow but distinct median furrow; clypeus with pair of processes connected by protruding arc (
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFC93F9F4FC6EE704" box="[825,984,1621,1646]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Figs 156–157</figureCitation>
|
|||
|
); sternum unmodified.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DF9DBFF79E7B0" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
CHELICERAE. As in
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFE17F9D8FDD6E7F8" box="[445,608,1657,1682]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Figs 156–157</figureCitation>
|
|||
|
, with pair of small apophyses proximally laterally and pair of long apophyses directed forwards, tips of apophyses distally directed downwards, distance between tips of apophyses: 0.64.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="422DA15AFFB4FFBAFF6DF947FCB7E600" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">
|
|||
|
PALPS. In general very similar to known congeners (cf.
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFCF4F944FBB7E794" box="[862,1025,1765,1790]" captionStart="FIGURES 150–155" captionStartId="36.[151,264,1893,1916]" captionTargetBox="[218,1340,196,1875]" captionTargetId="figure@36.[202,1387,195,1876]" captionTargetPageId="36" captionText="FIGURES 150–155. Wanniyala left male palps in prolateral and retrolateral views. 150–151. W. upekkha sp. n. from Dematagala (ZFMK Ar 20086). 152–153. W. viharekele sp. n. from Viharekele (ZFMK Ar 20091). 154–155. W. labugama sp. n. from Labugama (ZFMK Ar 20097). Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; p, procursus; pa, patella; pbs, proximal bulbal sclerite; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.5 mm." httpUri="https://zenodo.org/record/2625078/files/figure.png" pageId="33" pageNumber="34">Figs 150–155</figureCitation>
|
|||
|
); coxa unmodified; trochanter with several distinctive processes (
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFE40F8A8FDF8E648" box="[490,590,1801,1826]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Fig. 158</figureCitation>
|
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|
); femur with indistinct small dorsal apophysis proximally; tibia very long (0.86); procursus with several distinctive elements (
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFD46F88CFC38E62C" box="[748,910,1837,1862]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Figs 160–161</figureCitation>
|
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|
); bulb with simple membranous embolus and short, weakly curved and pointed apophysis (
|
|||
|
<figureCitation id="DAA9BDDFFFB4FFBAFD38F8F0FD42E600" box="[658,756,1873,1898]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="33" pageNumber="34">Fig. 159</figureCitation>
|
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).
|
|||
|
</paragraph>
|
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|
<paragraph id="422DA15AFFB4FFBAFF6DF8D7FD6EE6BC" blockId="33.[151,1437,757,2006]" pageId="33" pageNumber="34">LEGS. Without spines and curved hairs; with short vertical hairs in higher than usual density on all metatarsi (especially proximally); retrolateral trichobothrium on tibia 1 at 12%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~25 pseudosegments, distally distinct.</paragraph>
|
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|
<caption id="16EDF1D2FFB7FFB9FF3DFACAFC86E482" httpUri="https://zenodo.org/record/2625076/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,264,1387,1410]" targetBox="[151,1436,195,1366]" targetPageId="34">
|
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|
<paragraph id="422DA15AFFB7FFB9FF3DFACAFC86E482" blockId="34.[151,1437,1387,1512]" pageId="34" pageNumber="35">
|
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|
<emphasis id="70E67D48FFB7FFB9FF3DFACAFED1E4EB" bold="true" box="[151,359,1387,1410]" pageId="34" pageNumber="35">FIGURES 138–149</emphasis>
|
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.
|
|||
|
<taxonomicName id="8592DAD9FFB7FFB9FEDEFACDFE69E4E8" authorityName="Huber & Benjamin" authorityYear="2005" box="[372,479,1388,1410]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FEDEFACDFE69E4E8" box="[372,479,1388,1410]" italics="true" pageId="34" pageNumber="35">Wanniyala</emphasis>
|
|||
|
</taxonomicName>
|
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|
, live specimens, part 2. 138–140.
|
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|
<taxonomicName id="8592DAD9FFB7FFB9FC94FACCFC20E4E8" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[830,918,1388,1410]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ohiya" status="sp. nov.">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FC94FACCFC20E4E8" box="[830,918,1388,1410]" italics="true" pageId="34" pageNumber="35">W. ohiya</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="70E67D48FFB7FFB9FC37FACDFC61E4EB" bold="true" box="[925,983,1388,1409]" pageId="34" pageNumber="35">
|
|||
|
<taxonomicNameLabel id="6BD5C033FFB7FFB9FC37FACDFC61E4EB" box="[925,983,1388,1409]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, males and female with egg-sac from Ohiya (Fig. 139 shows a male in typical resting position). 141–143.
|
|||
|
<taxonomicName id="8592DAD9FFB7FFB9FCADFA2FFC26E4C9" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[775,912,1421,1443]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="viharekele" status="sp. nov.">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FCADFA2FFC26E4C9" box="[775,912,1421,1443]" italics="true" pageId="34" pageNumber="35">W. viharekele</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="70E67D48FFB7FFB9FC3CFA2FFC79E4C9" bold="true" box="[918,975,1422,1443]" pageId="34" pageNumber="35">
|
|||
|
<taxonomicNameLabel id="6BD5C033FFB7FFB9FC3CFA2FFC79E4C9" box="[918,975,1422,1443]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, males from Viharekele and Kottawa, female from Kottawa. 144–146.
|
|||
|
<taxonomicName id="8592DAD9FFB7FFB9FE35FA11FD90E4AE" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[415,550,1454,1477]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="mapalena" status="sp. nov.">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FE35FA11FD90E4AE" box="[415,550,1454,1477]" italics="true" pageId="34" pageNumber="35">W. mapalena</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="70E67D48FFB7FFB9FD85FA0EFDDDE4AE" bold="true" box="[559,619,1455,1476]" pageId="34" pageNumber="35">
|
|||
|
<taxonomicNameLabel id="6BD5C033FFB7FFB9FD85FA0EFDDDE4AE" box="[559,619,1455,1476]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, male and females from between Ratnapura and Mapalena Ella. 147–149.
|
|||
|
<taxonomicName id="8592DAD9FFB7FFB9FA29FA11FF4AE482" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="labugama" status="sp. nov.">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FA29FA11FF4AE482" italics="true" pageId="34" pageNumber="35">W. labugama</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="70E67D48FFB7FFB9FEA8FA73FE8DE48D" bold="true" box="[258,315,1490,1511]" pageId="34" pageNumber="35">
|
|||
|
<taxonomicNameLabel id="6BD5C033FFB7FFB9FEA8FA73FE8DE48D" box="[258,315,1490,1511]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, males and female with egg-sac from Labugama.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<paragraph id="422DA15AFFB7FFB9FF6DF99DFAC0E73E" blockId="34.[151,1437,1596,1909]" box="[199,1398,1596,1621]" pageId="34" pageNumber="35">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FF6DF99DFEB2E73F" bold="true" box="[199,260,1596,1621]" pageId="34" pageNumber="35">Male</emphasis>
|
|||
|
(variation). Tibia
|
|||
|
<quantity id="856A0CBFFFB7FFB9FE7AF99DFE4DE73F" box="[464,507,1596,1621]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="1.0">1 in</quantity>
|
|||
|
four other males from
|
|||
|
<typeStatus id="9D291FF8FFB7FFB9FD54F99CFC98E73F" box="[766,814,1597,1621]" pageId="34" pageNumber="35">type</typeStatus>
|
|||
|
locality: 3.1, 3.2, 3.2, 3.4; in male from Rattota 3.7.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="422DA15AFFB7FFB9FF6DF9C1FC85E663" blockId="34.[151,1437,1596,1909]" pageId="34" pageNumber="35">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FF6DF9C1FEA8E713" bold="true" box="[199,286,1632,1657]" pageId="34" pageNumber="35">Female</emphasis>
|
|||
|
. In general similar to male (
|
|||
|
<figureCitation id="DAA9BDDFFFB7FFB9FDF6F9C1FD4EE712" box="[604,760,1632,1657]" captionStart="FIGURES 126–137" captionStartId="32.[151,264,1429,1452]" captionTargetBox="[151,1436,193,1409]" captionTargetId="figure@32.[151,1436,193,1409]" captionTargetPageId="32" captionText="FIGURES 126–137. Wanniyala, live specimens, part 1. 126–128. W. agrabopath Huber & Benjamin, 2005, male and females with egg-sacs from Hakgala. 129–131. W. mudita sp. n., male and females with egg-sacs from Kandy. 132–134. W. orientalis sp. n., males and female with egg-sac from Inginiyagala. 135–137. W. upekkha sp. n., male from Dematagala and females with and without egg-sac from Ella." httpUri="https://zenodo.org/record/2625074/files/figure.png" pageId="34" pageNumber="35">Figs 130–131</figureCitation>
|
|||
|
) but clypeus unmodified and legs with usual low number of short vertical hairs; two females with entirely black sternum. Tibia
|
|||
|
<quantity id="856A0CBFFFB7FFB9FC31F925FC7EE7F7" box="[923,968,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="1.0">1 in</quantity>
|
|||
|
<specimenCount id="54946AD3FFB7FFB9FC7BF925FBF5E7F7" box="[977,1091,1668,1693]" pageId="34" pageNumber="35" type="female">3 females</specimenCount>
|
|||
|
: 2.6, 2.7, 2.8. Epigynum as in
|
|||
|
<figureCitation id="DAA9BDDFFFB7FFB9FF3DF909FE82E7AB" box="[151,308,1704,1729]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="34" pageNumber="35">Figs 162–163</figureCitation>
|
|||
|
, strongly protruding and pointed in lateral view; posterior plate apparently reduced(?) to two lateral sclerites mostly hidden behind epigynal plate and poorly visible in ventral view; internal genitalia as in
|
|||
|
<figureCitation id="DAA9BDDFFFB7FFB9FA99F96DFA21E78F" box="[1331,1431,1740,1765]" captionStart="FIGURES 156–164" captionStartId="37.[151,264,1806,1829]" captionTargetBox="[152,1432,193,1784]" captionTargetId="figure@37.[151,1436,193,1785]" captionTargetPageId="37" captionText="FIGURES 156–164. Wanniyala mudita sp. n., from Kandy (ZFMK Ar 20077). 156–157. Male chelicerae, clypeus and ocular area, frontal and lateral views. 158. Left male palpal trochanter and femur, retrolateral view. 159. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 160–161. Left procursus, prolateral and retrolateral views. 162–163. Epigynum, ventral and lateral views. 164. Cleared female genitalia, dorsal view (arrow points at anterior ‘valve’). Abbreviations: ba, bulbal apophysis; e, embolus; pbs, proximal bulbal sclerite; ps1, ps2, prolateral sclerites 1 and 2; ptp, prolateral transparent process; rs, retrolateral sclerite; vtp, ventral transparent process. Scale lines: 0.3 mm (158–161), 0.5 mm (156–157, 162–164)." httpUri="https://zenodo.org/record/2625080/files/figure.png" pageId="34" pageNumber="35">Figs 164</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="DAA9BDDFFFB7FFB9FF3DF951FF72E662" box="[151,196,1776,1800]" captionStart="FIGURES 213–219" captionStartId="49.[151,264,1720,1743]" captionTargetBox="[151,1436,412,1657]" captionTargetId="figure@49.[151,1436,412,1657]" captionTargetPageId="49" captionText="FIGURES 213–219. Wanniyala spp., cleared female genitalia in dorsal views. 213. W. mudita sp. n., from Kandy (ZFMK Ar 20077). 214. W. orientalis sp. n., from Dimbulagala (ZFMK Ar 20079). 215. W. upekkha sp. n., from Dematagala (ZFMK Ar 20086). 216. W. ohiya sp. n., from Ohiya (ZFMK Ar 20089). 217. W. viharekele sp. n., from Viharekele (ZFMK Ar 20092). 218. W. mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 219. W. labugama sp. n., from Labugama (ZFMK Ar 20097). At various scales." httpUri="https://zenodo.org/record/2625096/files/figure.png" pageId="34" pageNumber="35">213</figureCitation>
|
|||
|
, with wide anterior ‘valve’ and pair of oval pore plates.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="422DA15AFFB7FFB9FF6DF8B5FEB8E63B" blockId="34.[151,1437,1596,1909]" pageId="34" pageNumber="35">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FF6DF8B5FECBE647" bold="true" box="[199,381,1812,1837]" pageId="34" pageNumber="35">Natural history</emphasis>
|
|||
|
. The spiders were found in the leaf litter, with their small webs slightly projecting from under the leaves.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="0A88F2D1FFB7FFB9FF6DF8FDFC46E61F" box="[199,1008,1884,1909]" pageId="34" pageNumber="35" type="distribution">
|
|||
|
<paragraph id="422DA15AFFB7FFB9FF6DF8FDFC46E61F" blockId="34.[151,1437,1596,1909]" box="[199,1008,1884,1909]" pageId="34" pageNumber="35">
|
|||
|
<emphasis id="70E67D48FFB7FFB9FF6DF8FDFEEFE61F" bold="true" box="[199,345,1884,1909]" pageId="34" pageNumber="35">Distribution</emphasis>
|
|||
|
. Known from two localities in central
|
|||
|
<collectingCountry id="3A85E1CAFFB7FFB9FCA1F8FDFCCFE61F" box="[779,889,1884,1909]" name="Sri Lanka" pageId="34" pageNumber="35">Sri Lanka</collectingCountry>
|
|||
|
(
|
|||
|
<figureCitation id="DAA9BDDFFFB7FFB9FC22F8FDFC51E61F" box="[904,999,1884,1909]" captionStart="FIGURES 226–227" captionStartId="53.[151,264,898,921]" captionTargetBox="[151,1436,183,885]" captionTargetId="figure@53.[151,1436,183,885]" captionTargetPageId="53" captionText="FIGURES 226–227. Known distribution of Wanniyala." httpUri="https://zenodo.org/record/2625104/files/figure.png" pageId="34" pageNumber="35">Fig. 226</figureCitation>
|
|||
|
)
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|