treatments-xml/data/35/1A/AE/351AAE4D024CFFF93295B882FAAAFEA4.xml

<document id="4E33D5457D5F9084AEA329B7EBAD952D" ID-CLB-Dataset="7387" ID-DOI="10.11646/zootaxa.4951.3.3" ID-GBIF-Dataset="b4b99917-e609-4b32-aba7-62bbddd1f0cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668172" ID-ZooBank="ADA22E04-9548-4ECC-AF84-18E51955852B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617778947944" checkinUser="plazi" docAuthor="Souma, Jun &amp; Ishikawa, Tadashi" docDate="2021" docId="351AAE4D024CFFF93295B882FAAAFEA4" docLanguage="en" docName="zootaxa.4951.3.3.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Physopelta (Neophysopelta) quadriguttata Bergroth 1894" docType="treatment" docVersion="9" lastPageNumber="486" masterDocId="C923D635025BFFE03202B97CFF80FFB9" masterDocTitle="A taxonomic review of the genus Physopelta (Hemiptera: Heteroptera: Largidae) from Japan, Korea, and Taiwan, with a new record of Ph. parviceps from China" masterLastPageNumber="491" masterPageNumber="461" pageNumber="484" updateTime="1698936105298" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E715EBDC69437555E91C2F1700F7A356" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="329EE0AB5E0CE8DCE9872CFA127FAE0F">
<mods:title id="681D8CA90FB8857DF505F6A8346C6ACF">A taxonomic review of the genus Physopelta (Hemiptera: Heteroptera: Largidae) from Japan, Korea, and Taiwan, with a new record of Ph. parviceps from China</mods:title>
</mods:titleInfo>
<mods:name id="0375995DF5AF911C0757B9D3BECA3250" type="personal">
<mods:role id="C455B978FC84A3DDDA123CA70A7C61C3">
<mods:roleTerm id="F1DA53849D52A86D5657D644579E3C22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53AFBBE81BBCC9FEA8C691065626CECD">Souma, Jun</mods:namePart>
<mods:nameIdentifier id="A8E7C1B3F4FF9448BA8CB02F66A1D0E0" type="ORCID">0000-0002-2238-5015</mods:nameIdentifier>
<mods:affiliation id="642F8005BB9B020685C1A4E7AE673102">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan. kodokusignal @ gmail. com; https: // orcid. org / 0000 - 0002 - 2238 - 5015 &amp; Research Fellowship for Young Scientists (DC 1), Japan Society for the Promotion of Science, Tokyo, Japan</mods:affiliation>
<mods:nameIdentifier id="1C101C0DB47E76E7967658A887A4098F" type="email">kodokusignal@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="6D5255D98BC1A9E60D20957CAAFDFF58" type="personal">
<mods:role id="04CB988C605A556DA24B3EB1ADD1A36B">
<mods:roleTerm id="252384A6374F1ACD7EC6FEDB153B4C8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4458C56FA05166DD369B0A584B6A016C">Ishikawa, Tadashi</mods:namePart>
<mods:affiliation id="4C0860C6F6D44DDAD8A197A539F37897">Laboratory of Entomology, Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9ED096C9340044CCCF6AB935291FE7DB">text</mods:typeOfResource>
<mods:relatedItem id="4445D52DB70D571543DB569EC9755892" type="host">
<mods:titleInfo id="845A3833C2F2F6E2CCBF1D50F9F07AE4">
<mods:title id="CA2E918DB177CFAD7A096323F2A07E38">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DD9CF3A805021210B5AC1F6EF8AF492C">
<mods:date id="620F4E2593C0887807405184A60887DD">2021</mods:date>
<mods:detail id="105DBE7F0006649B82C065B61C0E6B6C" type="pubDate">
<mods:number id="50C8D62792FA1DB58E2F6C432E2FA1E7">2021-04-07</mods:number>
</mods:detail>
<mods:detail id="B1573D8957DFE7DEB4F43FCBAE3F0216" type="volume">
<mods:number id="3421C9D107B1451873200B3989D6A2D5">4951</mods:number>
</mods:detail>
<mods:detail id="984A515AEDCCF0D53E0CF939553499F2" type="issue">
<mods:number id="8FCA49216FD1EF1692F1399C355A6ABF">3</mods:number>
</mods:detail>
<mods:extent id="4B327BAC4C1CD86D7F69F356D86040C8" unit="page">
<mods:start id="E414F82FF35C9474EB032BEA502E6F64">461</mods:start>
<mods:end id="F5BEAEEF9B339494E44F27773122530E">491</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="23620E29C8F7491C571846B54B609DB8">journal article</mods:classification>
<mods:identifier id="61F458F61CDEFB12DCA3F297B2B04B93" type="CLB-Dataset">7387</mods:identifier>
<mods:identifier id="9B415FF8174B6D37CA2FB75E8E3EF9F1" type="DOI">10.11646/zootaxa.4951.3.3</mods:identifier>
<mods:identifier id="A70DB9CD6AFCA7233D184B5A56A41E38" type="GBIF-Dataset">b4b99917-e609-4b32-aba7-62bbddd1f0cf</mods:identifier>
<mods:identifier id="A6449786303E1587A746DFA3BE01A720" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8383D7B0F683BAD649D4C3777B15E6A1" type="Zenodo-Dep">4668172</mods:identifier>
<mods:identifier id="BB8808A542F3840A1F1B152095973F4A" type="ZooBank">ADA22E04-9548-4ECC-AF84-18E51955852B</mods:identifier>
</mods:mods>
<treatment id="351AAE4D024CFFF93295B882FAAAFEA4" ID-DOI="http://doi.org/10.5281/zenodo.4668185" ID-GBIF-Taxon="181003287" ID-Zenodo-Dep="4668185" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4" httpUri="http://treatment.plazi.org/id/351AAE4D024CFFF93295B882FAAAFEA4" lastPageId="25" lastPageNumber="486" pageId="23" pageNumber="484">
<subSubSection id="F5A94CD0024CFFF73295B882FCD6FDA1" box="[151,854,510,537]" pageId="23" pageNumber="484" type="nomenclature">
<paragraph id="BD0C1F5B024CFFF73295B882FCD6FDA1" blockId="23.[151,884,510,573]" box="[151,854,510,537]" pageId="23" pageNumber="484">
<heading id="E644A837024CFFF73295B882FCD6FDA1" bold="true" box="[151,854,510,537]" fontSize="11" level="1" pageId="23" pageNumber="484" reason="1">
<taxonomicName id="7AB364D8024CFFF73295B882FCD6FDA1" ID-CoL="7WG5G" authority="Bergroth, 1894" authorityName="Bergroth" authorityYear="1894" box="[151,854,510,537]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="484" phylum="Arthropoda" rank="species" species="quadriguttata" subGenus="Neophysopelta">
<emphasis id="8FC7C349024CFFF73295B882FCD6FDA1" bold="true" box="[151,854,510,537]" pageId="23" pageNumber="484">
<emphasis id="8FC7C349024CFFF73295B882FE9BFDA0" bold="true" box="[151,283,510,537]" italics="true" pageId="23" pageNumber="484">Physopelta</emphasis>
(
<emphasis id="8FC7C349024CFFF7332EB882FE5EFDA0" bold="true" box="[300,478,510,537]" italics="true" pageId="23" pageNumber="484">Neophysopelta</emphasis>
) 
<emphasis id="8FC7C349024CFFF733EDB882FD14FDA0" bold="true" box="[495,660,510,537]" italics="true" pageId="23" pageNumber="484">quadriguttata</emphasis>
<bibRefCitation id="D92262AA024CFFF73099B883FCD6FDA1" author="Bergroth, E." box="[667,854,510,537]" pageId="23" pageNumber="484" pagination="152 - 164" refId="ref21667" refString="Bergroth, E. (1894) Rhynchota orientalia. Revue d'Entomologie, 13, 152 - 164. https: // doi. org / 10.5962 / bhl. part. 13239" type="journal article" year="1894">Bergroth, 1894</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD0024CFFF7329CBB5FFCF4FD84" box="[158,884,546,573]" pageId="23" pageNumber="484" type="description">
<paragraph id="BD0C1F5B024CFFF7329CBB5FFCF4FD84" blockId="23.[151,884,510,573]" box="[158,884,546,573]" pageId="23" pageNumber="484">
(
<figureCitation id="258803DE024CFFF732A4BB5FFEA9FD84" box="[166,297,547,573]" captionStart="FIGURE 1" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[154,1430,425,1465]" captionTargetId="figure-130@3.[151,1436,417,1468]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of four Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta, male (A) and female (B) from Japan proper; Ph. (N.) fusciscutellata sp. nov., male from the main island of Taiwan (C) and female from the Ryukyu Islands, Japan (D); Ph. (N.) quadriguttata, male (E, G) and female (F) from the main island of Taiwan; Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (H). Arrows mark antenna with antennomeral oligomery and malformed calli. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668176" httpUri="https://zenodo.org/record/4668176/files/figure.png" pageId="23" pageNumber="484">Figs. 1E–G</figureCitation>
, 
<figureCitation id="258803DE024CFFF73337BB5FFED3FD84" box="[309,339,547,573]" captionStart="FIGURE 3" captionStartId="10.[151,250,1897,1921]" captionTargetBox="[415,1165,191,1860]" captionTargetId="figure-30@10.[392,1196,181,1874]" captionTargetPageId="10" captionText="FIGURE 3. Habitus images of five Physopelta (Neophysopelta) species, lateral view: Ph. (N.) gutta gutta, female from Japan proper (A); Ph. (N.) fusciscutellata sp. nov., female from the Ryukyu Islands, Japan (B); Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (C); Ph. (N.) parviceps, male from Japan proper (D) and female from the Ryukyu Islands (E); Ph. (N.) quadriguttata, male from the main island of Taiwan (F). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668183" httpUri="https://zenodo.org/record/4668183/files/figure.png" pageId="23" pageNumber="484">3F</figureCitation>
, 
<figureCitation id="258803DE024CFFF7335CBB5FFEFCFD84" box="[350,380,547,573]" captionStart="FIGURE 4" captionStartId="11.[151,250,1800,1824]" captionTargetBox="[162,1418,193,1763]" captionTargetId="figure-17@11.[155,1432,181,1776]" captionTargetPageId="11" captionText="FIGURE 4. Habitus images of five Physopelta (Neophysopelta) species, ventral view: Ph. (N.) gutta gutta, female from Japan proper (A); Ph. (N.) fusciscutellata sp. nov., female from the Ryukyu Islands, Japan (B); Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (C); Ph. (N.) parviceps, male from Japan proper (D) and female from the Ryukyu Islands (E); Ph. (N.) quadriguttata, male from the main island of Taiwan (F). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668187" httpUri="https://zenodo.org/record/4668187/files/figure.png" pageId="23" pageNumber="484">4F</figureCitation>
, 
<figureCitation id="258803DE024CFFF7338ABB5EFE2CFD84" box="[392,428,546,573]" captionStart="FIGURE 5" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[154,1437,1028,1838]" captionTargetId="figure-420@12.[151,1437,1028,1838]" captionTargetPageId="12" captionText="FIGURE 5. Male pronotum and scutellum of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); macropterous morph of Ph. (N.) parviceps from Japan proper (C, D); macropterous morph of Ph. (N.) parviceps from the Ryukyu Islands, Japan (E); brachypterous morph of Ph. (N.) parviceps from Japan proper (F); Ph. (N.) fusciscutellata sp. nov. (G); Ph. (N.) quadriguttata (H). Arrows mark malformed calli. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668189" httpUri="https://zenodo.org/record/4668189/files/figure.png" pageId="23" pageNumber="484">5H</figureCitation>
, 
<figureCitation id="258803DE024CFFF733BABB5FFE56FD84" box="[440,470,547,573]" captionStart="FIGURE 6" captionStartId="13.[151,250,1708,1732]" captionTargetBox="[162,1433,890,1675]" captionTargetId="figure-351@13.[151,1437,885,1685]" captionTargetPageId="13" captionText="FIGURE 6. Left fore wing of five macropterous species of Physopelta (Neophysopelta), dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668191" httpUri="https://zenodo.org/record/4668191/files/figure.png" pageId="23" pageNumber="484">6F</figureCitation>
, 
<figureCitation id="258803DE024CFFF733E3BB5FFD80FD84" box="[481,512,547,573]" captionStart="FIGURE 7" captionStartId="16.[151,250,1458,1482]" captionTargetBox="[151,1436,741,1435]" captionTargetId="figure-318@16.[151,1436,741,1435]" captionTargetPageId="16" captionText="FIGURE 7. Procoxa of five Physopelta (Neophysopelta) species, ventral view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviation: pp = procoxal projection. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4668193" httpUri="https://zenodo.org/record/4668193/files/figure.png" pageId="23" pageNumber="484">7F</figureCitation>
, 
<figureCitation id="258803DE024CFFF73009BB5FFDA8FD84" box="[523,552,547,573]" captionStart="FIGURE 8" captionStartId="17.[151,250,1278,1302]" captionTargetBox="[151,1436,561,1255]" captionTargetId="figure-215@17.[151,1436,561,1255]" captionTargetPageId="17" captionText="FIGURE 8. Left scent gland ostiole of five Physopelta (Neophysopelta) species, ventral view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviation: pe = peritreme. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668195" httpUri="https://zenodo.org/record/4668195/files/figure.png" pageId="23" pageNumber="484">8F</figureCitation>
, 
<figureCitation id="258803DE024CFFF73036BB5FFDD2FD84" box="[564,594,547,573]" captionStart="FIGURE 9" captionStartId="18.[151,250,899,923]" captionTargetBox="[151,1437,181,876]" captionTargetId="figure-30@18.[151,1437,181,876]" captionTargetPageId="18" captionText="FIGURE 9. Genital capsule of five Physopelta (Neophysopelta) species, posterior view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: vif = infolding of ventral rim; cr = crown of paramere. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668197" httpUri="https://zenodo.org/record/4668197/files/figure.png" pageId="23" pageNumber="484">9F</figureCitation>
, 
<figureCitation id="258803DE024CFFF7305FBB5FFD09FD84" box="[605,649,547,573]" captionStart="FIGURE 10" captionStartId="18.[151,250,1747,1771]" captionTargetBox="[156,1433,1060,1719]" captionTargetId="figure-119@18.[151,1436,1056,1724]" captionTargetPageId="18" captionText="FIGURE 10. Left scent gland ostiole of five Physopelta (Neophysopelta) species, lateral view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviation: pe = peritreme. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4668199" httpUri="https://zenodo.org/record/4668199/files/figure.png" pageId="23" pageNumber="484">10F</figureCitation>
, 
<figureCitation id="258803DE024CFFF73097BB5FFD3FFD84" box="[661,703,547,573]" captionStart="FIGURE 11" captionStartId="19.[151,250,1814,1838]" captionTargetBox="[247,1355,187,1784]" captionTargetId="figure-17@19.[224,1364,181,1790]" captionTargetPageId="19" captionText="FIGURE 11. Phallus of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: ad = aedeagus; bb = basal plate bridge; bp = basal plate; co = conjunctiva; cp = capitate process; dc = dorsal connective; ds = ductus seminis; ed = endophallic duct; eo = endophallic reservoir outgrowth; ep = erection fluid pump; er = endophallic reservoir; pt = phallotheca; 1ca, 2ca = first and second conjunctival process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668201" httpUri="https://zenodo.org/record/4668201/files/figure.png" pageId="23" pageNumber="484">11F</figureCitation>
, 
<figureCitation id="258803DE024CFFF730C9BB5FFD77FD84" box="[715,759,547,573]" captionStart="FIGURE 12" captionStartId="20.[151,250,1903,1927]" captionTargetBox="[160,1409,187,1865]" captionTargetId="figure-30@20.[151,1436,181,1880]" captionTargetPageId="20" captionText="FIGURE 12. Left paramere of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: ba = basal apodeme; cr = crown; ne = neck; st = stem. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668203" httpUri="https://zenodo.org/record/4668203/files/figure.png" pageId="23" pageNumber="484">12F</figureCitation>
, 
<figureCitation id="258803DE024CFFF73100BB5FFCB2FD84" box="[770,818,547,573]" captionStart="FIGURE 13" captionStartId="21.[151,250,1679,1703]" captionTargetBox="[164,1421,468,1646]" captionTargetId="figure-131@21.[151,1436,453,1656]" captionTargetPageId="21" captionText="FIGURE 13. Female terminalia of four Physopelta (Neophysopelta) species, anterior view: Ph. (N.) gutta gutta (A); Ph. (N.) parviceps from Japan proper (B); Ph. (N.) parviceps from the Ryukyu Islands, Japan (C); Ph. (N.) fusciscutellata sp. nov. (D); Ph. (N.) quadriguttata (E). Abbreviations: go = gonangulum; lt9 = laterotergite IX; mt8 = mediotergite VIII; ra8, ra9 = ramus of valvula XIII, IX; rs = ring sclerite; va8, va9 = valvulae VIII, IX; vf8, vf9 = valvifers VIII, IX. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668205" httpUri="https://zenodo.org/record/4668205/files/figure.png" pageId="23" pageNumber="484">13E</figureCitation>
, 
<figureCitation id="258803DE024CFFF7313FBB5FFCECFD84" box="[829,876,547,573]" captionStart="FIGURE 14" captionStartId="22.[151,250,1605,1629]" captionTargetBox="[166,1429,458,1575]" captionTargetId="figure-160@22.[151,1436,453,1581]" captionTargetPageId="22" captionText="FIGURE 14. Spermatheca of four Physopelta (Neophysopelta) species: Ph. (N.) gutta gutta (A); Ph. (N.) parviceps from Japan proper (B); Ph. (N.) parviceps from the Ryukyu Islands, Japan (C); Ph. (N.) fusciscutellata sp. nov. (D); Ph. (N.) quadriguttata (E). Abbreviations: ar = apical receptacle; ip = intermediate part; sd = spermathecal duct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668207" httpUri="https://zenodo.org/record/4668207/files/figure.png" pageId="23" pageNumber="484">14E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD0024CFFF73295BB1BFE1DFC8E" pageId="23" pageNumber="484" type="reference_group">
<paragraph id="BD0C1F5B024CFFF73295BB1BFAE2FDC6" blockId="23.[151,1378,615,639]" box="[151,1378,615,639]" pageId="23" pageNumber="484">
<treatmentCitationGroup id="9DA33875024CFFF73295BB1BFAE2FDC6" box="[151,1378,615,639]" pageId="23" pageNumber="484">
<treatmentCitation id="3C12394A024CFFF73295BB1BFDEFFDC6" author="Bergroth, E." box="[151,623,615,639]" page="160" pageId="23" pageNumber="484" year="1894">
<taxonomicName id="7AB364D8024CFFF73295BB1BFDEFFDC6" ID-CoL="7WG5G" authority="Bergroth, 1894: 160" authorityName="Bergroth" authorityPageNumber="160" authorityYear="1894" box="[151,623,615,639]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="484" phylum="Arthropoda" rank="species" species="quadriguttata">
<emphasis id="8FC7C349024CFFF73295BB1BFE19FDC6" box="[151,409,615,639]" italics="true" pageId="23" pageNumber="484">Physopelta quadriguttata</emphasis>
<bibRefCitation id="D92262AA024CFFF733A2BB1BFDEFFDC6" author="Bergroth, E." box="[416,623,615,639]" pageId="23" pageNumber="484" pagination="152 - 164" refId="ref21667" refString="Bergroth, E. (1894) Rhynchota orientalia. Revue d'Entomologie, 13, 152 - 164. https: // doi. org / 10.5962 / bhl. part. 13239" type="journal article" year="1894">Bergroth, 1894: 160</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="6208A1F9024CFFF7307BBB1BFD5AFDC6" box="[633,730,615,639]" pageId="23" pageNumber="484" type="syntype">Syntypes</typeStatus>
: macropterous 
<specimenCount id="ABB5D4D2024CFFF73171BB1BFC19FDC6" box="[883,921,615,639]" pageId="23" pageNumber="484" type="female">2 ♀</specimenCount>
, 
<collectingCountry id="C5A45FCB024CFFF731A4BB1BFC5EFDC6" box="[934,990,615,639]" name="India" pageId="23" pageNumber="484">India</collectingCountry>
: 
<collectingRegion id="7F77D1B9024CFFF731E5BB1BFBB4FDC6" box="[999,1076,615,639]" country="India" name="Sikkim" pageId="23" pageNumber="484">Sikkim</collectingRegion>
; MZHF? (
<treatmentCitation id="3C12394A024CFFF7369FBB1BFAD6FDC6" ID-DOI="http://doi.org/10.5281/zenodo.5906250" ID-Zenodo-Dep="5906250" author="Stehlik, J. L." box="[1181,1366,615,639]" httpUri="http://treatment.plazi.org/id/03B66912FF950441FE93FF1CFBD8FEAE" page="540" pageId="23" pageNumber="484" year="2013">
<bibRefCitation id="D92262AA024CFFF7369FBB1BFAD6FDC6" author="Stehlik, J. L." box="[1181,1366,615,639]" pageId="23" pageNumber="484" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík 2013: 540</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BD0C1F5B024CFFF73295BBD8FE1DFC8E" blockId="23.[151,1437,676,823]" pageId="23" pageNumber="484">
<emphasis id="8FC7C349024CFFF73295BBD8FE91FD04" bold="true" box="[151,273,676,701]" inLineHeading="true" pageId="23" pageNumber="484" reason="1">
<emphasis id="8FC7C349024CFFF73295BBD8FE8AFD05" bold="true" box="[151,266,676,700]" italics="true" pageId="23" pageNumber="484">References</emphasis>
.
</emphasis>
<bibRefCitation id="D92262AA024CFFF7331BBBD8FE52FD05" author="Esaki, T." box="[281,466,676,700]" pageId="23" pageNumber="484" pagination="136 - 189" refId="ref21862" refString="Esaki, T. (1926) Verzeichniss der Hemiptera-heteroptera der Insel Formosa. Annales Musei Nationalis Hungarici, 24, 136 - 189." type="journal article" year="1926">Esaki (1926: 157)</bibRefCitation>
(distribution); 
<bibRefCitation id="D92262AA024CFFF7306DBBD8FCA7FD04" author="Blote, H. C." box="[623,807,676,701]" pageId="23" pageNumber="484" pagination="97 - 117" refId="ref21700" refString="Blote, H. C. (1931) Catalogue of the Pyrrhocoridae in ' s Rijks Museum van Natuurlijke Historie. Zoologische Mededeelingen, Leiden, 14, 97 - 117." type="journal article" year="1931">Blöte (1931: 100)</bibRefCitation>
(distribution); 
<bibRefCitation id="D92262AA024CFFF731C6BBD8FB24FD05" author="Kerzhner, I. M." box="[964,1188,676,701]" pageId="23" pageNumber="484" pagination="1843" refId="ref22916" refString="Kerzhner, I. M. (2001) Superfamily Pyrrhocoroidea Amyot &amp; Serville, 1843. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. The Netherlands Entomological Society, Amster- dam, pp. 245 - 247." type="book chapter" year="2001">Kerzhner (2001: 246)</bibRefCitation>
(checklist: Palaearctic); 
<bibRefCitation id="D92262AA024CFFF732C5BBBFFEF4FD62" author="Stehlik, J. L." box="[199,372,707,731]" pageId="23" pageNumber="484" pagination="1 - 11" refId="ref24393" refString="Stehlik, J. L. (2004) Largidae and Pyrrhocoridae of Nepal (Heteroptera). Entomologica Basiliensia, 25, 1 - 11." type="journal article" year="2004">Stehlík (2004: 3)</bibRefCitation>
(distribution); 
<bibRefCitation id="D92262AA024CFFF7300DBBBFFCA3FD62" author="Miyamoto, S. &amp; Hayashi, M. &amp; Kohno, K." box="[527,803,707,731]" pageId="23" pageNumber="484" pagination="11 - 13" refId="ref23710" refString="Miyamoto, S., Hayashi, M. &amp; Kohno, K. (2005) New records of three Pentatomomorphan species (Heteroptera) from Japan. Japanese Journal of Systematic Entomology, 11, 11 - 13." type="journal article" year="2005">
Miyamoto 
<emphasis id="8FC7C349024CFFF7307DBBB8FD33FD62" box="[639,691,707,731]" italics="true" pageId="23" pageNumber="484">et al.</emphasis>
(2005: 11)
</bibRefCitation>
(distribution); 
<bibRefCitation id="D92262AA024CFFF731BCBBBFFB25FD62" author="Redei, D. &amp; Tsai, J. - F. &amp; Yang, M. - M." box="[958,1189,707,731]" pageId="23" pageNumber="484" refId="ref24020" refString="Redei, D., Tsai, J. - F. &amp; Yang, M. - M. (2009) Heteropteran Fauna of Taiwan: Cotton stainers and relatives (Hemiptera: Heteroptera: Pyrrhocoroidea). National Chung Hsing University, Taichung, 52 pp." type="book" year="2009">
Rédei 
<emphasis id="8FC7C349024CFFF73602BBB8FBB4FD62" box="[1024,1076,707,731]" italics="true" pageId="23" pageNumber="484">et al.</emphasis>
(2009: 13)
</bibRefCitation>
(monograph); 
<bibRefCitation id="D92262AA024CFFF7373CBBBFFEDFFD43" author="Kohno, K. &amp; Hayashi, M. &amp; Miyamoto, S." pageId="23" pageNumber="484" pagination="90 - 91" refId="ref22979" refString="Kohno, K., Hayashi, M. &amp; Miyamoto, S. (2012) Family Largidae Amyot &amp; Serville, 1843. In: Ishikawa, T., Takai, M. &amp; Yasunaga, T. (Eds.), A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, pp. 399 - 402, pls. 90 - 91. [in Japanese]" type="book chapter" year="2012">
Kohno 
<emphasis id="8FC7C349024CFFF73788BBB8FF62FD43" italics="true" pageId="23" pageNumber="484">et al.</emphasis>
(2012: 400)
</bibRefCitation>
(monograph); 
<bibRefCitation id="D92262AA024CFFF733F4BB9EFC82FD43" author="Aukema, B. &amp; Rieger, C. &amp; Rabitsch, W." box="[502,770,738,762]" pageId="23" pageNumber="484" refId="ref21623" refString="Aukema, B., Rieger, C. &amp; Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6, Supplement. The Netherlands Entomological Society, Amsterdam, xxiv + 629 pp." type="book" year="2013">
Aukema 
<emphasis id="8FC7C349024CFFF73050BB9FFDFFFD43" box="[594,639,738,762]" italics="true" pageId="23" pageNumber="484">et al</emphasis>
. (2013: 401)
</bibRefCitation>
(checklist: Palaearctic); 
<bibRefCitation id="D92262AA024CFFF731FCBB9EFB43FD43" author="Stehlik, J. L." box="[1022,1219,738,762]" pageId="23" pageNumber="484" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík (2013: 540)</bibRefCitation>
(monograph); 
<bibRefCitation id="D92262AA024CFFF7375EBB9EFEF2FCA1" author="Zheng, S. &amp; Lin, Y." pageId="23" pageNumber="484" refId="ref25901" refString="Zheng, S. &amp; Lin, Y. (2013) [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, 381 pp. [in Chinese]" type="book" year="2013">Zheng &amp; Lin (2013: 15)</bibRefCitation>
(distribution); 
<bibRefCitation id="D92262AA024CFFF73008BA7DFD14FCA1" author="Ito, T." box="[522,660,768,792]" pageId="23" pageNumber="484" pagination="55" refId="ref22607" refString="Ito, T. (2015) [A distributional record of Physopelta quadriguttata from Ishigaki Island, the Ryukyu Islands, Japan]. Gekkan- Mushi, 527, 55. [in Japanese]" type="journal article" year="2015">Ito (2015: 55)</bibRefCitation>
(distribution); 
<bibRefCitation id="D92262AA024CFFF7312EBA7CFB83FCA0" author="Ishikawa, T." box="[812,1027,768,793]" pageId="23" pageNumber="484" pagination="477 - 478" refId="ref22398" refString="Ishikawa, T. (2016) Family Largidae. In: Hayashi, M., Tomokuni, M., Yoshizawa, K. &amp; Ishikawa, T. (Eds.), Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, pp. 477 - 478. [in Japanese]" type="book chapter" year="2016">
<collectingRegion id="7F77D1B9024CFFF7312EBA7CFC07FCA1" box="[812,903,768,792]" country="Japan" name="Ishikawa" pageId="23" pageNumber="484">Ishikawa</collectingRegion>
(2016: 478)
</bibRefCitation>
(checklist: 
<collectingCountry id="C5A45FCB024CFFF73675BA7DFB35FCA0" box="[1143,1205,769,793]" name="Japan" pageId="23" pageNumber="484">Japan</collectingCountry>
); 
<bibRefCitation id="D92262AA024CFFF736C4BA7CFF5CFC8E" author="Zhou, Y. &amp; Redei, D." pageId="23" pageNumber="484" pagination="100914" refId="ref25939" refString="Zhou, Y. &amp; Redei, D. (2020) From lanceolate to plate-like: Gross morphology, terminology, and evolutionary trends of the trichophoran ovipositor. Arthropod Structure &amp; Development, 54 (2020), 100914. https: // doi. org / 10.1016 / j. asd. 2020.100914" type="journal article" year="2020">Zhou &amp; Rédei (2020: 5)</bibRefCitation>
(female genitalia).
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD0024CFFF73295BA19FB36FB7D" pageId="23" pageNumber="484" type="materials_examined">
<paragraph id="BD0C1F5B024CFFF73295BA19FB36FB7D" blockId="23.[151,1437,869,2012]" pageId="23" pageNumber="484">
<emphasis id="8FC7C349024CFFF73295BA19FE05FCC6" bold="true" box="[151,389,869,895]" pageId="23" pageNumber="484">Material examined.</emphasis>
<materialsCitation id="0DDB1506024CFFF73393BA1AFCF7FC39" ID-GBIF-Occurrence="3071281335" box="[401,887,869,896]" location="Non-types" pageId="23" pageNumber="484" specimenCount="107" specimenCount-female="41" specimenCount-male="66">
<location id="B86C4980024CFFF73393BA1AFD86FC39" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73393BA1AFD86FC39" box="[401,518,870,896]" name="Non-types" pageId="23" pageNumber="484">Non-types</location>
(macropterous 
<specimenCount id="ABB5D4D2024CFFF730BCBA19FC90FC39" box="[702,784,869,896]" pageId="23" pageNumber="484" type="male">66 ♂♂</specimenCount>
<specimenCount id="ABB5D4D2024CFFF7311EBA19FCECFC39" box="[796,876,869,896]" pageId="23" pageNumber="484" type="female">41 ♀♀</specimenCount>
)
</materialsCitation>
, 
<materialsCitation id="0DDB1506024CFFF7318BBA19FB00FC1D" ID-GBIF-Occurrence="3071281376" collectingDate="1999-04-21" collectionCode="TUA" collectorName="K. Takahashi" country="Japan" county="Ryukyu Islands" location="Ishigaki Is." municipality="Yaeyama Islands" pageId="23" pageNumber="484" specimenCount="1" specimenCount-female="1">
<emphasis id="8FC7C349024CFFF7318BBA19FA1CFCC6" bold="true" box="[905,1436,869,896]" pageId="23" pageNumber="484">
<collectingCountry id="C5A45FCB024CFFF7318BBA19FC67FCC6" box="[905,999,869,895]" name="Japan" pageId="23" pageNumber="484">JAPAN</collectingCountry>
: 
<collectingCounty id="546D67D7024CFFF731FABA19FB3BFCC6" box="[1016,1211,869,895]" pageId="23" pageNumber="484">Ryukyu Islands</collectingCounty>
: 
<emphasis id="8FC7C349024CFFF736CEBA1AFA13FC39" bold="true" box="[1228,1427,869,896]" italics="true" pageId="23" pageNumber="484">
<collectingMunicipality id="5D688521024CFFF736CEBA1AFA13FC39" box="[1228,1427,869,896]" pageId="23" pageNumber="484">Yaeyama Islands</collectingMunicipality>
</emphasis>
:
</emphasis>
<location id="B86C4980024CFFF73295BAF5FE98FC1D" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73295BAF5FE98FC1D" box="[151,280,905,932]" country="Japan" county="Ryukyu Islands" municipality="Yaeyama Islands" name="Ishigaki Is." pageId="23" pageNumber="484">Ishigaki Is.</location>
, 
<location id="B86C4980024CFFF73325BAF5FE4DFC1D" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73325BAF5FE4DFC1D" box="[295,461,905,932]" country="Japan" county="Ryukyu Islands" municipality="Yaeyama Islands" name="Nosoko Forest" pageId="23" pageNumber="484">Nosoko Forest</location>
Road, 
<date id="C90D399B024CFFF73021BAF5FD1BFC1A" box="[547,667,905,931]" pageId="23" pageNumber="484" value="1999-04-21">
<collectingDate id="D949C073024CFFF73021BAF5FD1BFC1A" box="[547,667,905,931]" pageId="23" pageNumber="484" value="1999-04-21">21.iv.1999</collectingDate>
</date>
, 
<collectingMethod id="64F2674C024CFFF730ABBAF5FC94FC1D" box="[681,788,905,932]" pageId="23" pageNumber="484">light trap</collectingMethod>
, leg. 
<collectorName id="10467A8D024CFFF73157BAF6FC6AFC1A" box="[853,1002,905,932]" pageId="23" pageNumber="484">K. Takahashi</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF731FDBAF5FBAAFC1D" box="[1023,1066,905,932]" pageId="23" pageNumber="484" type="female">1 ♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF73638BAF6FBF8FC1D" box="[1082,1144,906,932]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DDB1506024CFFF7368CBAF5FD5CFC71" ID-GBIF-Occurrence="3071281378" collectingDate="2004-11-18" collectionCode="TUA" collectorName="M. Hayashi" country="Japan" location="Hoshidate" municipality="Iriomote Is." pageId="23" pageNumber="484" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="5D688521024CFFF7368CBAF5FA95FC1D" box="[1166,1301,905,932]" pageId="23" pageNumber="484">Iriomote Is.</collectingMunicipality>
, 
<location id="B86C4980024CFFF73726BAF5FA18FC1A" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73726BAF5FA18FC1A" box="[1316,1432,905,931]" country="Japan" municipality="Iriomote Is." name="Hoshidate" pageId="23" pageNumber="484">Hoshidate</location>
, 
<date id="C90D399B024CFFF73295BAD1FE92FC7E" box="[151,274,941,967]" pageId="23" pageNumber="484" value="2004-11-18">
<collectingDate id="D949C073024CFFF73295BAD1FE92FC7E" box="[151,274,941,967]" pageId="23" pageNumber="484" value="2004-11-18">18.xi.2004</collectingDate>
</date>
, 
<collectingMethod id="64F2674C024CFFF7331CBAD1FE08FC71" box="[286,392,941,968]" pageId="23" pageNumber="484">light trap</collectingMethod>
, leg. 
<collectorName id="10467A8D024CFFF733C4BAD2FDCAFC7E" box="[454,586,941,968]" pageId="23" pageNumber="484">M. Hayashi</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF7305FBAD1FD07FC71" box="[605,647,941,968]" pageId="23" pageNumber="484" type="female">1 ♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF73097BAD2FD54FC71" box="[661,724,942,968]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="0DDB1506024CFFF730EABAD1FC5BFC55" ID-GBIF-Occurrence="3071281344" collectingDate="1932-07-26" collectionCode="ELKU" collectorName="K. Sato" country="Taiwan" location="Mt. Taiping-shan" municipality="Datong Township" pageId="23" pageNumber="484" specimenCount="1" specimenCount-male="1" stateProvince="Yilan">
<emphasis id="8FC7C349024CFFF730EABAD1FB95FC7E" bold="true" box="[744,1045,941,967]" pageId="23" pageNumber="484">
<collectingCountry id="C5A45FCB024CFFF730EABAD1FCDEFC7E" box="[744,862,941,967]" name="Taiwan" pageId="23" pageNumber="484">TAIWAN</collectingCountry>
: 
<collectingRegion id="7F77D1B9024CFFF73168BAD1FB90FC7E" box="[874,1040,941,967]" country="Taiwan" name="Yilan" pageId="23" pageNumber="484">Yilan County</collectingRegion>
:
</emphasis>
“Taiheizan (Taihoku-shû)” [= 
<collectingMunicipality id="5D688521024CFFF73770BAD2FEC0FC52" pageId="23" pageNumber="484">Datong Township</collectingMunicipality>
, 
<location id="B86C4980024CFFF73348BAAEFD92FC55" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73348BAAEFD92FC55" box="[330,530,977,1004]" country="Taiwan" municipality="Datong Township" name="Mt. Taiping-shan" pageId="23" pageNumber="484" stateProvince="Yilan">Mt. Taiping-shan</location>
?], 
<date id="C90D399B024CFFF7302FBAADFD2EFC52" box="[557,686,977,1003]" pageId="23" pageNumber="484" value="1932-07-26">
<collectingDate id="D949C073024CFFF7302FBAADFD2EFC52" box="[557,686,977,1003]" pageId="23" pageNumber="484" value="1932-07-26">26.vii.1932</collectingDate>
</date>
, leg. 
<collectorName id="10467A8D024CFFF730EBBAAEFCBDFC52" box="[745,829,977,1004]" pageId="23" pageNumber="484">K. Sato</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF7314CBAADFCF7FC55" box="[846,887,977,1004]" pageId="23" pageNumber="484" type="male">1 ♂</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF73181BAAEFC53FC55" box="[899,979,978,1004]" pageId="23" pageNumber="484">ELKU</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DDB1506024CFFF731E7BAAEFA19FC55" ID-GBIF-Occurrence="3071281373" box="[997,1433,977,1004]" collectingDate="1932-10-24" collectionCode="ELKU" country="Taiwan" pageId="23" pageNumber="484" specimenCount="2" specimenCount-female="2" stateProvince="Yilan">
as above but 
<date id="C90D399B024CFFF73674BAADFB66FC52" box="[1142,1254,977,1003]" pageId="23" pageNumber="484" value="1932-10-24">
<collectingDate id="D949C073024CFFF73674BAADFB66FC52" box="[1142,1254,977,1003]" pageId="23" pageNumber="484" value="1932-10-24">24.x.1932</collectingDate>
</date>
(
<specimenCount id="ABB5D4D2024CFFF736F7BAADFAB5FC55" box="[1269,1333,977,1004]" pageId="23" pageNumber="484" type="female">2 ♀♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF73743BAAEFA11FC55" box="[1345,1425,978,1004]" pageId="23" pageNumber="484">ELKU</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="0DDB1506024CFFF73295BA89FBD3FB8D" ID-GBIF-Occurrence="3071281409" collectingDate="2018-04-03" collectionCode="TUA" collectorName="Y. Fukuda &amp; S. Shimamoto &amp; S. Taru" country="Taiwan" county="Taoyuan District" elevation="1350" latitude="23.05211" location="Mt. Jianfu-shan" longLatPrecision="1" longitude="120.74519" municipality="Baoshan" pageId="23" pageNumber="484" specimenCount="55" specimenCount-female="22" specimenCount-male="33" stateProvince="Kaohsiung">
<emphasis id="8FC7C349024CFFF73295BA89FEE1FBB6" bold="true" box="[151,353,1013,1039]" pageId="23" pageNumber="484">
<collectingRegion id="7F77D1B9024CFFF73295BA89FEDCFBB6" box="[151,348,1013,1039]" country="Taiwan" name="Kaohsiung" pageId="23" pageNumber="484">Kaohsiung City</collectingRegion>
:
</emphasis>
<collectingCounty id="546D67D7024CFFF7336CBA8AFDB1FBB6" box="[366,561,1013,1040]" pageId="23" pageNumber="484">Taoyuan District</collectingCounty>
, 
<collectingMunicipality id="5D688521024CFFF73040BA89FD25FBB6" box="[578,677,1013,1039]" pageId="23" pageNumber="484">Baoshan</collectingMunicipality>
, near 
<location id="B86C4980024CFFF730F0BA8AFC2BFBB6" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF730F0BA8AFC2BFBB6" box="[754,939,1013,1040]" country="Taiwan" county="Taoyuan District" latitude="23.05211" longLatPrecision="1" longitude="120.74519" municipality="Baoshan" name="Mt. Jianfu-shan" pageId="23" pageNumber="484" stateProvince="Kaohsiung">Mt. Jianfu-shan</location>
, 
<geoCoordinate id="D887799C024CFFF731B9BA89FBD4FBB6" box="[955,1108,1013,1039]" degrees="23" direction="north" minutes="03" orientation="latitude" pageId="23" pageNumber="484" precision="1" seconds="07.6" value="23.05211">23°03'07.6&quot;N</geoCoordinate>
<geoCoordinate id="D887799C024CFFF73663BA89FA87FBB6" box="[1121,1287,1013,1039]" degrees="120" direction="east" minutes="44" orientation="longitude" pageId="23" pageNumber="484" precision="1" seconds="42.7" value="120.74519">120°44'42.7&quot;E</geoCoordinate>
, 
<quantity id="7A4BB2BE024CFFF7371ABA89FAE6FBA9" box="[1304,1382,1013,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="23" pageNumber="484" unit="m" value="1350.0">
<elevation id="369EF868024CFFF7371ABA89FAE6FBA9" box="[1304,1382,1013,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="23" pageNumber="484" unit="m" value="1350.0">1350m</elevation>
</quantity>
alt., 
<date id="C90D399B024CFFF73295BD65FE82FB8A" box="[151,258,1049,1075]" pageId="23" pageNumber="484" value="2018-04-03">
<collectingDate id="D949C073024CFFF73295BD65FE82FB8A" box="[151,258,1049,1075]" pageId="23" pageNumber="484" value="2018-04-03">3.iv.2018</collectingDate>
</date>
, 
<collectingMethod id="64F2674C024CFFF7330CBD65FEF7FB8D" box="[270,375,1049,1076]" pageId="23" pageNumber="484">light trap</collectingMethod>
, leg. 
<collectorName id="10467A8D024CFFF733B1BD66FDA9FB8A" box="[435,553,1049,1076]" pageId="23" pageNumber="484">Y. Fukuda</collectorName>
, 
<collectorName id="10467A8D024CFFF73037BD65FD54FB8A" box="[565,724,1049,1075]" pageId="23" pageNumber="484">S. Shimamoto</collectorName>
&amp; 
<collectorName id="10467A8D024CFFF730FBBD65FCCAFB8D" box="[761,842,1049,1076]" pageId="23" pageNumber="484">S. Taru</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF73158BD65FC29FB8D" box="[858,937,1049,1076]" pageId="23" pageNumber="484" type="male">33 ♂♂</specimenCount>
<specimenCount id="ABB5D4D2024CFFF731B2BD65FC7FFB8D" box="[944,1023,1049,1076]" pageId="23" pageNumber="484" type="female">22 ♀♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF7360EBD66FBCAFB8D" box="[1036,1098,1050,1076]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DDB1506024CFFF7365CBD66FB00FBE1" ID-GBIF-Occurrence="3071281358" collectingDate="2019-04-26" collectionCode="TUA" collectorName="S. Shimamoto" country="Taiwan" latitude="23.044277" location="Mt. Jianfu Shan" longLatPrecision="1" longitude="120.74519" municipality="Taoyuan District" pageId="23" pageNumber="484" specimenCount="19" specimenCount-female="4" specimenCount-male="15" stateProvince="Kaohsiung">
<collectingMunicipality id="5D688521024CFFF7365CBD66FA9DFB8A" box="[1118,1309,1049,1076]" pageId="23" pageNumber="484">Taoyuan District</collectingMunicipality>
, 
<location id="B86C4980024CFFF7372ABD66FF52FBEE" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF7372ABD66FF52FBEE" country="Taiwan" latitude="23.044277" longLatPrecision="1" longitude="120.74519" municipality="Taoyuan District" name="Mt. Jianfu Shan" pageId="23" pageNumber="484" stateProvince="Kaohsiung">Mt. Jianfu Shan</location>
, 
<geoCoordinate id="D887799C024CFFF732DCBD41FEF7FBEE" box="[222,375,1085,1111]" degrees="23" direction="north" minutes="02" orientation="latitude" pageId="23" pageNumber="484" precision="1" seconds="39.4" value="23.044277">23°02'39.4&quot;N</geoCoordinate>
<geoCoordinate id="D887799C024CFFF7337CBD41FDA5FBEE" box="[382,549,1085,1111]" degrees="120" direction="east" minutes="44" orientation="longitude" pageId="23" pageNumber="484" precision="1" seconds="42.7" value="120.74519">120°44'42.7&quot;E</geoCoordinate>
, 
<date id="C90D399B024CFFF73033BD41FD29FBEE" box="[561,681,1085,1111]" pageId="23" pageNumber="484" value="2019-04-26">
<collectingDate id="D949C073024CFFF73033BD41FD29FBEE" box="[561,681,1085,1111]" pageId="23" pageNumber="484" value="2019-04-26">26.iv.2019</collectingDate>
</date>
, leg. 
<collectorName id="10467A8D024CFFF730E4BD41FC06FBEE" box="[742,902,1085,1111]" pageId="23" pageNumber="484">S. Shimamoto</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF73194BD41FC64FBE1" box="[918,996,1085,1112]" pageId="23" pageNumber="484" type="male">15 ♂♂</specimenCount>
<specimenCount id="ABB5D4D2024CFFF731EEBD41FBADFBE1" box="[1004,1069,1085,1112]" pageId="23" pageNumber="484" type="female">4 ♀♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF73638BD42FBF9FBE1" box="[1082,1145,1086,1112]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="0DDB1506024CFFF73689BD41FD03FBC5" ID-GBIF-Occurrence="3071281327" collectingDate="1968-05-04" collectionCode="TUA" collectorName="Y. Watanabe" country="Taiwan" location="Wulai District" municipality="City" pageId="23" pageNumber="484" specimenCount="1" specimenCount-female="1" stateProvince="Taipei">
<emphasis id="8FC7C349024CFFF73689BD41FA99FBEE" bold="true" box="[1163,1305,1085,1111]" pageId="23" pageNumber="484">
<collectingRegion id="7F77D1B9024CFFF73689BD41FB55FBEE" box="[1163,1237,1085,1111]" country="Taiwan" name="Taipei" pageId="23" pageNumber="484">Taipei</collectingRegion>
<collectingMunicipality id="5D688521024CFFF736DFBD41FA94FBEE" box="[1245,1300,1085,1111]" pageId="23" pageNumber="484">City</collectingMunicipality>
:
</emphasis>
<location id="B86C4980024CFFF73723BD41FF47FBC2" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73723BD41FF47FBC2" country="Taiwan" municipality="City" name="Wulai District" pageId="23" pageNumber="484" stateProvince="Taipei">Wulai District</location>
, 
<date id="C90D399B024CFFF732D0BD1DFEB5FBC2" box="[210,309,1121,1147]" pageId="23" pageNumber="484" value="1968-05-04">
<collectingDate id="D949C073024CFFF732D0BD1DFEB5FBC2" box="[210,309,1121,1147]" pageId="23" pageNumber="484" value="1968-05-04">4.v.1968</collectingDate>
</date>
, leg. 
<collectorName id="10467A8D024CFFF73372BD1EFE7CFBC2" box="[368,508,1121,1148]" pageId="23" pageNumber="484">Y. Watanabe</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF7300DBD1DFDB0FBC2" box="[527,560,1121,1147]" pageId="23" pageNumber="484" type="female">1♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF7303FBD1EFDFCFBC5" box="[573,636,1122,1148]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="0DDB1506024CFFF7308CBD1DFE81FB19" ID-GBIF-Occurrence="3071281341" collectingDate="1968-05-17" collectionCode="TUA" collectorName="H. Sawada" country="Taiwan" location="Alishan Township" pageId="23" pageNumber="484" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiayi">
<emphasis id="8FC7C349024CFFF7308CBD1DFCC6FBC2" bold="true" box="[654,838,1121,1147]" pageId="23" pageNumber="484">
<collectingRegion id="7F77D1B9024CFFF7308CBD1DFCC1FBC2" box="[654,833,1121,1147]" country="Taiwan" name="Chiayi" pageId="23" pageNumber="484">Chiayi County</collectingRegion>
:
</emphasis>
<location id="B86C4980024CFFF7314FBD1DFB9CFBC2" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF7314FBD1DFB9CFBC2" box="[845,1052,1121,1147]" country="Taiwan" name="Alishan Township" pageId="23" pageNumber="484" stateProvince="Chiayi">Alishan Township</location>
, 
<date id="C90D399B024CFFF73625BD1DFB18FBC2" box="[1063,1176,1121,1147]" pageId="23" pageNumber="484" value="1968-05-17">
<collectingDate id="D949C073024CFFF73625BD1DFB18FBC2" box="[1063,1176,1121,1147]" pageId="23" pageNumber="484" value="1968-05-17">17.v.1968</collectingDate>
</date>
, leg. 
<collectorName id="10467A8D024CFFF736D1BD1EFACDFBC2" box="[1235,1357,1121,1148]" pageId="23" pageNumber="484">H. Sawada</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF7375DBD1DFA07FBC5" box="[1375,1415,1121,1148]" pageId="23" pageNumber="484" type="male">1 ♂</specimenCount>
<specimenCount id="ABB5D4D2024CFFF7378CBD1DFF2CFB19" pageId="23" pageNumber="484" type="female">1 ♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF732B9BDFAFF7AFB19" box="[187,250,1158,1184]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="0DDB1506024CFFF7330FBDF9FDC9FB7D" ID-GBIF-Occurrence="3071281417" collectingDate="2016-05-19" collectionCode="TUA" collectorName="N. Kaneko &amp; Y. Tamadera" country="Taiwan" latitude="24.127333" location="Ren'ai Township" pageId="23" pageNumber="484" specimenCount="23" specimenCount-female="8" specimenCount-male="15" stateProvince="Nantou">
<emphasis id="8FC7C349024CFFF7330FBDF9FE50FB26" bold="true" box="[269,464,1157,1183]" pageId="23" pageNumber="484">
<collectingRegion id="7F77D1B9024CFFF7330FBDF9FE4BFB26" box="[269,459,1157,1183]" country="Taiwan" name="Nantou" pageId="23" pageNumber="484">Nantou County</collectingRegion>
:
</emphasis>
<location id="B86C4980024CFFF733DABDF9FD1CFB26" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF733DABDF9FD1CFB26" box="[472,668,1157,1183]" country="Taiwan" latitude="24.127333" name="Ren'ai Township" pageId="23" pageNumber="484" stateProvince="Nantou">Ren’ai Township</location>
, 
<geoCoordinate id="D887799C024CFFF730AABDF9FCC1FB26" box="[680,833,1157,1183]" degrees="24" direction="north" minutes="07" orientation="latitude" pageId="23" pageNumber="484" precision="1" seconds="38.4" value="24.127333">24°07'38.4&quot;N</geoCoordinate>
<geoCoordinate id="D887799C024CFFF7314BBDF9FC70FB19" box="[841,1008,1157,1184]" pageId="23" pageNumber="484">121°15'88.8&quot;E</geoCoordinate>
, 
<date id="C90D399B024CFFF731FEBDF9FBEDFB26" box="[1020,1133,1157,1183]" pageId="23" pageNumber="484" value="2016-05-19">
<collectingDate id="D949C073024CFFF731FEBDF9FBEDFB26" box="[1020,1133,1157,1183]" pageId="23" pageNumber="484" value="2016-05-19">19.v.2016</collectingDate>
</date>
, 
<collectingMethod id="64F2674C024CFFF73678BDF9FB63FB19" box="[1146,1251,1157,1184]" pageId="23" pageNumber="484">light trap</collectingMethod>
, leg. 
<collectorName id="10467A8D024CFFF73720BDFAFA1CFB26" box="[1314,1436,1157,1184]" pageId="23" pageNumber="484">N. Kaneko</collectorName>
and 
<collectorName id="10467A8D024CFFF732C4BDD6FED2FB7A" box="[198,338,1193,1220]" pageId="23" pageNumber="484">Y. Tamadera</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF73363BDD5FE2FFB7D" box="[353,431,1193,1220]" pageId="23" pageNumber="484" type="male">15 ♂♂</specimenCount>
<specimenCount id="ABB5D4D2024CFFF733B4BDD5FE76FB7D" box="[438,502,1193,1220]" pageId="23" pageNumber="484" type="female">8 ♀♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF73000BDD6FDC0FB7D" box="[514,576,1194,1220]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DDB1506024CFFF73056BDD5FB32FB7D" ID-GBIF-Occurrence="3071281364" box="[596,1202,1193,1220]" collectingDate="1971-05-16" collectionCode="TUA" collectorName="K. Sakai" country="Taiwan" location="Nanshanchi" pageId="23" pageNumber="484" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nantou">
<location id="B86C4980024CFFF73056BDD5FD5BFB7A" LSID="urn:lsid:plazi:treatment:351AAE4D024CFFF93295B882FAAAFEA4:B86C4980024CFFF73056BDD5FD5BFB7A" box="[596,731,1193,1219]" country="Taiwan" name="Nanshanchi" pageId="23" pageNumber="484" stateProvince="Nantou">Nanshanchi</location>
, 
<date id="C90D399B024CFFF730E4BDD5FCD6FB7A" box="[742,854,1193,1219]" pageId="23" pageNumber="484" value="1971-05-16">
<collectingDate id="D949C073024CFFF730E4BDD5FCD6FB7A" box="[742,854,1193,1219]" pageId="23" pageNumber="484" value="1971-05-16">16.v.1971</collectingDate>
</date>
, leg. 
<collectorName id="10467A8D024CFFF73191BDD6FC73FB7A" box="[915,1011,1193,1220]" pageId="23" pageNumber="484">K. Sakai</collectorName>
(
<specimenCount id="ABB5D4D2024CFFF73607BDD5FBADFB7D" box="[1029,1069,1193,1220]" pageId="23" pageNumber="484" type="male">1 ♂</specimenCount>
<specimenCount id="ABB5D4D2024CFFF73636BDD5FBDEFB7D" box="[1076,1118,1193,1220]" pageId="23" pageNumber="484" type="female">1 ♀</specimenCount>
, 
<collectionCode id="DBA2879E024CFFF7366EBDD6FB2BFB7D" box="[1132,1195,1194,1220]" pageId="23" pageNumber="484">TUA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD0024CFFF732C5BDB1FC35F902" pageId="23" pageNumber="484" type="diagnosis">
<paragraph id="BD0C1F5B024CFFF732C5BDB1FC35F902" blockId="23.[151,1437,869,2012]" pageId="23" pageNumber="484">
<emphasis id="8FC7C349024CFFF732C5BDB1FEC2FB5E" bold="true" box="[199,322,1229,1255]" pageId="23" pageNumber="484">Diagnosis.</emphasis>
Recognized among other species of 
<taxonomicName id="7AB364D8024CFFF730DDBDB1FCDBFB5E" box="[735,859,1229,1255]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="8FC7C349024CFFF730DDBDB1FCDBFB5E" box="[735,859,1229,1255]" italics="true" pageId="23" pageNumber="484">Physopelta</emphasis>
</taxonomicName>
by a combination of the following characters: only macropterous morph known; body 2.8 times as long as maximum width across fore wings (
<figureCitation id="258803DE024CFFF736AEBD8DFAA5FAB2" box="[1196,1317,1265,1291]" captionStart="FIGURE 1" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[154,1430,425,1465]" captionTargetId="figure-130@3.[151,1436,417,1468]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of four Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta, male (A) and female (B) from Japan proper; Ph. (N.) fusciscutellata sp. nov., male from the main island of Taiwan (C) and female from the Ryukyu Islands, Japan (D); Ph. (N.) quadriguttata, male (E, G) and female (F) from the main island of Taiwan; Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (H). Arrows mark antenna with antennomeral oligomery and malformed calli. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668176" httpUri="https://zenodo.org/record/4668176/files/figure.png" pageId="23" pageNumber="484">Fig. 1C–E</figureCitation>
); calli and disc of pronotum pale brown, with orange margins (
<figureCitation id="258803DE024CFFF730F7BC69FCD2FA89" box="[757,850,1301,1328]" captionStart="FIGURE 5" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[154,1437,1028,1838]" captionTargetId="figure-420@12.[151,1437,1028,1838]" captionTargetPageId="12" captionText="FIGURE 5. Male pronotum and scutellum of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); macropterous morph of Ph. (N.) parviceps from Japan proper (C, D); macropterous morph of Ph. (N.) parviceps from the Ryukyu Islands, Japan (E); brachypterous morph of Ph. (N.) parviceps from Japan proper (F); Ph. (N.) fusciscutellata sp. nov. (G); Ph. (N.) quadriguttata (H). Arrows mark malformed calli. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668189" httpUri="https://zenodo.org/record/4668189/files/figure.png" pageId="23" pageNumber="484">Fig. 5H</figureCitation>
); scutellum brown, with numerous pale callosities; membrane of fore wing pale brown (
<figureCitation id="258803DE024CFFF73036BC45FD0AFAED" box="[564,650,1337,1364]" captionStart="FIGURE 6" captionStartId="13.[151,250,1708,1732]" captionTargetBox="[162,1433,890,1675]" captionTargetId="figure-351@13.[151,1437,885,1685]" captionTargetPageId="13" captionText="FIGURE 6. Left fore wing of five macropterous species of Physopelta (Neophysopelta), dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668191" httpUri="https://zenodo.org/record/4668191/files/figure.png" pageId="23" pageNumber="484">Fig. 6F</figureCitation>
); abdominal sternites paler brown with dark brown lunulae on sutures (
<figureCitation id="258803DE024CFFF7329DBC21FF7FFACE" box="[159,255,1373,1399]" captionStart="FIGURE 3" captionStartId="10.[151,250,1897,1921]" captionTargetBox="[415,1165,191,1860]" captionTargetId="figure-30@10.[392,1196,181,1874]" captionTargetPageId="10" captionText="FIGURE 3. Habitus images of five Physopelta (Neophysopelta) species, lateral view: Ph. (N.) gutta gutta, female from Japan proper (A); Ph. (N.) fusciscutellata sp. nov., female from the Ryukyu Islands, Japan (B); Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (C); Ph. (N.) parviceps, male from Japan proper (D) and female from the Ryukyu Islands (E); Ph. (N.) quadriguttata, male from the main island of Taiwan (F). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668183" httpUri="https://zenodo.org/record/4668183/files/figure.png" pageId="23" pageNumber="484">Figs. 3F</figureCitation>
, 
<figureCitation id="258803DE024CFFF7330FBC21FEAEFACE" box="[269,302,1373,1399]" captionStart="FIGURE 4" captionStartId="11.[151,250,1800,1824]" captionTargetBox="[162,1418,193,1763]" captionTargetId="figure-17@11.[155,1432,181,1776]" captionTargetPageId="11" captionText="FIGURE 4. Habitus images of five Physopelta (Neophysopelta) species, ventral view: Ph. (N.) gutta gutta, female from Japan proper (A); Ph. (N.) fusciscutellata sp. nov., female from the Ryukyu Islands, Japan (B); Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (C); Ph. (N.) parviceps, male from Japan proper (D) and female from the Ryukyu Islands (E); Ph. (N.) quadriguttata, male from the main island of Taiwan (F). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668187" httpUri="https://zenodo.org/record/4668187/files/figure.png" pageId="23" pageNumber="484">4F</figureCitation>
); compound eye more than 0.3 times as wide as vertex in dorsal view; antennomere I shorter than antennomere II; antennomere II nearly clavate; calli in male convex; punctures of scutellum denser in central part than in marginal part, as large as punctures of pronotum; anterior margins of fore wings not parallel to each other in rest; procoxal projection present, horn-shaped, curved, less than 1.5 times as long as its maximum width (
<figureCitation id="258803DE024CFFF7373BBCB5FA0EFA5D" box="[1337,1422,1481,1508]" captionStart="FIGURE 7" captionStartId="16.[151,250,1458,1482]" captionTargetBox="[151,1436,741,1435]" captionTargetId="figure-318@16.[151,1436,741,1435]" captionTargetPageId="16" captionText="FIGURE 7. Procoxa of five Physopelta (Neophysopelta) species, ventral view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviation: pp = procoxal projection. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4668193" httpUri="https://zenodo.org/record/4668193/files/figure.png" pageId="23" pageNumber="484">Fig. 7F</figureCitation>
); protrochanteral wrinkles present; male profemur more than 2 times as wide as mesofemur at widest part of each; male protibia lacking tooth at apex, with a single row of denticles throughout its length ventrally; peritreme of scent gland ostiole crescent-shaped, protruding posterolaterad (
<figureCitation id="258803DE024CFFF73119BF49FCF8F9F6" box="[795,888,1589,1615]" captionStart="FIGURE 8" captionStartId="17.[151,250,1278,1302]" captionTargetBox="[151,1436,561,1255]" captionTargetId="figure-215@17.[151,1436,561,1255]" captionTargetPageId="17" captionText="FIGURE 8. Left scent gland ostiole of five Physopelta (Neophysopelta) species, ventral view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviation: pe = peritreme. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668195" httpUri="https://zenodo.org/record/4668195/files/figure.png" pageId="23" pageNumber="484">Figs. 8F</figureCitation>
, 
<figureCitation id="258803DE024CFFF73186BF49FC33F9F6" box="[900,947,1589,1615]" captionStart="FIGURE 10" captionStartId="18.[151,250,1747,1771]" captionTargetBox="[156,1433,1060,1719]" captionTargetId="figure-119@18.[151,1436,1056,1724]" captionTargetPageId="18" captionText="FIGURE 10. Left scent gland ostiole of five Physopelta (Neophysopelta) species, lateral view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviation: pe = peritreme. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4668199" httpUri="https://zenodo.org/record/4668199/files/figure.png" pageId="23" pageNumber="484">10F</figureCitation>
); infolding of ventral rim of genital capsule less convex in middle part (
<figureCitation id="258803DE024CFFF733CABF25FD9CF9CA" box="[456,540,1625,1651]" captionStart="FIGURE 9" captionStartId="18.[151,250,899,923]" captionTargetBox="[151,1437,181,876]" captionTargetId="figure-30@18.[151,1437,181,876]" captionTargetPageId="18" captionText="FIGURE 9. Genital capsule of five Physopelta (Neophysopelta) species, posterior view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: vif = infolding of ventral rim; cr = crown of paramere. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668197" httpUri="https://zenodo.org/record/4668197/files/figure.png" pageId="23" pageNumber="484">Fig. 9F</figureCitation>
); outer margin of endophallic reservoir outgrowth emarginate in apical part (
<figureCitation id="258803DE024CFFF7376DBF25FF45F92E" captionStart="FIGURE 11" captionStartId="19.[151,250,1814,1838]" captionTargetBox="[247,1355,187,1784]" captionTargetId="figure-17@19.[224,1364,181,1790]" captionTargetPageId="19" captionText="FIGURE 11. Phallus of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: ad = aedeagus; bb = basal plate bridge; bp = basal plate; co = conjunctiva; cp = capitate process; dc = dorsal connective; ds = ductus seminis; ed = endophallic duct; eo = endophallic reservoir outgrowth; ep = erection fluid pump; er = endophallic reservoir; pt = phallotheca; 1ca, 2ca = first and second conjunctival process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668201" httpUri="https://zenodo.org/record/4668201/files/figure.png" pageId="23" pageNumber="484">Fig. 11F</figureCitation>
); stem of paramere not emarginate in basal part (
<figureCitation id="258803DE024CFFF730E2BF01FCC2F92E" box="[736,834,1661,1687]" captionStart="FIGURE 12" captionStartId="20.[151,250,1903,1927]" captionTargetBox="[160,1409,187,1865]" captionTargetId="figure-30@20.[151,1436,181,1880]" captionTargetPageId="20" captionText="FIGURE 12. Left paramere of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: ba = basal apodeme; cr = crown; ne = neck; st = stem. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668203" httpUri="https://zenodo.org/record/4668203/files/figure.png" pageId="23" pageNumber="484">Fig. 12F</figureCitation>
); crown of paramere at apex convex in posterior view; and inner margins of ring sclerites not parallel to each other (
<figureCitation id="258803DE024CFFF73141BFDDFC28F902" box="[835,936,1697,1723]" captionStart="FIGURE 13" captionStartId="21.[151,250,1679,1703]" captionTargetBox="[164,1421,468,1646]" captionTargetId="figure-131@21.[151,1436,453,1656]" captionTargetPageId="21" captionText="FIGURE 13. Female terminalia of four Physopelta (Neophysopelta) species, anterior view: Ph. (N.) gutta gutta (A); Ph. (N.) parviceps from Japan proper (B); Ph. (N.) parviceps from the Ryukyu Islands, Japan (C); Ph. (N.) fusciscutellata sp. nov. (D); Ph. (N.) quadriguttata (E). Abbreviations: go = gonangulum; lt9 = laterotergite IX; mt8 = mediotergite VIII; ra8, ra9 = ramus of valvula XIII, IX; rs = ring sclerite; va8, va9 = valvulae VIII, IX; vf8, vf9 = valvifers VIII, IX. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668205" httpUri="https://zenodo.org/record/4668205/files/figure.png" pageId="23" pageNumber="484">Fig. 13E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD0024CFFF832C5BFB9FB2EFF6C" lastPageId="24" lastPageNumber="485" pageId="23" pageNumber="484" type="description">
<paragraph id="BD0C1F5B024CFFF732C5BFB9FF53F82A" blockId="23.[151,1437,869,2012]" pageId="23" pageNumber="484">
<emphasis id="8FC7C349024CFFF732C5BFB9FE65F966" bold="true" box="[199,485,1733,1759]" pageId="23" pageNumber="484">Description of genitalia.</emphasis>
Genital capsule (
<figureCitation id="258803DE024CFFF730A8BFB9FD7DF966" box="[682,765,1733,1759]" captionStart="FIGURE 9" captionStartId="18.[151,250,899,923]" captionTargetBox="[151,1437,181,876]" captionTargetId="figure-30@18.[151,1437,181,876]" captionTargetPageId="18" captionText="FIGURE 9. Genital capsule of five Physopelta (Neophysopelta) species, posterior view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: vif = infolding of ventral rim; cr = crown of paramere. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668197" httpUri="https://zenodo.org/record/4668197/files/figure.png" pageId="23" pageNumber="484">Fig. 9F</figureCitation>
) spherical, semicircular in ventral view, smooth on surface; infolding of ventral rim less convex in middle part. Phallus (
<figureCitation id="258803DE024CFFF7313FBF95FC1EF8BA" box="[829,926,1769,1795]" captionStart="FIGURE 11" captionStartId="19.[151,250,1814,1838]" captionTargetBox="[247,1355,187,1784]" captionTargetId="figure-17@19.[224,1364,181,1790]" captionTargetPageId="19" captionText="FIGURE 11. Phallus of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: ad = aedeagus; bb = basal plate bridge; bp = basal plate; co = conjunctiva; cp = capitate process; dc = dorsal connective; ds = ductus seminis; ed = endophallic duct; eo = endophallic reservoir outgrowth; ep = erection fluid pump; er = endophallic reservoir; pt = phallotheca; 1ca, 2ca = first and second conjunctival process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668201" httpUri="https://zenodo.org/record/4668201/files/figure.png" pageId="23" pageNumber="484">Fig. 11F</figureCitation>
) oblong; capitate process membranous; basal plate and phallotheca coriaceous; conjunctiva with two pairs of partly sclerotized conjunctival appendages; endophallic reservoir with a pair of outgrowths; outer margin of outgrowth emarginate in apical part. Paramere (
<figureCitation id="258803DE024CFFF73736BE4DFA14F8F2" box="[1332,1428,1841,1867]" captionStart="FIGURE 12" captionStartId="20.[151,250,1903,1927]" captionTargetBox="[160,1409,187,1865]" captionTargetId="figure-30@20.[151,1436,181,1880]" captionTargetPageId="20" captionText="FIGURE 12. Left paramere of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); Ph. (N.) parviceps from Japan proper (C); Ph. (N.) parviceps from the Ryukyu Islands, Japan (D); Ph. (N.) fusciscutellata sp. nov. (E); Ph. (N.) quadriguttata (F). Abbreviations: ba = basal apodeme; cr = crown; ne = neck; st = stem. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668203" httpUri="https://zenodo.org/record/4668203/files/figure.png" pageId="23" pageNumber="484">Fig. 12F</figureCitation>
) longer than its maximum width across crown; stem not emarginate in basal part; crown at apex convex in posterior view.
</paragraph>
<paragraph id="BD0C1F5B024CFFF832C5BEE1FB2EFF6C" blockId="23.[151,1437,869,2012]" lastBlockId="24.[151,1436,151,357]" lastPageId="24" lastPageNumber="485" pageId="23" pageNumber="484">
Female terminalia (
<figureCitation id="258803DE024CFFF733A2BEE1FD81F80E" box="[416,513,1949,1975]" captionStart="FIGURE 13" captionStartId="21.[151,250,1679,1703]" captionTargetBox="[164,1421,468,1646]" captionTargetId="figure-131@21.[151,1436,453,1656]" captionTargetPageId="21" captionText="FIGURE 13. Female terminalia of four Physopelta (Neophysopelta) species, anterior view: Ph. (N.) gutta gutta (A); Ph. (N.) parviceps from Japan proper (B); Ph. (N.) parviceps from the Ryukyu Islands, Japan (C); Ph. (N.) fusciscutellata sp. nov. (D); Ph. (N.) quadriguttata (E). Abbreviations: go = gonangulum; lt9 = laterotergite IX; mt8 = mediotergite VIII; ra8, ra9 = ramus of valvula XIII, IX; rs = ring sclerite; va8, va9 = valvulae VIII, IX; vf8, vf9 = valvifers VIII, IX. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4668205" httpUri="https://zenodo.org/record/4668205/files/figure.png" pageId="23" pageNumber="484">Fig. 13E</figureCitation>
) triangular in anterior view; valvulae VIII and IX combining to form an ovipositor, fused with valvifers VIII and IX, respectively; ring sclerites coriaceous in marginal part, membranous in central part, protruding inward, with inner margins not parallel to each other. Spermatheca (
<figureCitation id="258803DE0243FFF8364AB9EBFB2BFF08" box="[1096,1195,151,177]" captionStart="FIGURE 14" captionStartId="22.[151,250,1605,1629]" captionTargetBox="[166,1429,458,1575]" captionTargetId="figure-160@22.[151,1436,453,1581]" captionTargetPageId="22" captionText="FIGURE 14. Spermatheca of four Physopelta (Neophysopelta) species: Ph. (N.) gutta gutta (A); Ph. (N.) parviceps from Japan proper (B); Ph. (N.) parviceps from the Ryukyu Islands, Japan (C); Ph. (N.) fusciscutellata sp. nov. (D); Ph. (N.) quadriguttata (E). Abbreviations: ar = apical receptacle; ip = intermediate part; sd = spermathecal duct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4668207" httpUri="https://zenodo.org/record/4668207/files/figure.png" pageId="24" pageNumber="485">Fig. 14E</figureCitation>
) membranous; apical receptacle spherical; intermediate part in width uniform; spermathecal duct apically widened.
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD00243FFF832C5B9A3FD95F8E2" pageId="24" pageNumber="485" type="discussion">
<paragraph id="BD0C1F5B0243FFF832C5B9A3FE0CFEDC" blockId="24.[151,1436,151,357]" pageId="24" pageNumber="485">
<emphasis id="8FC7C3490243FFF832C5B9A3FEBBFF40" bold="true" box="[199,315,223,249]" pageId="24" pageNumber="485">Remarks.</emphasis>
In terms of general appearance, 
<taxonomicName id="7AB364D80243FFF830B3B9A3FB11FF40" authorityName="Bergroth" authorityYear="1894" box="[689,1169,223,249]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="485" phylum="Arthropoda" rank="species" species="quadriguttata" subGenus="Neophysopelta">
<emphasis id="8FC7C3490243FFF830B3B9A3FCADFF40" box="[689,813,223,249]" italics="true" pageId="24" pageNumber="485">Physopelta</emphasis>
(
<emphasis id="8FC7C3490243FFF8313DB9A3FC65FF40" box="[831,997,223,249]" italics="true" pageId="24" pageNumber="485">Neophysopelta</emphasis>
) 
<emphasis id="8FC7C3490243FFF831F5B9A3FB11FF40" box="[1015,1169,223,249]" italics="true" pageId="24" pageNumber="485">quadriguttata</emphasis>
</taxonomicName>
strongly resembles 
<emphasis id="8FC7C3490243FFF83774B9A3FA15FF40" box="[1398,1429,223,249]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF832A3B87FFF34FEA5" box="[161,180,259,284]" italics="true" pageId="24" pageNumber="485">N</emphasis>
.) 
<emphasis id="8FC7C3490243FFF832C8B87FFE8AFEA4" box="[202,266,259,285]" italics="true" pageId="24" pageNumber="485">redeii</emphasis>
, but the former can be distinguished from the latter by the body form, which is 2.8 times as long as maximum width across the fore wings. In contrast, 
<emphasis id="8FC7C3490243FFF8309FB85BFD3CFEF8" box="[669,700,295,321]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF830D6B85BFD67FEF9" box="[724,743,295,320]" italics="true" pageId="24" pageNumber="485">N</emphasis>
.) 
<emphasis id="8FC7C3490243FFF830FCB85BFCBEFEF8" box="[766,830,295,321]" italics="true" pageId="24" pageNumber="485">redeii</emphasis>
has the body 2.5 times as long as the maximum width across the fore wings.
</paragraph>
<caption id="E9CC4FD30243FFF83295BF5EFBE0F93B" ID-DOI="http://doi.org/10.5281/zenodo.4668209" ID-Zenodo-Dep="4668209" httpUri="https://zenodo.org/record/4668209/files/figure.png" pageId="24" pageNumber="485" startId="24.[151,250,1570,1594]" targetBox="[155,1430,385,1539]" targetPageId="24">
<paragraph id="BD0C1F5B0243FFF83295BF5EFBE0F93B" blockId="24.[151,1437,1570,1667]" pageId="24" pageNumber="485">
<emphasis id="8FC7C3490243FFF83295BF5EFEA0F983" bold="true" box="[151,288,1570,1594]" pageId="24" pageNumber="485">FIGURE 15.</emphasis>
Habitus images of three 
<taxonomicName id="7AB364D80243FFF8301DBF5EFD10F983" box="[543,656,1570,1594]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="8FC7C3490243FFF8301DBF5EFD10F983" box="[543,656,1570,1594]" italics="true" pageId="24" pageNumber="485">Physopelta</emphasis>
</taxonomicName>
species confused with 
<taxonomicName id="7AB364D80243FFF8317EBF5EFB35F983" authorityName="Blote" authorityYear="1931" box="[892,1205,1570,1595]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="485" phylum="Arthropoda" rank="species" species="parviceps" subGenus="Neophysopelta">
<emphasis id="8FC7C3490243FFF8317EBF5EFC18F983" box="[892,920,1570,1594]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF831AFBF5EFBC4F983" box="[941,1092,1570,1594]" italics="true" pageId="24" pageNumber="485">Neophysopelta</emphasis>
) 
<emphasis id="8FC7C3490243FFF83651BF5FFB35F983" box="[1107,1205,1571,1594]" italics="true" pageId="24" pageNumber="485">parviceps</emphasis>
</taxonomicName>
in eastern Asia, dorsal view: 
<taxonomicName id="7AB364D80243FFF832DABF3AFD88F9E7" authorityName="DeGeer" authorityYear="1773" box="[216,520,1606,1631]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="485" phylum="Arthropoda" rank="species" species="albofasciata" subGenus="Physopelta">
<emphasis id="8FC7C3490243FFF832DABF3AFF74F9E7" box="[216,244,1606,1630]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF83308BF3AFEFBF9E7" box="[266,379,1606,1630]" italics="true" pageId="24" pageNumber="485">Physopelta</emphasis>
) 
<emphasis id="8FC7C3490243FFF83389BF3AFD88F9E7" box="[395,520,1606,1630]" italics="true" pageId="24" pageNumber="485">albofasciata</emphasis>
</taxonomicName>
, male (A) and female (B) from Malaysia; 
<taxonomicName id="7AB364D80243FFF831C6BF3AFB0EF9E7" authorityName="Stal" authorityYear="1863" box="[964,1166,1606,1631]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="462" phylum="Arthropoda" rank="species" species="cincticollis" subGenus="Neophysopelta">
<emphasis id="8FC7C3490243FFF831C6BF3AFC60F9E7" box="[964,992,1606,1630]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF831F5BF3BFB88F9E7" box="[1015,1032,1607,1630]" italics="true" pageId="24" pageNumber="485">N</emphasis>
.) 
<emphasis id="8FC7C3490243FFF8361DBF3AFB0EF9E7" box="[1055,1166,1606,1630]" italics="true" pageId="24" pageNumber="485">cincticollis</emphasis>
</taxonomicName>
, male (C) and female (D) from Vietnam; 
<taxonomicName id="7AB364D80243FFF83333BF16FE7CF93B" authorityName="Fabricius" authorityYear="1787" box="[305,508,1642,1667]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="462" phylum="Arthropoda" rank="species" species="slanbuschii" subGenus="Neophysopelta">
<emphasis id="8FC7C3490243FFF83333BF16FECDF93B" box="[305,333,1642,1666]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF83361BF17FEF4F93B" box="[355,372,1643,1666]" italics="true" pageId="24" pageNumber="485">N</emphasis>
.) 
<emphasis id="8FC7C3490243FFF8338BBF16FE7CF93B" box="[393,508,1642,1666]" italics="true" pageId="24" pageNumber="485">slanbuschii</emphasis>
</taxonomicName>
, male (E) and female (F) from Thailand. Scale bar: 5.0 mm.
</paragraph>
</caption>
<paragraph id="BD0C1F5B0243FFF832C5BFCDFD95F8E2" blockId="24.[151,1437,1713,2027]" pageId="24" pageNumber="485">
The teratological from in pronotum is found in 
<taxonomicName id="7AB364D80243FFF830CDBFCDFC46F972" authorityName="Bergroth" authorityYear="1894" box="[719,966,1713,1739]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="461" phylum="Arthropoda" rank="species" species="quadriguttata" subGenus="Neophysopelta">
<emphasis id="8FC7C3490243FFF830CDBFCDFD6EF972" box="[719,750,1713,1739]" italics="true" pageId="24" pageNumber="485">Ph</emphasis>
. (
<emphasis id="8FC7C3490243FFF83106BFCEFC97F972" box="[772,791,1714,1739]" italics="true" pageId="24" pageNumber="485">N</emphasis>
.) 
<emphasis id="8FC7C3490243FFF8312EBFCDFC46F972" box="[812,966,1713,1739]" italics="true" pageId="24" pageNumber="485">quadriguttata</emphasis>
</taxonomicName>
(
<figureCitation id="258803DE0243FFF831D6BFCDFBB6F972" box="[980,1078,1713,1739]" captionStart="FIGURE 1" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[154,1430,425,1465]" captionTargetId="figure-130@3.[151,1436,417,1468]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of four Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta, male (A) and female (B) from Japan proper; Ph. (N.) fusciscutellata sp. nov., male from the main island of Taiwan (C) and female from the Ryukyu Islands, Japan (D); Ph. (N.) quadriguttata, male (E, G) and female (F) from the main island of Taiwan; Ph. (N.) lutaspidata sp. nov., male from the main island of Taiwan (H). Arrows mark antenna with antennomeral oligomery and malformed calli. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668176" httpUri="https://zenodo.org/record/4668176/files/figure.png" pageId="24" pageNumber="485">Figs. 1G</figureCitation>
, 
<figureCitation id="258803DE0243FFF83643BFCDFBE7F975" box="[1089,1127,1713,1740]" captionStart="FIGURE 5" captionStartId="12.[151,250,1861,1885]" captionTargetBox="[154,1437,1028,1838]" captionTargetId="figure-420@12.[151,1437,1028,1838]" captionTargetPageId="12" captionText="FIGURE 5. Male pronotum and scutellum of five Physopelta (Neophysopelta) species, dorsal view: Ph. (N.) gutta gutta (A); Ph. (N.) lutaspidata sp. nov. (B); macropterous morph of Ph. (N.) parviceps from Japan proper (C, D); macropterous morph of Ph. (N.) parviceps from the Ryukyu Islands, Japan (E); brachypterous morph of Ph. (N.) parviceps from Japan proper (F); Ph. (N.) fusciscutellata sp. nov. (G); Ph. (N.) quadriguttata (H). Arrows mark malformed calli. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4668189" httpUri="https://zenodo.org/record/4668189/files/figure.png" pageId="24" pageNumber="485">5H</figureCitation>
). The calli of this individual is more convex in the left part than in the right part. In 
<taxonomicName id="7AB364D80243FFF8310CBFA9FC34F956" authorityName="Ahmad &amp; Abbas" authorityYear="1987" box="[782,948,1749,1775]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="478" phylum="Arthropoda" rank="subGenus" subGenus="Neophysopelta">
<emphasis id="8FC7C3490243FFF8310CBFA9FC34F956" box="[782,948,1749,1775]" italics="true" pageId="24" pageNumber="485">Neophysopelta</emphasis>
</taxonomicName>
, sexual dimorphism in the swelling degree of the calli and similar dimorphism between larger and smaller male has been known as mentioned in the previous section. In this individual, the left and right parts of the calli represents dimorphism between larger male and female/smaller male, respectively.
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD00243FFF832C5BE18FC8FF852" pageId="24" pageNumber="485" type="distribution">
<paragraph id="BD0C1F5B0243FFF832C5BE18FC8FF852" blockId="24.[151,1437,1713,2027]" pageId="24" pageNumber="485">
<emphasis id="8FC7C3490243FFF832C5BE18FEE1F8C7" bold="true" box="[199,353,1892,1918]" pageId="24" pageNumber="485">Distribution.</emphasis>
<collectingCountry id="C5A45FCB0243FFF83368BE19FE2EF8C6" box="[362,430,1893,1919]" name="Japan" pageId="24" pageNumber="485">Japan</collectingCountry>
: Ryukyu Islands (Ishigaki Island, Iriomote Island) (
<bibRefCitation id="D92262AA0243FFF831FDBE19FB7EF839" author="Miyamoto, S. &amp; Hayashi, M. &amp; Kohno, K." box="[1023,1278,1893,1920]" pageId="24" pageNumber="485" pagination="11 - 13" refId="ref23710" refString="Miyamoto, S., Hayashi, M. &amp; Kohno, K. (2005) New records of three Pentatomomorphan species (Heteroptera) from Japan. Japanese Journal of Systematic Entomology, 11, 11 - 13." type="journal article" year="2005">
Miyamoto 
<emphasis id="8FC7C3490243FFF8367DBE1AFB33F8C6" box="[1151,1203,1893,1919]" italics="true" pageId="24" pageNumber="485">et al</emphasis>
. 2005
</bibRefCitation>
; 
<bibRefCitation id="D92262AA0243FFF8370EBE19FF53F81A" author="Kohno, K. &amp; Hayashi, M. &amp; Miyamoto, S." pageId="24" pageNumber="485" pagination="90 - 91" refId="ref22979" refString="Kohno, K., Hayashi, M. &amp; Miyamoto, S. (2012) Family Largidae Amyot &amp; Serville, 1843. In: Ishikawa, T., Takai, M. &amp; Yasunaga, T. (Eds.), A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, pp. 399 - 402, pls. 90 - 91. [in Japanese]" type="book chapter" year="2012">
Kohno 
<emphasis id="8FC7C3490243FFF83760BE1AFA15F8C6" box="[1378,1429,1893,1919]" italics="true" pageId="24" pageNumber="485">et al</emphasis>
. 2012
</bibRefCitation>
; 
<bibRefCitation id="D92262AA0243FFF832DCBEF5FEBFF81D" author="Ito, T." box="[222,319,1929,1956]" pageId="24" pageNumber="485" pagination="55" refId="ref22607" refString="Ito, T. (2015) [A distributional record of Physopelta quadriguttata from Ishigaki Island, the Ryukyu Islands, Japan]. Gekkan- Mushi, 527, 55. [in Japanese]" type="journal article" year="2015">Ito 2015</bibRefCitation>
; present study); 
<collectingCountry id="C5A45FCB0243FFF833F5BEF5FDC9F81A" box="[503,585,1929,1955]" name="Taiwan" pageId="24" pageNumber="485">Taiwan</collectingCountry>
(main island) (
<bibRefCitation id="D92262AA0243FFF830F6BEF5FCF6F81A" author="Esaki, T." box="[756,886,1929,1955]" pageId="24" pageNumber="485" pagination="136 - 189" refId="ref21862" refString="Esaki, T. (1926) Verzeichniss der Hemiptera-heteroptera der Insel Formosa. Annales Musei Nationalis Hungarici, 24, 136 - 189." type="journal article" year="1926">Esaki 1926</bibRefCitation>
; 
<bibRefCitation id="D92262AA0243FFF83183BEF5FBACF81A" author="Kerzhner, I. M." box="[897,1068,1929,1955]" pageId="24" pageNumber="485" pagination="1843" refId="ref22916" refString="Kerzhner, I. M. (2001) Superfamily Pyrrhocoroidea Amyot &amp; Serville, 1843. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. The Netherlands Entomological Society, Amster- dam, pp. 245 - 247." type="book chapter" year="2001">Kerzhner 2001</bibRefCitation>
; 
<bibRefCitation id="D92262AA0243FFF8363ABEF5FB7AF81A" author="Redei, D. &amp; Tsai, J. - F. &amp; Yang, M. - M." box="[1080,1274,1929,1955]" pageId="24" pageNumber="485" refId="ref24020" refString="Redei, D., Tsai, J. - F. &amp; Yang, M. - M. (2009) Heteropteran Fauna of Taiwan: Cotton stainers and relatives (Hemiptera: Heteroptera: Pyrrhocoroidea). National Chung Hsing University, Taichung, 52 pp." type="book" year="2009">
Rédei 
<emphasis id="8FC7C3490243FFF83682BEF6FB31F81A" box="[1152,1201,1929,1955]" italics="true" pageId="24" pageNumber="485">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="D92262AA0243FFF83704BEF5FA17F81A" author="Stehlik, J. L." box="[1286,1431,1929,1955]" pageId="24" pageNumber="485" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík 2013</bibRefCitation>
; 
<bibRefCitation id="D92262AA0243FFF83295BED1FEE7F87E" author="Zheng, S. &amp; Lin, Y." box="[151,359,1965,1991]" pageId="24" pageNumber="485" refId="ref25901" refString="Zheng, S. &amp; Lin, Y. (2013) [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, 381 pp. [in Chinese]" type="book" year="2013">Zheng &amp; Lin 2013</bibRefCitation>
; present study); 
<collectingCountry id="C5A45FCB0243FFF8301EBED1FDDFF87E" box="[540,607,1965,1991]" name="China" pageId="24" pageNumber="485">China</collectingCountry>
(
<bibRefCitation id="D92262AA0243FFF8306EBED1FC80F87E" author="Stehlik, J. L." box="[620,768,1965,1991]" pageId="24" pageNumber="485" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík 2013</bibRefCitation>
); 
<collectingCountry id="C5A45FCB0243FFF83110BED1FCCCF87E" box="[786,844,1965,1991]" name="India" pageId="24" pageNumber="485">India</collectingCountry>
(
<bibRefCitation id="D92262AA0243FFF8315ABED1FC7FF87E" author="Bergroth, E." box="[856,1023,1965,1991]" pageId="24" pageNumber="485" pagination="152 - 164" refId="ref21667" refString="Bergroth, E. (1894) Rhynchota orientalia. Revue d'Entomologie, 13, 152 - 164. https: // doi. org / 10.5962 / bhl. part. 13239" type="journal article" year="1894">Bergroth 1894</bibRefCitation>
); 
<collectingCountry id="C5A45FCB0243FFF83610BED1FBC8F87E" box="[1042,1096,1965,1991]" name="Laos" pageId="24" pageNumber="485">Laos</collectingCountry>
(
<bibRefCitation id="D92262AA0243FFF83657BED1FB69F87E" author="Stehlik, J. L." box="[1109,1257,1965,1991]" pageId="24" pageNumber="485" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík 2013</bibRefCitation>
); 
<collectingCountry id="C5A45FCB0243FFF836F9BED1FABEF87E" box="[1275,1342,1965,1991]" name="Nepal" pageId="24" pageNumber="485">Nepal</collectingCountry>
(
<bibRefCitation id="D92262AA0243FFF83749BED1FF54F852" author="Stehlik, J. L." pageId="24" pageNumber="485" pagination="1 - 11" refId="ref24393" refString="Stehlik, J. L. (2004) Largidae and Pyrrhocoridae of Nepal (Heteroptera). Entomologica Basiliensia, 25, 1 - 11." type="journal article" year="2004">Stehlík 2004</bibRefCitation>
); 
<collectingCountry id="C5A45FCB0243FFF832E5BEADFECBF852" box="[231,331,2001,2027]" name="Thailand" pageId="24" pageNumber="485">Thailand</collectingCountry>
(
<bibRefCitation id="D92262AA0243FFF8335BBEADFE6EF852" author="Stehlik, J. L." box="[345,494,2001,2027]" pageId="24" pageNumber="485" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík 2013</bibRefCitation>
); 
<collectingCountry id="C5A45FCB0243FFF83003BEADFDE0F852" box="[513,608,2001,2027]" name="Vietnam" pageId="24" pageNumber="485">Vietnam</collectingCountry>
(
<bibRefCitation id="D92262AA0243FFF8306CBEADFC83F852" author="Stehlik, J. L." box="[622,771,2001,2027]" pageId="24" pageNumber="485" pagination="505 - 584" refId="ref24490" refString="Stehlik, J. L. (2013) Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae). Acta Entomologica Musei Nationalis Pragae, 53 (2), 505 - 584." type="journal article" year="2013">Stehlík 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F5A94CD00242FFF932C5B9EAFAAAFEA4" pageId="25" pageNumber="486" type="biology_ecology">
<paragraph id="BD0C1F5B0242FFF932C5B9EAFA80FF41" blockId="25.[151,1436,150,285]" pageId="25" pageNumber="486">
<emphasis id="8FC7C3490242FFF932C5B9EAFECEFF09" bold="true" box="[199,334,150,176]" pageId="25" pageNumber="486">Host plant.</emphasis>
In 
<collectingCountry id="C5A45FCB0242FFF93375B9EAFE4CFF09" box="[375,460,150,176]" name="Taiwan" pageId="25" pageNumber="486">Taiwan</collectingCountry>
, this species was collected from 
<taxonomicName id="7AB364D80242FFF93144B9EAFF55FF6C" authority="(Lam.) Mull. Arg." authorityName="Mull. Arg." baseAuthorityName="Lam." class="Actinopterygii" family="Osmeridae" genus="Mallotus" kingdom="Animalia" order="Osmeriformes" pageId="25" pageNumber="486" phylum="Chordata" rank="species" species="paniculatus">
<emphasis id="8FC7C3490242FFF93144B9EAFBB6FF09" box="[838,1078,150,176]" italics="true" pageId="25" pageNumber="486">Mallotus paniculatus</emphasis>
(Lam.) Müll. Arg. (Euphorbiaceae)
</taxonomicName>
, and 
<taxonomicName id="7AB364D80242FFF9330CB9C6FD7CFF6D" authority="Munro." authorityName="Munro." box="[270,764,186,213]" class="Liliopsida" family="Poaceae" genus="Dendrocalamus" kingdom="Plantae" order="Poales" pageId="25" pageNumber="486" phylum="Tracheophyta" rank="species" species="latiflorus">
<emphasis id="8FC7C3490242FFF9330CB9C6FDAEFF6D" box="[270,558,186,212]" italics="true" pageId="25" pageNumber="486">Dendrocalamus latiflorus</emphasis>
Munro. (Poaceae)
</taxonomicName>
(
<bibRefCitation id="D92262AA0242FFF9310BB9C6FC5FFF6D" author="Zheng, S. &amp; Lin, Y." box="[777,991,186,212]" pageId="25" pageNumber="486" refId="ref25901" refString="Zheng, S. &amp; Lin, Y. (2013) [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, 381 pp. [in Chinese]" type="book" year="2013">Zheng &amp; Lin 2013</bibRefCitation>
), but nymphs have not been collected to date. Therefore, the “true” host plant for 
<taxonomicName id="7AB364D80242FFF93063B9A2FBBEFF41" authorityName="Bergroth" authorityYear="1894" box="[609,1086,222,249]" class="Insecta" family="Largidae" genus="Physopelta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="486" phylum="Arthropoda" rank="species" species="quadriguttata" subGenus="Neophysopelta">
<emphasis id="8FC7C3490242FFF93063B9A2FD5DFF41" box="[609,733,222,248]" italics="true" pageId="25" pageNumber="486">Physopelta</emphasis>
(
<emphasis id="8FC7C3490242FFF930EFB9A2FC13FF41" box="[749,915,222,248]" italics="true" pageId="25" pageNumber="486">Neophysopelta</emphasis>
) 
<emphasis id="8FC7C3490242FFF931A6B9A2FBBEFF41" box="[932,1086,222,248]" italics="true" pageId="25" pageNumber="486">quadriguttata</emphasis>
</taxonomicName>
is still unknown.
</paragraph>
<paragraph id="BD0C1F5B0242FFF932C5B87EFAAAFEA4" blockId="25.[151,1436,150,285]" box="[199,1322,258,285]" pageId="25" pageNumber="486">
<emphasis id="8FC7C3490242FFF932C5B87EFEA7FEA5" bold="true" box="[199,295,258,284]" pageId="25" pageNumber="486">Biology.</emphasis>
Adults are attracted to artificial light (
<bibRefCitation id="D92262AA0242FFF930D1B87EFC17FEA5" author="Redei, D. &amp; Tsai, J. - F. &amp; Yang, M. - M." box="[723,919,258,284]" pageId="25" pageNumber="486" refId="ref24020" refString="Redei, D., Tsai, J. - F. &amp; Yang, M. - M. (2009) Heteropteran Fauna of Taiwan: Cotton stainers and relatives (Hemiptera: Heteroptera: Pyrrhocoroidea). National Chung Hsing University, Taichung, 52 pp." type="book" year="2009">
Rédei 
<emphasis id="8FC7C3490242FFF9311FB87FFCD5FEA5" box="[797,853,258,284]" italics="true" pageId="25" pageNumber="486">et al.</emphasis>
2009
</bibRefCitation>
; 
<bibRefCitation id="D92262AA0242FFF931A1B87EFBF7FEA5" author="Zheng, S. &amp; Lin, Y." box="[931,1143,258,284]" pageId="25" pageNumber="486" refId="ref25901" refString="Zheng, S. &amp; Lin, Y. (2013) [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, 381 pp. [in Chinese]" type="book" year="2013">Zheng &amp; Lin 2013</bibRefCitation>
; present paper).
</paragraph>
</subSubSection>
</treatment>
</document>