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<document id="29B64CBC62EE1A2660E3D46BDBF6646D" ID-CLB-Dataset="5003" ID-DOI="10.1649/0010-065X-71.3.532" ID-GBIF-Dataset="8e549dbe-957f-47cc-91ff-dea6e2fc272a" ID-ISSN="1938-4394" ID-Zenodo-Dep="4788691" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621946305265" checkinUser="carolina" docAuthor="Smith, Andrew B. T. &amp; Paulsen, M. J." docDate="2017" docId="B4615401FF862C18FF67AA66FF35FC2B" docLanguage="en" docName="TheColeopBull.71.3.532-536.pdf" docOrigin="The Coleopterists Bulletin 71 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-71.3.532" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Podolasia parapilosa Smith and Paulsen 2017" docType="treatment" docVersion="11" lastPageNumber="536" masterDocId="48582C79FF842C1CFFDCAF1EFFDCFFF2" masterDocTitle="Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico" masterLastPageNumber="536" masterPageNumber="532" pageNumber="534" updateTime="1698989653070" updateUser="ExternalLinkService">
<mods:mods id="2F3F0158D0805936E200028505227DC4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EF98EFA4E68F811CE99118AABFE9400A">Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico</mods:title>
</mods:titleInfo>
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<mods:namePart id="4CB7EBA937AACDBA381010C8A039D8D8">Smith, Andrew B. T.</mods:namePart>
</mods:name>
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<mods:role id="5BAFF4A9088CA8801634113AFE5F7047">
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</mods:role>
<mods:namePart id="F14BAE56E3058A5088DFF3E2DCDB041A">Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="98B1E1C1D131EDF1B56575EE122155EC">text</mods:typeOfResource>
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<mods:title id="BD8BB4A5C2F51AFFFE4134B1C94E94B2">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="8B2BBF5D84A0DD4B1F126D574385D362">
<mods:date id="4203F2568CACF336713F8C8A788022A8">2017</mods:date>
<mods:detail id="44387EC4C7878AF6C382D1C2BB5BEF50" type="pubDate">
<mods:number id="6F4717561979F895833DDE9F45FA54C4">2017-09-18</mods:number>
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<mods:number id="DE03093051B5E6B850414CACEF329B50">71</mods:number>
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<mods:start id="F0DD6124C52EACEC8F5CF6E045F6708C">532</mods:start>
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<mods:url id="68B2E300262EE59E3079A087433DE19E">http://dx.doi.org/10.1649/0010-065x-71.3.532</mods:url>
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<mods:classification id="A6FF3D2F65AA33646A1CFD5F533C92E1">journal article</mods:classification>
<mods:identifier id="B38A95A306FD3F89FE99934D4B63DCF6" type="CLB-Dataset">5003</mods:identifier>
<mods:identifier id="7F643E2800CAB815C09C908588637CD9" type="DOI">10.1649/0010-065X-71.3.532</mods:identifier>
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<treatment id="B4615401FF862C18FF67AA66FF35FC2B" ID-DOI="http://doi.org/10.5281/zenodo.5191749" ID-GBIF-Taxon="184097535" ID-Zenodo-Dep="5191749" LSID="urn:lsid:plazi:treatment:B4615401FF862C18FF67AA66FF35FC2B" httpUri="http://treatment.plazi.org/id/B4615401FF862C18FF67AA66FF35FC2B" lastPageId="4" lastPageNumber="536" pageId="2" pageNumber="534">
<subSubSection id="74D2B69CFF862C1EFF67AA66FE19FA5A" pageId="2" pageNumber="534" type="nomenclature">
<paragraph id="3C77E517FF862C1EFF67AA66FE19FA5A" blockId="2.[151,639,1400,1730]" pageId="2" pageNumber="534">
<emphasis id="0EBC3905FF862C1EFF67AA66FE19FA5A" bold="true" pageId="2" pageNumber="534">
<taxonomicName id="FBC89E94FF862C1EFF67AA66FD89FA7C" ID-CoL="4KQRM" authority="Smith and Paulsen" authorityName="Smith and Paulsen" authorityYear="2017" box="[187,597,1400,1422]" class="Insecta" family="Scarabaeidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="534" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF862C1EFF67AA66FE50FA7F" bold="true" box="[187,396,1400,1421]" italics="true" pageId="2" pageNumber="534">Podolasia parapilosa</emphasis>
Smith and Paulsen
</taxonomicName>
, new species
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="74D2B69CFF862C1EFE8CAAB0FE19FA31" box="[336,453,1453,1475]" pageId="2" pageNumber="534" type="description">
<paragraph id="3C77E517FF862C1EFE8CAAB0FE19FA31" blockId="2.[151,639,1400,1730]" box="[336,453,1453,1475]" pageId="2" pageNumber="534">
(
<figureCitation id="A4F3F992FF862C1EFE8BAAB0FE61FA31" box="[343,445,1453,1475]" captionStart="Figs" captionStartId="3.[197,236,1492,1511]" captionTargetBox="[177,1188,196,1474]" captionTargetId="figure-71@3.[175,1190,194,1476]" captionTargetPageId="3" captionText="Figs. 6–11. Podolasia parapilosa, holotype and paratype. 6) Holotype, dorsal habitus; 7) Holotype, oblique habitus; 8) Paratype, ventral habitus; 9) Holotype sclerotized genital capsule; 10) Holotype, apical view of the head; 11) Holotype parameres." figureDoi="http://doi.org/10.5281/zenodo.4788723" httpUri="https://zenodo.org/record/4788723/files/figure.png" pageId="2" pageNumber="534">Figs. 6–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74D2B69CFF862C1EFF73AAC6FC11FD97" pageId="2" pageNumber="534" type="materials_examined">
<paragraph id="3C77E517FF862C1EFF73AAC6FC11FD97" blockId="2.[151,639,1400,1730]" lastBlockId="2.[685,1173,193,1730]" pageId="2" pageNumber="534">
<emphasis id="0EBC3905FF862C1EFF73AAC6FE9FFA1F" bold="true" box="[175,323,1496,1517]" pageId="2" pageNumber="534">Type Material.</emphasis>
<materialsCitation id="8CA0EF4AFF862C1EFE94AAC6FD9CF97F" ID-GBIF-Occurrence="3333868302" collectingDate="2011-06-21" collectionCode="UNMC" collectorName="D. C. Lightfoot" country="Mexico" county="Cuatro" latitude="26.84484" location="Cienegas Nat'" longLatPrecision="1" longitude="-102.17866" municipality="Cienegas Nat'" pageId="2" pageNumber="534" specimenCount="1" specimenCount-male="1" stateProvince="Coahuila" typeStatus="holotype">
<typeStatus id="E3735BB5FF862C1EFE94AAC6FE7CFA1F" box="[328,416,1496,1517]" pageId="2" pageNumber="534" type="holotype">Holotype</typeStatus>
<specimenCount id="2ACE2E9EFF862C1EFE79AAC6FE0FFA1F" box="[421,467,1496,1517]" pageId="2" pageNumber="534" type="male">male</specimenCount>
(
<collectionCode id="5AD97DD2FF862C1EFE02AAC6FDF4FA1F" box="[478,552,1496,1517]" pageId="2" pageNumber="534">UNMC</collectionCode>
) labeled: a) “
<collectingCountry id="44DFA587FF862C1EFF1EAAEDFED2F9FA" box="[194,270,1523,1544]" name="Mexico" pageId="2" pageNumber="534">Mexico</collectingCountry>
:
<collectingRegion id="FE0C2BF5FF862C1EFEC1AAEDFEAAF9FA" box="[285,374,1523,1544]" country="Mexico" name="Coahuila de Zaragoza" pageId="2" pageNumber="534">Coahuila</collectingRegion>
:
<collectingCounty id="D5169D9BFF862C1EFE59AAEDFE19F9FA" box="[389,453,1523,1544]" pageId="2" pageNumber="534">Cuatro</collectingCounty>
/
<collectingMunicipality id="DC137F6DFF862C1EFE38AAEDFDADF9FA" box="[484,625,1522,1544]" pageId="2" pageNumber="534">Cienegas Nat’</collectingMunicipality>
l. Preserve /
<date id="4876C3D7FF862C1EFF24A913FEB8F9D0" box="[248,356,1549,1570]" pageId="2" pageNumber="534" value="2011-06-21">
<collectingDate id="58323A3FFF862C1EFF24A913FEB8F9D0" box="[248,356,1549,1570]" pageId="2" pageNumber="534" value="2011-06-21">21.VI.2011</collectingDate>
</date>
/
<geoCoordinate id="59FC83D0FF862C1EFEA8A913FE34F9D0" box="[372,488,1549,1570]" degrees="26.84484" direction="north" orientation="latitude" pageId="2" pageNumber="534" precision="1" value="26.84484">26.84484°N</geoCoordinate>
<geoCoordinate id="59FC83D0FF862C1EFE31A913FDAEF9D0" box="[493,626,1549,1570]" degrees="102.17866" direction="west" orientation="longitude" pageId="2" pageNumber="534" precision="1" value="-102.17866">102.17866°W</geoCoordinate>
/
<collectorName id="913D80C1FF862C1EFF4BA936FEC3F9CF" box="[151,287,1576,1597]" pageId="2" pageNumber="534">D.C. Lightfoot</collectorName>
et al., colrs”; b) “Site B3 / ~
<locationDeviation id="2DC5FC35FF862C1EFDFAA936FF09F9AA" bearing="225.0" distance="19430.0" pageId="2" pageNumber="534">
<quantity id="FB3048F2FF862C1EFDFAA936FDA1F9CF" box="[550,637,1576,1597]" metricMagnitude="4" metricUnit="m" metricValue="1.943" pageId="2" pageNumber="534" unit="km" value="19.43">19.43 km</quantity>
SW of
</locationDeviation>
Cuatro / gypsum dunes, pedestals / general collecting”; c) on red paper, “HOLOTYPE /
<taxonomicName id="FBC89E94FF862C1EFF4BA969FEB5F97F" authorityName="Smith and Paulsen" authorityYear="2017" box="[151,361,1655,1677]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="534" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF862C1EFF4BA969FEB5F97F" box="[151,361,1655,1677]" italics="true" pageId="2" pageNumber="534">Podolasia parapilosa</emphasis>
</taxonomicName>
/ Smith &amp; Paulsen”.
</materialsCitation>
<materialsCitation id="8CA0EF4AFF862C1EFD96A966FC7EFF03" ID-GBIF-Occurrence="3333868303" pageId="2" pageNumber="534" typeStatus="paratype">
Eight male
<typeStatus id="E3735BB5FF862C1EFF13A98CFEF6F955" box="[207,298,1682,1703]" pageId="2" pageNumber="534" type="paratype">paratypes</typeStatus>
(3
<collectionCode id="5AD97DD2FF862C1EFE93A98CFE44F955" box="[335,408,1682,1703]" pageId="2" pageNumber="534">CMNC</collectionCode>
, 1
<collectionCode id="5AD97DD2FF862C1EFE66A98CFE24F955" box="[442,504,1682,1703]" pageId="2" pageNumber="534">MJPC</collectionCode>
, 1
<collectionCode id="5AD97DD2FF862C1EFDC6A98CFDB9F955" box="[538,613,1682,1703]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="2" pageNumber="534">UNAM</collectionCode>
, 2
<collectionCode id="5AD97DD2FF862C1EFF4BA9B3FF3DF930" box="[151,225,1709,1730]" pageId="2" pageNumber="534">UNMC</collectionCode>
, 1
<collectionCode id="5AD97DD2FF862C1EFEDEA9B3FE94F930" box="[258,328,1709,1730]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="2" pageNumber="534">UNSM</collectionCode>
) labeled as holotype except for c) on yellow paper “PARATYPE /
<taxonomicName id="FBC89E94FF862C1EFC21AFDFFD38FF03" authorityName="Howden" authorityYear="1954" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="534" phylum="Arthropoda" rank="species" species="paradistinct">
<emphasis id="0EBC3905FF862C1EFC21AFDFFD38FF03" italics="true" pageId="2" pageNumber="534">Podolasia parapilosa</emphasis>
</taxonomicName>
/ Smith &amp; Paulsen”.
</materialsCitation>
<materialsCitation id="8CA0EF4AFF862C1EFC74AFC2FCDEFE34" ID-GBIF-Occurrence="3333868301" pageId="2" pageNumber="534" typeStatus="paratype">
Twenty male
<typeStatus id="E3735BB5FF862C1EFBF8AFC2FBA6FF03" box="[1060,1146,220,241]" pageId="2" pageNumber="534" type="paratype">paratypes</typeStatus>
(4 CMNC, 2 MJPC, 1 UNAM, 10 UNMC, 3 UNSM) labeled: a) “
<collectingCountry id="44DFA587FF862C1EFCC6AE0FFCBFFED4" box="[794,867,273,294]" name="Mexico" pageId="2" pageNumber="534">Mexico</collectingCountry>
:
<collectingRegion id="FE0C2BF5FF862C1EFCB5AE0FFC62FED4" box="[873,958,273,294]" country="Mexico" name="Coahuila de Zaragoza" pageId="2" pageNumber="534">Coahuila</collectingRegion>
: Cuatro / Cienegas Prot. Area /
<date id="4876C3D7FF862C1EFD26AE32FC8BFEB3" box="[762,855,300,321]" pageId="2" pageNumber="534" value="2010-05-18">18.V.2010</date>
/
<geoCoordinate id="59FC83D0FF862C1EFCABAE32FC07FEB3" box="[887,987,300,321]" degrees="26.8336" direction="north" orientation="latitude" pageId="2" pageNumber="534" precision="5" value="26.8336">26.8336°N</geoCoordinate>
<geoCoordinate id="59FC83D0FF862C1EFC34AE32FB80FEB3" box="[1000,1116,300,321]" degrees="102.1620" direction="west" orientation="longitude" pageId="2" pageNumber="534" precision="5" value="-102.162">102.1620°W</geoCoordinate>
/ D. Lightfoot, K.R. Wetherill, et al.”; b) “Site B1 / ~
<quantity id="FB3048F2FF862C1EFB83AE58FB48FEA9" box="[1119,1172,326,347]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="534" unit="km" value="19.0">19km</quantity>
SW Cuatrocienegas / gypsum dune / light trap, oatmeal line / general collecting”; c) on yellow paper “PARATYPE /
<taxonomicName id="FBC89E94FF862C1EFC8CAE88FBC6FE5E" authorityName="Smith and Paulsen" authorityYear="2017" box="[848,1050,406,428]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="534" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF862C1EFC8CAE88FBC6FE5E" box="[848,1050,406,428]" italics="true" pageId="2" pageNumber="534">Podolasia parapilosa</emphasis>
</taxonomicName>
/ Smith &amp; Paulsen”.
</materialsCitation>
<materialsCitation id="8CA0EF4AFF862C1EFCDAAEAFFC11FD97" ID-GBIF-Occurrence="3333868304" pageId="2" pageNumber="534" typeStatus="paratype">
Twelve male
<typeStatus id="E3735BB5FF862C1EFCA0AEAFFC0DFE34" box="[892,977,433,454]" pageId="2" pageNumber="534" type="paratype">paratypes</typeStatus>
(2 CMNC, 8 UNMC, 2 UNSM) labeled: a) “
<collectingCountry id="44DFA587FF862C1EFC50AED5FC08FE12" box="[908,980,459,480]" name="Mexico" pageId="2" pageNumber="534">Mexico</collectingCountry>
:
<collectingRegion id="FE0C2BF5FF862C1EFC3CAED5FBE8FE12" box="[992,1076,459,480]" country="Mexico" name="Coahuila de Zaragoza" pageId="2" pageNumber="534">Coahuila</collectingRegion>
: Cuatro- / Cienegas Prot. Area
<date id="4876C3D7FF862C1EFCBDAEF8FC62FE09" box="[865,958,486,507]" pageId="2" pageNumber="534" value="2010-05-18">18.V.2010</date>
/
<geoCoordinate id="59FC83D0FF862C1EFC10AEF8FBF8FE09" box="[972,1060,486,507]" degrees="26.833" direction="north" orientation="latitude" pageId="2" pageNumber="534" precision="55" value="26.833">26.833°N</geoCoordinate>
<geoCoordinate id="59FC83D0FF862C1EFBF4AEF8FB4CFE09" box="[1064,1168,486,507]" degrees="102.162" direction="west" orientation="longitude" pageId="2" pageNumber="534" precision="55" value="-102.162">102.162°W</geoCoordinate>
, ~
<quantity id="FB3048F2FF862C1EFD1DAD1FFD21FDE4" box="[705,765,513,534]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="534" unit="km" value="20.0">20 km</quantity>
/ SW Cuatrocienegas, colr. D.C. / Lightfoot, K.R. Wetherill, et al.”; b) “Site B1 light trap / Gypsum Dunes”; c) on yellow paper “PARATYPE /
<taxonomicName id="FBC89E94FF862C1EFBE5AD2BFCD2FD94" authorityName="Harold" authorityYear="1869" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="0EBC3905FF862C1EFBE5AD2BFCD2FD94" italics="true" pageId="2" pageNumber="534">Podolasia parapilosa</emphasis>
</taxonomicName>
/ Smith &amp; Paulsen”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="74D2B69CFF862C18FD19AD75FEC7FEFE" lastPageId="4" lastPageNumber="536" pageId="2" pageNumber="534" type="description">
<paragraph id="3C77E517FF862C1FFD19AD75FBB7F955" blockId="2.[685,1173,193,1730]" lastBlockId="3.[707,1193,1603,1730]" lastPageId="3" lastPageNumber="535" pageId="2" pageNumber="534">
<emphasis id="0EBC3905FF862C1EFD19AD75FC9CFD72" bold="true" box="[709,832,619,640]" pageId="2" pageNumber="534">Description.</emphasis>
Male, length 6.5 mm, greatest width 2.5 mm. Color dorsally and ventrally reddish brown with head and pronotum slightly darker. Head apically arcuate (not subcircular), clypeal and genal margins not continuous. Clypeal length and width subequal (
<figureCitation id="A4F3F992FF862C1EFCD6ADEEFC97FCF7" box="[778,843,752,773]" captionStart="Figs" captionStartId="3.[197,236,1492,1511]" captionTargetBox="[177,1188,196,1474]" captionTargetId="figure-71@3.[175,1190,194,1476]" captionTargetPageId="3" captionText="Figs. 6–11. Podolasia parapilosa, holotype and paratype. 6) Holotype, dorsal habitus; 7) Holotype, oblique habitus; 8) Paratype, ventral habitus; 9) Holotype sclerotized genital capsule; 10) Holotype, apical view of the head; 11) Holotype parameres." figureDoi="http://doi.org/10.5281/zenodo.4788723" httpUri="https://zenodo.org/record/4788723/files/figure.png" pageId="2" pageNumber="534">Fig. 10</figureCitation>
), sides reflexed, anteriorly thickened either side of weak median emargination, posteriorly not coplanar with frons; frontoclypeal suture carinate; clypeal disc with scattered large punctures, each with a long seta, surface between punctures roughened with micropunctures. Labrum not visible, presumably fused with clypeus. Genal margin distinctly produced, rounded; raised edge thickened anteriorly. Frons lacking secondary transverse carina, instead with oblique irregular carinae parallel to eyes; surface rugopunctate. Mouthparts reduced, except for labial and maxillary palpomeres. Antenna with 9 antennomeres (
<figureCitation id="A4F3F992FF862C1EFC56AB31FC0FFBB6" box="[906,979,1071,1092]" captionStart="Figs" captionStartId="3.[197,236,1492,1511]" captionTargetBox="[177,1188,196,1474]" captionTargetId="figure-71@3.[175,1190,194,1476]" captionTargetPageId="3" captionText="Figs. 6–11. Podolasia parapilosa, holotype and paratype. 6) Holotype, dorsal habitus; 7) Holotype, oblique habitus; 8) Paratype, ventral habitus; 9) Holotype sclerotized genital capsule; 10) Holotype, apical view of the head; 11) Holotype parameres." figureDoi="http://doi.org/10.5281/zenodo.4788723" httpUri="https://zenodo.org/record/4788723/files/figure.png" pageId="2" pageNumber="534">Fig. 10</figureCitation>
). Pronotum weakly convex (
<figureCitation id="A4F3F992FF862C1EFD23AB57FC85FBAC" box="[767,857,1097,1119]" captionStart="Figs" captionStartId="3.[197,236,1492,1511]" captionTargetBox="[177,1188,196,1474]" captionTargetId="figure-71@3.[175,1190,194,1476]" captionTargetPageId="3" captionText="Figs. 6–11. Podolasia parapilosa, holotype and paratype. 6) Holotype, dorsal habitus; 7) Holotype, oblique habitus; 8) Paratype, ventral habitus; 9) Holotype sclerotized genital capsule; 10) Holotype, apical view of the head; 11) Holotype parameres." figureDoi="http://doi.org/10.5281/zenodo.4788723" httpUri="https://zenodo.org/record/4788723/files/figure.png" pageId="2" pageNumber="534">Figs. 6–7</figureCitation>
), not declivous anteriorly, widest at middle, with marginal bead complete, thin laterally, not distinctly crenate near anterior angles; lacking distinct transverse groove behind anterior margin; lateral margin with dense, long setae; setae becoming short and sparse on posterior margin. Anterior angles slightly produced near eyes, acute; posterior angles obsolete; pronotal disc except near anterior and posterior margins with sparse, large punctures, punctures closer laterally than on disc, surface between punctures shiny; some lateral punctures with setae. Scutellum longer than wide, with bluntly rounded apex, surface smooth, impunctate. Elytron lacking impressed striae, 4 striae indicated by irregular rows of large punctures, some punctures with a fine, semi-erect seta; intervals between rows of punctures with irregular, sparse punctures lacking setae; interval punctures mixed fine to large; surface shiny; apex of elytron rounded near suture. Pygidium relatively short and broad, surface with scattered punctures, each with a long seta; surface shiny, feebly reticulate basally. Abdomen with ventral sutures mostly effaced (connate), only apical and basal segments with defined sutures. Protibia with 2 teeth. Metafemur oval, less than 2X long as wide. Metatibia elongate, about 2X longer than width at apex; apex with basotarsal flange narrowly produced in lateral view; slanted carina extending across 1/2 of outer face near apical third of tibia, outer surface without distinct tubercles.Tarsal claws simple. Each paramere of genitalia (
<figureCitation id="A4F3F992FF872C1FFCC2A966FCBDF97F" box="[798,865,1656,1677]" captionStart="Figs" captionStartId="3.[197,236,1492,1511]" captionTargetBox="[177,1188,196,1474]" captionTargetId="figure-71@3.[175,1190,194,1476]" captionTargetPageId="3" captionText="Figs. 6–11. Podolasia parapilosa, holotype and paratype. 6) Holotype, dorsal habitus; 7) Holotype, oblique habitus; 8) Paratype, ventral habitus; 9) Holotype sclerotized genital capsule; 10) Holotype, apical view of the head; 11) Holotype parameres." figureDoi="http://doi.org/10.5281/zenodo.4788723" httpUri="https://zenodo.org/record/4788723/files/figure.png" pageId="3" pageNumber="535">Fig. 11</figureCitation>
) straight externally with distinctive triangular expansion apically. Wings present.
</paragraph>
<caption id="68B7B59FFF872C1FFF19AACAFE5DF9E5" ID-DOI="http://doi.org/10.5281/zenodo.4788723" ID-Zenodo-Dep="4788723" httpUri="https://zenodo.org/record/4788723/files/figure.png" pageId="3" pageNumber="535" startId="3.[197,236,1492,1511]" targetBox="[177,1188,196,1474]" targetPageId="3">
<paragraph id="3C77E517FF872C1FFF19AACAFE5DF9E5" blockId="3.[173,1194,1491,1559]" pageId="3" pageNumber="535">
<emphasis id="0EBC3905FF872C1FFF19AACAFEF5FA15" bold="true" box="[197,297,1492,1511]" pageId="3" pageNumber="535">Figs. 6–11.</emphasis>
<taxonomicName id="FBC89E94FF872C1FFEE3AACDFE27FA14" authorityName="Smith and Paulsen" authorityYear="2017" box="[319,507,1491,1510]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="535" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF872C1FFEE3AACDFE27FA14" box="[319,507,1491,1510]" italics="true" pageId="3" pageNumber="535">Podolasia parapilosa</emphasis>
</taxonomicName>
, holotype and paratype.
<emphasis id="0EBC3905FF872C1FFD38AACAFD29FA15" bold="true" box="[740,757,1492,1511]" pageId="3" pageNumber="535">6)</emphasis>
Holotype, dorsal habitus;
<emphasis id="0EBC3905FF872C1FFC31AACAFC23FA15" bold="true" box="[1005,1023,1492,1511]" pageId="3" pageNumber="535">7)</emphasis>
Holotype, oblique habitus;
<emphasis id="0EBC3905FF872C1FFF2BAAF2FED4FA0D" bold="true" box="[247,264,1516,1535]" pageId="3" pageNumber="535">8)</emphasis>
Paratype, ventral habitus;
<emphasis id="0EBC3905FF872C1FFE28AAF2FDD9FA0D" bold="true" box="[500,517,1516,1535]" pageId="3" pageNumber="535">9)</emphasis>
Holotype sclerotized genital capsule;
<emphasis id="0EBC3905FF872C1FFC89AAF2FCADFA0D" bold="true" box="[853,881,1516,1535]" pageId="3" pageNumber="535">10)</emphasis>
Holotype, apical view of the head;
<emphasis id="0EBC3905FF872C1FFF71A91AFF1BF9E5" bold="true" box="[173,199,1540,1559]" pageId="3" pageNumber="535">11)</emphasis>
Holotype parameres.
</paragraph>
</caption>
<paragraph id="3C77E517FF872C1FFD07A9B3FC4BF930" blockId="3.[707,1193,1603,1730]" box="[731,919,1709,1730]" pageId="3" pageNumber="535">
<emphasis id="0EBC3905FF872C1FFD07A9B3FCF6F930" bold="true" box="[731,810,1709,1730]" pageId="3" pageNumber="535">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="3C77E517FF802C18FF73AFDCFEC7FEFE" blockId="4.[151,639,193,985]" pageId="4" pageNumber="536">
<emphasis id="0EBC3905FF802C18FF73AFDCFEC9FF25" bold="true" box="[175,277,194,215]" pageId="4" pageNumber="536">Variation.</emphasis>
Length
<quantity id="FB3048F2FF802C18FEB1AFDCFE3AFF25" box="[365,486,193,215]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="6.3" metricValueMin="4.5" pageId="4" pageNumber="536" unit="mm" value="5.4" valueMax="6.3" valueMin="4.5">4.5–6.3 mm</quantity>
, greatest width
<quantity id="FB3048F2FF802C18FF4BAFC2FEC9FF03" box="[151,277,220,242]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="4" pageNumber="536" unit="mm" value="2.05" valueMax="2.6" valueMin="1.5">1.5–2.6 mm</quantity>
. Smaller specimens lack a clypeal emargination.
</paragraph>
</subSubSection>
<subSubSection id="74D2B69CFF802C18FF73AE0FFF12FCA6" pageId="4" pageNumber="536" type="discussion">
<paragraph id="3C77E517FF802C18FF73AE0FFDD4FDC2" blockId="4.[151,639,193,985]" pageId="4" pageNumber="536">
<emphasis id="0EBC3905FF802C18FF73AE0FFECDFED4" bold="true" box="[175,273,273,294]" pageId="4" pageNumber="536">Remarks.</emphasis>
<taxonomicName id="FBC89E94FF802C18FECAAE0EFE3EFED4" authorityName="Smith and Paulsen" authorityYear="2017" box="[278,482,272,294]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF802C18FECAAE0EFE3EFED4" box="[278,482,272,294]" italics="true" pageId="4" pageNumber="536">Podolasia parapilosa</emphasis>
</taxonomicName>
keys to
<taxonomicName id="FBC89E94FF802C18FDF2AE0FFDA2FED4" authorityName="Howden" authorityYear="1954" box="[558,638,272,294]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FDF2AE0FFDA2FED4" box="[558,638,272,294]" italics="true" pageId="4" pageNumber="536">P. pilosa</emphasis>
</taxonomicName>
in
<taxonomicName id="FBC89E94FF802C18FF69AE32FE8FFEB3" authority="Smith &amp; Paulsen, 2017" authorityName="Smith &amp; Paulsen" authorityYear="2017" box="[181,339,300,321]" class="Insecta" family="Scarabaeidae" genus="Howden" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="genus" status="gen. nov.">
<bibRefCitation id="585998E6FF802C18FF69AE32FE8FFEB3" author="Howden, H. F." box="[181,339,300,321]" pageId="4" pageNumber="536" pagination="275 - 277" refId="ref2360" refString="Howden, H. F. 1997. Podolasiini Howden, new tribe, When describing the tribe Podolasiini, Howden and a revision of the included genera, Podolasia (1997) provided a list of other &quot; primitive &quot; melo- Harold and Podostena Howden, new genus (Colonthine taxa that shared some characters. One leoptera: Scarabaeidae: Melolonthinae). The Coadditional character that we observed and warrants leopterists Bulletin 51: 223 - 255. mention is the sclerotized male genital capsule (Fig. Howden, H. F. 1998. More on Podolasia Harold: a new 9). This structure has two apical knobs with forward species and notes on another (Coleoptera: Scarprojecting setae, which is remarkable for a taxon in abaeidae: Melolonthinae). The Coleopterists Bulthe phytophagous scarab clade (Smith et al. 2006). letin 52: 275 - 277." type="journal article" year="1997">Howden (1997)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="FBC89E94FF802C18FEBDAE35FDE8FEB3" authorityName="Smith and Paulsen" authorityYear="2017" box="[353,564,299,321]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF802C18FEBDAE35FDE8FEB3" box="[353,564,299,321]" italics="true" pageId="4" pageNumber="536">Podolasia parapilosa</emphasis>
</taxonomicName>
is distinguished by the clypeus being longer than wide (wider than long in
<taxonomicName id="FBC89E94FF802C18FE81AE7FFE6DFE84" authorityName="Howden" authorityYear="1954" box="[349,433,352,374]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FE81AE7FFEB5FE84" box="[349,361,353,374]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FEABAE7EFE6DFE84" box="[375,433,352,374]" italics="true" pageId="4" pageNumber="536">pilosa</emphasis>
</taxonomicName>
) and usually weakly emarginate (never emarginate in
<taxonomicName id="FBC89E94FF802C18FE31AE62FD9AFE63" authorityName="Howden" authorityYear="1954" box="[493,582,379,401]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FE31AE62FE25FE63" box="[493,505,380,401]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FDD0AE65FD9AFE63" box="[524,582,379,401]" italics="true" pageId="4" pageNumber="536">pilosa</emphasis>
</taxonomicName>
), the metatibiae being 2–3X as thick at the apex compared to the base (at least 3X thicker in
<taxonomicName id="FBC89E94FF802C18FE23AEAFFD93FE34" authorityName="Howden" authorityYear="1954" box="[511,591,432,454]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FE23AEAFFDD7FE34" box="[511,523,433,454]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FDC9AEAEFD93FE34" box="[533,591,432,454]" italics="true" pageId="4" pageNumber="536">pilosa</emphasis>
</taxonomicName>
), the straight outer edges of the parameres (slightly bowed medially in
<taxonomicName id="FBC89E94FF802C18FE90AEF8FE41FE09" authorityName="Howden" authorityYear="1954" box="[332,413,485,507]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FE90AEF8FE84FE09" box="[332,344,486,507]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FEBFAEFBFE41FE09" box="[355,413,485,507]" italics="true" pageId="4" pageNumber="536">pilosa</emphasis>
</taxonomicName>
), and the distribution in
<collectingRegion id="FE0C2BF5FF802C18FF4BAD1EFF33FDE7" box="[151,239,512,533]" country="Mexico" name="Coahuila de Zaragoza" pageId="4" pageNumber="536">Coahuila</collectingRegion>
,
<collectingCountry id="44DFA587FF802C18FF21AD1EFE99FDE7" box="[253,325,512,533]" name="Mexico" pageId="4" pageNumber="536">Mexico</collectingCountry>
(New
<collectingRegion id="FE0C2BF5FF802C18FE51AD1EFE04FDE7" box="[397,472,512,533]" country="Mexico" name="Mexico" pageId="4" pageNumber="536">Mexico</collectingRegion>
,
<collectingRegion id="FE0C2BF5FF802C18FE39AD1EFD93FDE7" box="[485,591,512,533]" country="Mexico" name="Chihuahua" pageId="4" pageNumber="536">Chihuahua</collectingRegion>
, and bordering areas of Texas in
<taxonomicName id="FBC89E94FF802C18FE7BAD05FE27FDC2" authorityName="Howden" authorityYear="1954" box="[423,507,538,560]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FE7BAD05FE6FFDC2" box="[423,435,539,560]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FE1DAD04FE27FDC2" box="[449,507,538,560]" italics="true" pageId="4" pageNumber="536">pilosa</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3C77E517FF802C18FF73AD2BFF12FCA6" blockId="4.[151,639,193,985]" pageId="4" pageNumber="536">
When describing the tribe Podolasiini, Howden (1997) provided a list of other “primitive” melolonthine taxa that shared some characters. One additional character that we observed and warrants mention is the sclerotized male genital capsule (Fig. 9). This structure has two apical knobs with forward projecting setae, which is remarkable for a taxon in the phytophagous scarab clade (Smith
<emphasis id="0EBC3905FF802C18FDD6ADEEFDE6FCF7" box="[522,570,751,773]" italics="true" pageId="4" pageNumber="536">et al.</emphasis>
2006).
<journalOrPublisher id="E609EFDBFF802C18FF4BAC14FDA1FCED" box="[151,637,778,799]" pageId="4" pageNumber="536">The vast majority of phytophagous scarabs have a</journalOrPublisher>
soft genital capsule without hard sclerotized parts or setae.
</paragraph>
</subSubSection>
<subSubSection id="74D2B69CFF802C18FF73AC44FF35FC2B" pageId="4" pageNumber="536" type="etymology">
<paragraph id="3C77E517FF802C18FF73AC44FF35FC2B" blockId="4.[151,639,193,985]" pageId="4" pageNumber="536">
<emphasis id="0EBC3905FF802C18FF73AC44FEFEFC9D" bold="true" box="[175,290,858,879]" pageId="4" pageNumber="536">Etymology.</emphasis>
We name this species
<taxonomicName id="FBC89E94FF802C18FE25AC44FDA4FC9D" authorityName="Smith and Paulsen" authorityYear="2017" box="[505,632,857,879]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="parapilosa">
<emphasis id="0EBC3905FF802C18FE25AC44FDD9FC9D" box="[505,517,858,879]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FDCDAC47FDA4FC9D" box="[529,632,857,879]" italics="true" pageId="4" pageNumber="536">parapilosa</emphasis>
</taxonomicName>
, meaning “close to
<taxonomicName id="FBC89E94FF802C18FE91AC6AFE7CFC78" authorityName="Howden" authorityYear="1954" box="[333,416,884,906]" class="Insecta" family="Melolonthidae" genus="Podolasia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="536" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0EBC3905FF802C18FE91AC6AFE85FC7B" box="[333,345,884,905]" italics="true" pageId="4" pageNumber="536">P</emphasis>
.
<emphasis id="0EBC3905FF802C18FEBAAC6AFE7CFC78" box="[358,416,884,906]" italics="true" pageId="4" pageNumber="536">pilosa</emphasis>
</taxonomicName>
”, to indicate the many similarities between these two species. This name should be treated as an adjective in the nominative singular.
</paragraph>
</subSubSection>
</treatment>
</document>
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