306 lines
35 KiB
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306 lines
35 KiB
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<document id="D5BC50D7430455D6A850B33B86E03B70" ID-DOI="10.1206/0003-0082(2000)293<0001:TEFOSB>2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="10110982" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630265931376" checkinUser="carolina" docAuthor="CAMARGO, JOÃO M. F., GRIMALDI, DAVID, A. & PEDRO, SILVIA R. M." docDate="2000" docId="03C09D45FFD8FF86AC0B2036F603FA64" docLanguage="en" docName="N3293.pdf" docOrigin="American Museum Novitates 3293" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29293%3C0001%3ATEFOSB%3E2.0.CO%3B2" docStyle="DocumentStyle:059C9B7006157ECE636E30DEF6793B3A.3:AmMusNovit.2000-2010.journal_article.type2" docStyleId="059C9B7006157ECE636E30DEF6793B3A" docStyleName="AmMusNovit.2000-2010.journal_article.type2" docStyleVersion="3" docTitle="Proplebeia vetusta CAMARGO & GRIMALDI & PEDRO 2000, new species" docType="treatment" docVersion="1" masterDocId="FFF9E53DFFD2FF8AAD512547F43BFFE4" masterDocTitle="The Extinct Fauna of Stingless Bees (Hymenoptera: Apidae: Meliponini) in Dominican Amber: Two New Species and Redescription of the Male of Proplebeia dominicana (Wille and Chandler)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1730300627421" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="EE4088191C5644B61E370189FA17A1CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6DB3B4F35EC61AACFDB6EF3013347AA5">
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<mods:title id="E35DEB8224AC89F31F56225E2C1126D0">The Extinct Fauna of Stingless Bees (Hymenoptera: Apidae: Meliponini) in Dominican Amber: Two New Species and Redescription of the Male of Proplebeia dominicana (Wille and Chandler)</mods:title>
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</mods:titleInfo>
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<mods:name id="A67C6AFBE3C5A695ED0F0809A432183E" type="personal">
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<mods:role id="BC13435E96FC2D0370AF6FF3980D1609">
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<mods:roleTerm id="8F27E37FF3D2375BF16F3B776F9314D4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E3D072DF55D7C5A783B078BCF9D5BAF4">CAMARGO, JOÃO M. F.</mods:namePart>
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<mods:affiliation id="6C88091B3BBAF5ADA0FB163ED12263FE">Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto — USP, Av. Bandeirantes, 3900, 14040 901 — Ribeirão Preto, SP, Brasil (JMFC is CNPq fellow researcher, Proc. 300014 / 84 8 RN)</mods:affiliation>
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<mods:nameIdentifier id="10968AEE4ADE0721B9BA34C12FF37BBC" type="email">jmfdcama@usp.br & jmafcama@ffclrp.usp.br</mods:nameIdentifier>
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</mods:name>
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<mods:name id="419E330FCF80514AB54A0019347E54B7" type="personal">
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<mods:role id="B595936C98CD1F668057E15FCDDB32C2">
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<mods:roleTerm id="79AE2979E18001701CADCF8632F01E46">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9A0476A5AEE37F0820DD7D5FD0B0392C">GRIMALDI, DAVID, A.</mods:namePart>
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<mods:affiliation id="890CEC2FB1544294397226791254BAAF">Division of Invertebrate Zoology, American Museum of Natural History</mods:affiliation>
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<mods:nameIdentifier id="091FDFDA3A95C2F5088BC67C43956B72" type="email">grimaldi@amnh.org</mods:nameIdentifier>
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</mods:name>
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<mods:name id="999A2D484F9A2A56B41998E6121877A0" type="personal">
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<mods:role id="50681DFB3499B8ED8E19D942E092B0F2">
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<mods:roleTerm id="9800B03D13862AC1857FDE42334DE6E8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="81145B798647B7C379BAD45EFB16CF8A">PEDRO, SILVIA R. M.</mods:namePart>
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<mods:affiliation id="3693FEB5DF4B1DCC1DF88ED868CC1776">Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto — USP, Av. Bandeirantes, 3900, 14040 901 — Ribeirão Preto, SP, Brasil (JMFC is CNPq fellow researcher, Proc. 300014 / 84 8 RN)</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="A9ABBC47044BC45DD49620474101942E">text</mods:typeOfResource>
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<mods:relatedItem id="BAF023F1905B721BCE652E0A5C733FCB" type="host">
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<mods:titleInfo id="FCDF32F7BBA93C7CC4B13C4D1F4B998F">
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<mods:title id="AB4F5FE23873520BD8338274366D4309">American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part id="9A9A1E01CE1D00477A4F3215D73B35BA">
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<mods:date id="B08402DBFC81DDFCC95A7CF1139CB96E">2000</mods:date>
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<mods:detail id="F035FE265AC6539973C2106C913357BE" type="pubDate">
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<mods:number id="41BAE2EEB662D2E509482F8B18076563">2000-03-16</mods:number>
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</mods:detail>
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<mods:detail id="E4F2D7033028F0139CB80E84FA0F5E5B" type="volume">
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<mods:number id="EEDD1BE7B6673623BBC395454A673E07">3293</mods:number>
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</mods:detail>
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<mods:extent id="479D0ABB448800F40DEF389A381B30B3" unit="page">
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<mods:start id="5EBC0AF241BBDF985EBC2BD0A5C87844">1</mods:start>
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<mods:end id="326BC1CE319001B3726CC6F7449621F7">1</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="55DFB6DEAA96FB03C81CA0230E333760">
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<mods:url id="5A0230103055B85C57168059F6D66924">http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29293%3C0001%3ATEFOSB%3E2.0.CO%3B2</mods:url>
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</mods:location>
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<mods:classification id="5B0B16081F47D9A4687E0EF5157DCE19">journal article</mods:classification>
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<mods:identifier id="7CC147C752A2C4B4E9D4C8C44DEE35BF" type="DOI">10.1206/0003-0082(2000)293<0001:TEFOSB>2.0.CO;2</mods:identifier>
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<mods:identifier id="CE80E2E88D0FAD1BF7E39A1CBD1AB945" type="ISSN">0003-0082</mods:identifier>
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<mods:identifier id="6734ABE9F6A77D20ED163DADC5FAAA8F" type="Zenodo-Dep">10110982</mods:identifier>
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</mods:mods>
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<treatment id="03C09D45FFD8FF86AC0B2036F603FA64" LSID="urn:lsid:plazi:treatment:03C09D45FFD8FF86AC0B2036F603FA64" httpUri="http://treatment.plazi.org/id/03C09D45FFD8FF86AC0B2036F603FA64" lastPageId="12" pageId="10">
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<subSubSection id="C3737FD8FFD8FF80AC0B2036F6EDFA6C" box="[346,726,1393,1416]" pageId="10" type="nomenclature">
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<paragraph id="8BD62C53FFD8FF80AC0B2036F6EDFA6C" blockId="10.[346,726,1393,1416]" box="[346,726,1393,1416]" pageId="10">
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<taxonomicName id="4C6957D0FFD8FF80AC0B2036F60CFA6C" authority="CAMARGO & GRIMALDI & PEDRO, 2000" authorityName="CAMARGO & GRIMALDI & PEDRO" authorityYear="2000" box="[346,567,1393,1416]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="vetusta" status="sp. nov.">
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<emphasis id="B91DF041FFD8FF80AC0B2036F60CFA6C" box="[346,567,1393,1416]" italics="true" pageId="10">Proplebeia vetusta</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22E4D3AFFD8FF80AF192036F6EDFA6C" box="[584,726,1393,1416]" pageId="10" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3737FD8FFD8FF80ACF820D7F6BCFA41" box="[425,647,1424,1445]" pageId="10" type="description">
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<paragraph id="8BD62C53FFD8FF80ACF820D7F6BCFA41" blockId="10.[425,647,1424,1445]" box="[425,647,1424,1445]" pageId="10">
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<heading id="D09E9B3FFFD8FF80ACF820D7F6BCFA41" box="[425,647,1424,1445]" centered="true" fontSize="36" level="3" pageId="10" reason="7">
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<figureCitation id="135230D6FFD8FF80ACF820D7F635FA41" box="[425,526,1424,1445]" captionStart="Figs" captionStartId="10.[298,346,1267,1288]" captionTargetBox="[340,1310,189,1236]" captionTargetId="figure-39@10.[331,1318,176,1245]" captionTargetPageId="10" captionText="Figs. 8–10. Forewing, worker; 8. Proplebeia vetusta, sp.n., holotype, specimen, AMNHDR14854; 9. Proplebeia dominicana, specimen AMNHDR141175; 10. Proplebeia tantilla, sp. n., paratype, specimen AMNHDR14911. Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10110992" httpUri="https://zenodo.org/record/10110992/files/figure.png" pageId="10">Figures 8</figureCitation>
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,
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<figureCitation id="135230D6FFD8FF80AF4D20D7F65BFA41" box="[540,608,1424,1445]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="13.[264,312,1205,1226]" captionStartId-1="14.[341,379,1605,1626]" captionTargetBox-0="[257,1323,189,1173]" captionTargetBox-1="[484,1174,184,1574]" captionTargetId-0="figure-36@13.[250,1331,176,1184]" captionTargetId-1="figure-37@14.[468,1180,176,1584]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Figs. 11–13. Proplebeia vetusta, sp. n., holotype, AMNHDR141481, head, detail of mandible, tibia and tarsus III, detail of penicillum and rastellum. Minor scale bar = 1.0 mm (fig. 11), large scale bar = 1.0 mm (figs. 12–13)." captionText-1="Fig. 14. Proplebeia vetusta, sp. n., holotype, AMNHDR141481. Scale bar = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10110994" figureDoi-1="http://doi.org/10.5281/zenodo.10110996" httpUri-0="https://zenodo.org/record/10110994/files/figure.png" httpUri-1="https://zenodo.org/record/10110996/files/figure.png" pageId="10">11–14</figureCitation>
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,
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<figureCitation id="135230D6FFD8FF80AF3C20D7F6BCFA41" box="[621,647,1424,1445]" captionStart="Fig" captionStartId="19.[264,302,1683,1704]" captionTargetBox="[352,1227,189,1661]" captionTargetId="figure-34@19.[352,1227,189,1661]" captionTargetPageId="19" captionText="Fig. 22. Photomicrographs of Proplebeia vetusta, sp. n., worker, paratype, AMNHDR141440. Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10111004" httpUri="https://zenodo.org/record/10111004/files/figure.png" pageId="10">22</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3737FD8FFD8FF80AC4120FFF550FA0C" pageId="10" type="reference_group">
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<paragraph id="8BD62C53FFD8FF80AC4120FFF550FA0C" blockId="10.[272,800,1464,1512]" pageId="10">
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<taxonomicName id="4C6957D0FFD8FF80AC4120FFF5B7FA29" box="[272,396,1464,1485]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B91DF041FFD8FF80AC4120FFF55BFA29" box="[272,352,1464,1485]" italics="true" pageId="10">Plebeia</emphasis>
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sp.
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</taxonomicName>
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;
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<bibRefCitation id="EFF851A2FFD8FF80ACCC20FFF612FA29" author="Poinar, G. O., Jr." box="[413,553,1464,1485]" pageId="10" refId="ref11819" refString="1994" type="journal article" year="1994">Poinar, 1994</bibRefCitation>
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b: 73, fig 2 (fossil review).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3737FD8FFD8FF80AC7C20BBF7C6FA1F" pageId="10" type="diagnosis">
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<paragraph id="8BD62C53FFD8FF80AC7C20BBF7C6FA1F" blockId="10.[272,801,1531,1731]" lastBlockId="10.[848,1377,1393,1731]" pageId="10">
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DIAGNOSIS:
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<emphasis id="B91DF041FFD8FF80ACE720BCF61BF9F7" bold="true" box="[438,544,1531,1555]" pageId="10">Worker.</emphasis>
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Differs from
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<taxonomicName id="4C6957D0FFD8FF80AF9220BBF55CF9D4" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="dominicana">
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<emphasis id="B91DF041FFD8FF80AF9220BBF55CF9D4" italics="true" pageId="10">P. dominicana</emphasis>
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</taxonomicName>
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by the large body size (
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<quantity id="4C9181B6FFD8FF80AFF6235EF57AF9A9" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="10" unit="mm" value="4.1" valueMax="4.4" valueMin="3.8">3.80–4.40 mm</quantity>
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;
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<taxonomicName id="4C6957D0FFD8FF80AC0B2370F62BF9AA" box="[346,528,1591,1614]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="dominicana">
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<emphasis id="B91DF041FFD8FF80AC0B2370F62BF9AA" box="[346,528,1591,1614]" italics="true" pageId="10">P. dominicana</emphasis>
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</taxonomicName>
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, 3.0 mm), forewing length (2.88–3.16;
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<taxonomicName id="4C6957D0FFD8FF80ACA12313F6A0F98F" box="[496,667,1620,1643]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="dominicana">
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<emphasis id="B91DF041FFD8FF80ACA12313F6A0F98F" box="[496,667,1620,1643]" italics="true" pageId="10">P. dominicana</emphasis>
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</taxonomicName>
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,
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<quantity id="4C9181B6FFD8FF80AFFB2313F579F96C" metricMagnitude="-3" metricUnit="m" metricValue="2.625" metricValueMax="2.75" metricValueMin="2.5" pageId="10" unit="mm" value="2.625" valueMax="2.75" valueMin="2.5">2.50–2.75 mm</quantity>
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); malar area a little longer than diameter of scape; clypeus yellowish on distal third; parocular areas with a yellow stripe, slightly truncate above the tentorial pits (in
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<typeStatus id="54D292F1FFD8FF80A8722036F7BCFA41" pageId="10" type="holotype">holotype</typeStatus>
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); and mandible with only a slight emargination separating the two denticles (in
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<taxonomicName id="4C6957D0FFD8FF80A81620ECF7E2FA3B" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="dominicana">
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<emphasis id="B91DF041FFD8FF80A81620ECF7E2FA3B" italics="true" pageId="10">P. dominicana</emphasis>
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</taxonomicName>
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the emargination between denticles is deep).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3737FD8FFD8FF81AE3F2347F0B6F941" lastPageId="11" pageId="10" type="description">
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<paragraph id="8BD62C53FFD8FF81AE3F2347F0B6F941" blockId="10.[848,1377,1393,1731]" lastBlockId="11.[814,1343,182,1731]" lastPageId="11" pageId="10">
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DESCRIPTION:
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<emphasis id="B91DF041FFD8FF80A94F2347F0BEF9FC" bold="true" box="[1054,1157,1536,1560]" pageId="10">Worker</emphasis>
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(AMNHDR14 1481).
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<emphasis id="B91DF041FFD8FF80AEE5235AF063F9D1" bold="true" box="[948,1112,1565,1589]" pageId="10">Dimensions:</emphasis>
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Approximate body length
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<quantity id="4C9181B6FFD8FF80AEE1237DF01DF9B5" box="[944,1062,1594,1617]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="10" unit="mm" value="3.8">3.80 mm</quantity>
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(metasoma quite compressed; in
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<typeStatus id="54D292F1FFD8FF80AEB92311F05BF989" box="[1000,1120,1622,1645]" pageId="10" type="paratype">paratypes</typeStatus>
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AMNHDR14854 and DR141440 approximate body length 4.40 and
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<quantity id="4C9181B6FFD8FF80AE9823C8F001F942" box="[969,1082,1679,1702]" metricMagnitude="-3" metricUnit="m" metricValue="4.32" pageId="10" unit="mm" value="4.32">4.32 mm</quantity>
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, respectively); forewing length
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<quantity id="4C9181B6FFD8FF80AEF223EBF035F927" box="[931,1038,1708,1731]" metricMagnitude="-3" metricUnit="m" metricValue="2.88" pageId="10" unit="mm" value="2.88">2.88 mm</quantity>
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from apex of costal sclerite to wing tip (including tegula,
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<quantity id="4C9181B6FFD9FF81AF0E25F1F6F5FF29" box="[607,718,182,205]" metricMagnitude="-3" metricUnit="m" metricValue="3.24" pageId="11" unit="mm" value="3.24">3.24 mm</quantity>
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), in the
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<typeStatus id="54D292F1FFD9FF81AC482593F546FF0F" box="[281,381,212,235]" pageId="11" type="paratype">paratype</typeStatus>
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DR14854,
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|
|
<quantity id="4C9181B6FFD9FF81AF492593F644FF0F" box="[536,639,212,235]" metricMagnitude="-3" metricUnit="m" metricValue="3.16" pageId="11" unit="mm" value="3.16">3.16 mm</quantity>
|
|||
|
|
(including tegula,
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AC1625B6F58DFEEC" box="[327,438,241,264]" metricMagnitude="-3" metricUnit="m" metricValue="3.72" pageId="11" unit="mm" value="3.72">3.72 mm</quantity>
|
|||
|
|
), and in DR141440, 3.00 mm (including tegula,
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AF502449F64AFEC1" box="[513,625,270,293]" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="11" unit="mm" value="3.44">3.44 mm</quantity>
|
|||
|
|
); maximum head width
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AC2A246BF5D3FEA7" box="[379,488,300,323]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="11" unit="mm" value="1.55">1.55 mm</quantity>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B91DF041FFD9FF81ACA7246CF6C5FEA7" bold="true" box="[502,766,299,323]" pageId="11">Color of integument:</emphasis>
|
|||
|
|
Black, including antennal scapes, except tarsomeres, chestnut and trochanters discolored, translucent; head with following parts diffuse yellowish: distal third of clypeus, lower parocular area to tentorial pit, supraclypeal area between antennal alveolus, and labrum; mandibles dark chestnut. Mesonotum black, devoid of yellow markings. Forewing C and R veins and tegulae dark chestnut; pterostigma with translucent spot in middle; wing membrane hyaline.
|
|||
|
|
<emphasis id="B91DF041FFD9FF81ACCD2729F639FD62" bold="true" box="[412,514,622,646]" pageId="11">Pilosity:</emphasis>
|
|||
|
|
Pale yellow over entire body. Decumbent, pale, minute plumose hairs covering entire face. Erect, unbranched hairs sparse on upper half of frons and increasing in number and length toward vertex; longest hairs on vertex ca.
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AF7A2646F6ACFCFC" box="[555,663,769,792]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="11" unit="mm" value="0.17">0.17 mm</quantity>
|
|||
|
|
; ones on scape
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AC682659F592FCD1" box="[313,425,798,821]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="11" unit="mm" value="0.04">0.04 mm</quantity>
|
|||
|
|
. Labial palpi with 3–4 long hairs (
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AC6B267BF5D4FCB7" box="[314,495,828,851]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" pageId="11" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18 mm</quantity>
|
|||
|
|
). Longest hairs on labrum
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AC69261EF593FC94" box="[312,424,857,880]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="11" unit="mm" value="0.22">0.22 mm</quantity>
|
|||
|
|
. Mesoscutum with fine, decumbent, unbranched micropilosity a little sparser than on head; erect hairs sparser on discal area and more abundant and longer (
|
|||
|
|
<quantity id="4C9181B6FFD9FF81ADA72689F558FC01" box="[246,355,974,997]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="11" unit="mm" value="0.14">0.14 mm</quantity>
|
|||
|
|
) on anterior corners; longest hairs on scutellum
|
|||
|
|
<quantity id="4C9181B6FFD9FF81ACDC26ABF5CEFBE7" box="[397,501,1004,1027]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="11" unit="mm" value="0.18">0.18 mm</quantity>
|
|||
|
|
on ventral part of mesepisterna
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AC3D214EF620FBC4" box="[364,539,1033,1056]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="11" unit="mm" value="0.14" valueMax="0.16" valueMin="0.12">0.12–0.16 mm</quantity>
|
|||
|
|
. Basal area of propodeum glabrous. Posterior border of tibia III with unbranched hairs, the longest ones ca.
|
|||
|
|
<quantity id="4C9181B6FFD9FF81ADBF2126F561FB9C" box="[238,346,1121,1144]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="11" unit="mm" value="0.18">0.18 mm</quantity>
|
|||
|
|
. Only basal half of inner surface of tibia III visible (fig. 14); in wider visible part, corresponding approximately to middle of tibia, keirotrichiate area ca.
|
|||
|
|
<quantity id="4C9181B6FFD9FF81AF1721FEF689FB34" box="[582,690,1209,1232]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="11" unit="mm" value="0.2">0.20 mm</quantity>
|
|||
|
|
wide, posterior rim glabrous (clearly depressed, forming a step), ca.
|
|||
|
|
<quantity id="4C9181B6FFD9FF81ACBE21B3F667FAEF" box="[495,604,1268,1291]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="11" unit="mm" value="0.08">0.08 mm</quantity>
|
|||
|
|
wide. Externally tibiae partially covered by amorphous substance (pollen?) and air bubbles, but it is possible to verify that the penicillum is normal, as in
|
|||
|
|
<taxonomicName id="4C6957D0FFD9FF81AC21202EF5F2FA64" authorityName="Schwarz" authorityYear="1938" box="[368,457,1385,1408]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B91DF041FFD9FF81AC21202EF5F2FA64" box="[368,457,1385,1408]" italics="true" pageId="11">Plebeia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(s.s.); posterior parapenicillum with long, slender hairs and anterior parapenicillum well differentiated with a row of at least eight stiff hairs; rastellum apex visible with at least seven stiff, spinelike hairs. Microtrichia on wing membrane ferruginous and as long as hair interspaces (ca.
|
|||
|
|
<quantity id="4C9181B6FFD9FF81ADBF2371F55CF9A9" box="[238,359,1590,1613]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="11" unit="mm" value="0.02">0.02 mm</quantity>
|
|||
|
|
). T1 practically glabrous, only some minute hairs on sides; other terga and sterna covered by wings, legs, bubbles, and a whitish substance.
|
|||
|
|
<emphasis id="B91DF041FFD9FF81ACAB23C9F6A8F942" bold="true" box="[506,659,1678,1702]" pageId="11">Integument:</emphasis>
|
|||
|
|
Smooth and shiny as in
|
|||
|
|
<taxonomicName id="4C6957D0FFD9FF81ACF623EBF681F927" box="[423,698,1708,1731]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="species" species="dominicana">
|
|||
|
|
<emphasis id="B91DF041FFD9FF81ACF623EBF681F927" box="[423,698,1708,1731]" italics="true" pageId="11">Proplebeia dominicana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; only piligerous punctures present.
|
|||
|
|
<emphasis id="B91DF041FFD9FF81A9D425F1F7A5FF0F" bold="true" pageId="11">Form and proportions:</emphasis>
|
|||
|
|
Head 1.2× wider than long (1.55: 1.28, length measured from the apex of clypeus to upper tangent of median ocellus), a little narrower than mesosoma (1.66, measured across mesepisterna). Eyes 2.38× longer than wide (1.00:0.42), slightly convergent below; upper interorbital distance 1.01, maximum distance 1.10 and lower distance 0.89. Malar area a little longer than diameter of scape (0.12:0.10). Clypeus 2.23× wider than long (0.80:0.36), slightly convex. Labrum normal. Distance between clypeus and median ocellus 0.81. Interalveolar distance a little longer than diameter of antennal alveolus (0.19: 0.16), but shorter than alveolorbital distance (0.22). Frons slightly depressed along median line. Distance between lateral ocelli ca. 3.0× diameter of median ocellus and 1.5× ocellorbital distance (0.36:0.12: 0.24). Vertex rounded behind ocelli; preoccipital ridge slightly rounded (as in
|
|||
|
|
<taxonomicName id="4C6957D0FFD9FF81A98F2658F10CFCD2" authorityName="Schwarz" authorityYear="1938" box="[1246,1335,799,822]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|||
|
|
<emphasis id="B91DF041FFD9FF81A98F2658F10CFCD2" box="[1246,1335,799,822]" italics="true" pageId="11">Plebeia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicNameLabel id="A22E4D3AFFD9FF81AE7F267BF768FCB7" box="[814,851,828,851]" pageId="11" sensu="stricto">s.s.</taxonomicNameLabel>
|
|||
|
|
). Scape length 5.4× its diameter (0.54: 0.10), and ca.
|
|||
|
|
<superScript id="7C1C811BFFD9FF81AEB2261FF7D1FC82" attach="right" box="[995,1002,856,870]" fontSize="6" pageId="11">5</superScript>
|
|||
|
|
<emphasis id="B91DF041FFD9FF81AEBB261DF7D5FC96" bold="true" box="[1002,1006,858,882]" pageId="11">⁄</emphasis>
|
|||
|
|
<subScript id="17ED2E16FFD9FF81AEBC2622F7CDFC97" attach="left" box="[1005,1014,869,883]" fontSize="6" pageId="11">6</subScript>
|
|||
|
|
distance between antennal alveolus and lateral ocellus (0.64). Flagellum plus pedicel (1.08) ca. 2.0× length of scape; flagellomeres shorter than wide (the second one, 0.08:0.12). Scutellum with distal border semicircular, slightly projecting over metanotum, 2.06× wider than long (0.70: 0.34), basal pit normal (as in
|
|||
|
|
<emphasis id="B91DF041FFD9FF81A9002160F08AFBDA" box="[1105,1201,1063,1086]" italics="true" pageId="11">
|
|||
|
|
<taxonomicName id="4C6957D0FFD9FF81A9002160F096FBDA" authorityName="Schwarz" authorityYear="1938" box="[1105,1197,1063,1086]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="genus" sensu="stricto">Plebeia</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicNameLabel id="A22E4D3AFFD9FF81A9EC2161F0D9FBD9" box="[1213,1250,1062,1085]" pageId="11" sensu="stricto">s.s.</taxonomicNameLabel>
|
|||
|
|
). Forewing 2.32× longer than wide (2.88:1.24), pterostigma 3.43× longer than wide (0.48: 0.14); marginal cell 3.76× longer than wide (0.94:0.25), slightly open at apex, Rs slightly sinuate toward apex; right angle between Rs and Rs + M (ca. 90°); first abscissa of M ca.
|
|||
|
|
<emphasis id="B91DF041FFD9FF81AE7F21B4F779FAEF" bold="true" box="[814,834,1267,1291]" pageId="11">⅔</emphasis>
|
|||
|
|
length of first abscissa of Cu (0.42: 0.62) and slightly shorter than Rs + M + second M (0.46). First submarginal cell with limits relatively clear; second submarginal cell completely absent. Hamuli, 5. Tibia III subtriangular (as
|
|||
|
|
<taxonomicName id="4C6957D0FFD9FF81AEB020C0F0A8FA7A" box="[993,1171,1415,1438]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="species" species="dominicana">
|
|||
|
|
<emphasis id="B91DF041FFD9FF81AEB020C0F0A8FA7A" box="[993,1171,1415,1438]" italics="true" pageId="11">P. dominicana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C6957D0FFD9FF81A98F20C0F10CFA7A" authorityName="Schwarz" authorityYear="1938" box="[1246,1335,1415,1438]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|||
|
|
<emphasis id="B91DF041FFD9FF81A98F20C0F10CFA7A" box="[1246,1335,1415,1438]" italics="true" pageId="11">Plebeia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicNameLabel id="A22E4D3AFFD9FF81AE7F20E3F769FA5F" box="[814,850,1444,1467]" pageId="11" sensu="stricto">s.s.</taxonomicNameLabel>
|
|||
|
|
), with posterodistal corner approximately right angled (position of tibiae, and presence of residues and bubbles do not permit reliable measures and details of shape); depressed rim on posterior border of inner surface
|
|||
|
|
<superScript id="7C1C811BFFD9FF81AE382372F74BF9A7" attach="right" box="[873,880,1589,1603]" fontSize="6" pageId="11">2</superScript>
|
|||
|
|
<emphasis id="B91DF041FFD9FF81AE212370F74FF9AB" bold="true" box="[880,884,1591,1615]" pageId="11">⁄</emphasis>
|
|||
|
|
<subScript id="17ED2E16FFD9FF81AE222304F747F9B5" attach="left" box="[883,892,1603,1617]" fontSize="6" pageId="11">5</subScript>
|
|||
|
|
width of elevated keirotrichiate area. Basitarsus III 1.93× longer than wide (0.56: 0.29), slightly broadened toward apex, posterodistal corner right angled.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3737FD8FFD9FF86AE1D23EBF69AFEC1" lastPageId="12" pageId="11" type="materials_examined">
|
|||
|
|
<paragraph id="8BD62C53FFD9FF86AE1D23EBF69AFEC1" blockId="11.[814,1343,182,1731]" lastBlockId="12.[272,801,182,1408]" lastPageId="12" pageId="11">
|
|||
|
|
<materialsCitation id="3B01260EFFD9FF86AE1D23EBF626FF29" collectionCode="AMNH-DR" country="Dominican Republic" lastPageId="12" location="Miocene" pageId="11" specimenCode="AMNH-DR-14-1481" specimenCount="1" typeStatus="holotype">
|
|||
|
|
TYPES:
|
|||
|
|
<typeStatus id="54D292F1FFD9FF81AEEB23EBF00AF927" box="[954,1073,1708,1731]" pageId="11" type="holotype">Holotype</typeStatus>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D6FE7DAFFD9FF81A90023EBF091F927" box="[1105,1194,1708,1731]" pageId="11" type="worker">worker</specimenCount>
|
|||
|
|
, specimen
|
|||
|
|
<specimenCode id="DBCF8428FFDEFF86AC4125F1F623FF29" box="[272,536,182,205]" pageId="12">AMNHDR141481</specimenCode>
|
|||
|
|
,
|
|||
|
|
</materialsCitation>
|
|||
|
|
<materialsCitation id="3B01260EFFDEFF86AF7C25F1F69AFEC1" collectionCode="AMNH-DR" country="Dominican Republic" location="Miocene" pageId="12" specimenCode="AMNH-DR-14-854, DR-14-1440" specimenCount="1" typeStatus="paratype">
|
|||
|
|
<typeStatus id="54D292F1FFDEFF86AF7C25F1F692FF29" box="[557,681,182,205]" pageId="12" type="paratype">paratypes</typeStatus>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D6FE7DAFFDEFF86AFEC25F1F71AFF29" box="[701,801,182,205]" pageId="12" type="worker">workers</specimenCount>
|
|||
|
|
<specimenCode id="DBCF8428FFDEFF86AC412593F5CCFF0F" box="[272,503,212,235]" pageId="12">AMNHDR14854</specimenCode>
|
|||
|
|
and
|
|||
|
|
<specimenCode id="DBCF8428FFDEFF86AF632593F6FCFF0F" box="[562,711,212,235]" pageId="12">DR141440</specimenCode>
|
|||
|
|
, in amber from
|
|||
|
|
<location id="8EB67A88FFDEFF86ACD825B6F5D4FEEC" LSID="urn:lsid:plazi:treatment:03C09D45FFD8FF86AC0B2036F603FA64:8EB67A88FFDEFF86ACD825B6F5D4FEEC" box="[393,495,241,264]" country="Dominican Republic" municipality="Specific" name="Miocene" pageId="12">Miocene</location>
|
|||
|
|
of
|
|||
|
|
<collectingCountry id="F37E6CC3FFDEFF86AF7325B6F726FEEC" box="[546,797,241,264]" name="Dominican Republic" pageId="12">Dominican Republic</collectingCountry>
|
|||
|
|
. Specific mine of origin unknown.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3737FD8FFDEFF86AC7C246BF557FE84" pageId="12" type="etymology">
|
|||
|
|
<paragraph id="8BD62C53FFDEFF86AC7C246BF557FE84" blockId="12.[272,801,182,1408]" pageId="12">
|
|||
|
|
ETYMOLOGY: From the Latin,
|
|||
|
|
<emphasis id="B91DF041FFDEFF86AFD9246BF6DDFEA7" box="[648,742,300,323]" italics="true" pageId="12">vetustus</emphasis>
|
|||
|
|
, old, ancient.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3737FD8FFDEFF86AC7C2421F603FA64" pageId="12" type="discussion">
|
|||
|
|
<paragraph id="8BD62C53FFDEFF86AC7C2421F726FC94" blockId="12.[272,801,182,1408]" pageId="12">
|
|||
|
|
REMARKS:
|
|||
|
|
<bibRefCitation id="EFF851A2FFDEFF86ACE12421F66AFE99" author="Poinar, G. O., Jr." box="[432,593,358,381]" pageId="12" refId="ref11819" refString="1994" type="journal article" year="1994">Poinar (1994</bibRefCitation>
|
|||
|
|
b: 73, fig. 2) referred to specimen AMNHDR141440 (here designated as a
|
|||
|
|
<typeStatus id="54D292F1FFDEFF86AF5A24E6F64AFE5C" box="[523,625,417,440]" pageId="12" type="paratype">paratype</typeStatus>
|
|||
|
|
) as
|
|||
|
|
<taxonomicName id="4C6957D0FFDEFF86AFCE24E6F71BFE5C" box="[671,800,417,440]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="B91DF041FFDEFF86AFCE24E6F6C3FE5C" box="[671,760,417,440]" italics="true" pageId="12">Plebeia</emphasis>
|
|||
|
|
sp.
|
|||
|
|
</taxonomicName>
|
|||
|
|
<bibRefCitation id="EFF851A2FFDEFF86AC4124F9F69FFE31" author="Michener, C. D. & G. O. Poinar, Jr." box="[272,676,446,469]" journalOrPublisher="J. Kansas Entomol. Soc." pageId="12" pagination="353 - 361" part="69" refId="ref11664" refString="Michener, C. D., and G. O. Poinar, Jr. 1996. The known bee fauna of the Dominican Amber. J. Kansas Entomol. Soc. 69 (4) suppl.: 353 - 361." title="The known bee fauna of the Dominican Amber." type="journal article" year="1996">Michener and Poinar (1996: 354)</bibRefCitation>
|
|||
|
|
examined this same specimen and considered it a species distinct from
|
|||
|
|
<taxonomicName id="4C6957D0FFDEFF86ACA724BEF69CFDF4" box="[502,679,505,528]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="species" species="dominicana">
|
|||
|
|
<emphasis id="B91DF041FFDEFF86ACA724BEF69CFDF4" box="[502,679,505,528]" italics="true" pageId="12">P. dominicana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, however these authors disagreed about its placement in the genus
|
|||
|
|
<taxonomicName id="4C6957D0FFDEFF86ACFF2773F63CFDAF" authorityName="Schwarz" authorityYear="1938" box="[430,519,564,587]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B91DF041FFDEFF86ACFF2773F63CFDAF" box="[430,519,564,587]" italics="true" pageId="12">Plebeia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C6957D0FFDEFF86AF482773F738FDAF" box="[537,771,564,587]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="species" species="vestusta">
|
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<emphasis id="B91DF041FFDEFF86AF482773F738FDAF" box="[537,771,564,587]" italics="true" pageId="12">Proplebeia vestusta</emphasis>
|
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|
</taxonomicName>
|
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|
|
is placed in the genus tentatively, based on the form of the forewing medial cell, the surrounding veins, integument structure, and general body form (which is as robust as in
|
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|
|
<taxonomicName id="4C6957D0FFDEFF86AC412780F581FD3A" box="[272,442,711,734]" class="Insecta" family="Apidae" genus="Proplebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="species" species="dominicana">
|
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|
|
<emphasis id="B91DF041FFDEFF86AC412780F581FD3A" box="[272,442,711,734]" italics="true" pageId="12">P. dominicana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<emphasis id="B91DF041FFDEFF86ACA22780F66FFD3A" box="[499,596,711,734]" italics="true" pageId="12">
|
|||
|
|
<taxonomicName id="4C6957D0FFDEFF86ACA22780F674FD3A" authorityName="Schwarz" authorityYear="1938" box="[499,591,711,734]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="genus" sensu="stricto">Plebeia</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicNameLabel id="A22E4D3AFFDEFF86AF0A2781F644FD39" box="[603,639,710,733]" pageId="12" sensu="stricto">s.s.</taxonomicNameLabel>
|
|||
|
|
) and, mainly, by the depression of the posterior rim of the inner surface of tibia III, with ca.
|
|||
|
|
<superScript id="7C1C811BFFDEFF86AFC82647F69BFCEA" attach="right" box="[665,672,768,782]" fontSize="6" pageId="12">2</superScript>
|
|||
|
|
<emphasis id="B91DF041FFDEFF86AFF12645F69FFCFE" bold="true" box="[672,676,770,794]" pageId="12">⁄</emphasis>
|
|||
|
|
<subScript id="17ED2E16FFDEFF86AFF5264AF696FCFF" attach="left" box="[676,685,781,795]" fontSize="6" pageId="12">5</subScript>
|
|||
|
|
the width of the keirotrichiate area. The rastellum and penicillum apparently do not differ from what is seen in the species of
|
|||
|
|
<taxonomicName id="4C6957D0FFDEFF86AF2A261EF6EFFC94" authorityName="Schwarz" authorityYear="1938" box="[635,724,857,880]" class="Insecta" family="Apidae" genus="Plebeia" kingdom="Animalia" order="Hymenoptera" pageId="12" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B91DF041FFDEFF86AF2A261EF6EFFC94" box="[635,724,857,880]" italics="true" pageId="12">Plebeia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(s.s.).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD62C53FFDEFF86AC7C2631F603FA64" blockId="12.[272,801,182,1408]" pageId="12">
|
|||
|
|
In the
|
|||
|
|
<typeStatus id="54D292F1FFDEFF86ACDF2631F5C9FC69" box="[398,498,886,909]" pageId="12" type="paratype">paratype</typeStatus>
|
|||
|
|
AMNHDR14854 the clypeus has only a pale yellow stripe along the distal border, a little narrower than the diameter of the scape, and the supraclypeal area is also pale yellow, but the parocular areas are black. In the
|
|||
|
|
<typeStatus id="54D292F1FFDEFF86AF4E214EF6B8FBC4" box="[543,643,1033,1056]" pageId="12" type="paratype">paratype</typeStatus>
|
|||
|
|
AMNHDR 141440 only one premarginal, fine, yellow stripe is visible in the clypeus; the other areas are blackish. On both
|
|||
|
|
<typeStatus id="54D292F1FFDEFF86AF4B2126F6B2FB9C" box="[538,649,1121,1144]" pageId="12" type="paratype">paratypes</typeStatus>
|
|||
|
|
(extensively surrounded by air bubbles and the legs tangled in such a way that is impossible to examine important characters), the vertex is arched and well elevated above the lateral ocelli, but this condition can be due to the lenticular deformation of the amber. In the remaining characters (e.g., size and proportions of structures), these specimens do not differ from the
|
|||
|
|
<typeStatus id="54D292F1FFDEFF86AC9B202EF60FFA64" box="[458,564,1385,1408]" pageId="12" type="holotype">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|