149 lines
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149 lines
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<document id="3721E5B280F662BF97691C840D84E604" ID-DOI="10.11646/zootaxa.5375.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184367" ID-ZooBank="FA90BCF7-C67D-44FA-B6B8-3BEC7531F7DB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1700643407716" checkinUser="plazi" docAuthor="Deschodt, Christian M. & Sole, Catherine L." docDate="2023" docId="810E963FFFC2FFE6FF47FF05D5EAA930" docLanguage="en" docName="zootaxa.5375.2.9.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hathor spinosa Deschodt 2023, new species" docType="treatment" docVersion="3" lastPageNumber="282" masterDocId="7D37EE47FFC1FFE5FFD0FF92D700AE6B" masterDocTitle="A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa" masterLastPageNumber="284" masterPageNumber="279" pageNumber="282" updateTime="1700730964351" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:title id="B6CA70E601646B5F184AABF353379343">A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa</mods:title>
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<mods:namePart id="EF5DABD2640931CA2F83C74E6BD106C5">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="3C5BC9446CCFEEA166F961F791CEFA4F">Sole, Catherine L.</mods:namePart>
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<mods:title id="87712EB93861A42280D04601DFD15639">Zootaxa</mods:title>
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<mods:part id="ECF6258E0CF66DAD06EEC0C739824519">
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<mods:date id="1A21E02FC46A4BD0614C931FD47AEA8B">2023</mods:date>
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<mods:number id="5A9ED2A166DEC3C540E5A45FB15840D9">2023-11-22</mods:number>
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<mods:number id="5070211267A4EF4066A152ADA120ADA4">2</mods:number>
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<mods:identifier id="05565D45AB8186C9D89A56E1644246EE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="060BF3EF075366D42D175E102B80B022" type="Zenodo-Dep">10184367</mods:identifier>
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<treatment id="810E963FFFC2FFE6FF47FF05D5EAA930" LSID="urn:lsid:plazi:treatment:810E963FFFC2FFE6FF47FF05D5EAA930" httpUri="http://treatment.plazi.org/id/810E963FFFC2FFE6FF47FF05D5EAA930" lastPageNumber="282" pageId="3" pageNumber="282">
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<subSubSection id="41BD74A2FFC2FFE6FF47FF05D56BAEDA" box="[151,619,151,178]" pageId="3" pageNumber="282" type="nomenclature">
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<paragraph id="09182729FFC2FFE6FF47FF05D56BAEDA" blockId="3.[151,619,151,213]" box="[151,619,151,178]" pageId="3" pageNumber="282">
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<heading id="52509045FFC2FFE6FF47FF05D56BAEDA" bold="true" box="[151,619,151,178]" fontSize="11" level="1" pageId="3" pageNumber="282" reason="1">
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<emphasis id="3BD3FB3BFFC2FFE6FF47FF05D56BAEDA" bold="true" box="[151,619,151,178]" pageId="3" pageNumber="282">
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<taxonomicName id="CEA75CAAFFC2FFE6FF47FF05D6CFAEDA" authority="Deschodt" authorityName="Deschodt" authorityYear="2023" box="[151,463,151,178]" class="Insecta" family="Scarabaeidae" genus="Hathor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
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<emphasis id="3BD3FB3BFFC2FFE6FF47FF05D652AEDA" bold="true" box="[151,338,151,178]" italics="true" pageId="3" pageNumber="282">Hathor spinosa</emphasis>
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Deschodt
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</taxonomicName>
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,
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<taxonomicNameLabel id="20E04640FFC2FFE6FE0CFF05D56BAEDA" box="[476,619,151,177]" pageId="3" pageNumber="282" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="41BD74A2FFC2FFE6FF47FF29D5EAA930" pageId="3" pageNumber="282" type="description">
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<paragraph id="09182729FFC2FFE6FF47FF29D683AEBE" blockId="3.[151,619,151,213]" box="[151,387,187,213]" pageId="3" pageNumber="282">
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<figureCitation id="919C3BACFFC2FFE6FF47FF29D639AEBE" box="[151,313,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1563,1587]" captionTargetBox="[158,1435,195,1540]" captionTargetId="figure-25@2.[151,1435,181,1540]" captionTargetPageId="2" captionText="FIGURE 1. Hathor spinosa Deschodt, new genus and species. (A) habitus in dorsal view showing the reduced fourth denticle of the protibiae; (B) habitus in ventral view; (C) habitus in lateral view showing the spine on the pronotum and; (D) frontal view showing the characteristic horns on the vertex and the triangular spine-like process situated anteromedially on the prothorax." figureDoi="http://doi.org/10.5281/zenodo.10184369" httpUri="https://zenodo.org/record/10184369/files/figure.png" pageId="3" pageNumber="282">Figures 1A–D</figureCitation>
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,
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<figureCitation id="919C3BACFFC2FFE6FE95FF29D669AEBE" box="[325,361,187,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1570,1594]" captionTargetBox="[179,1417,777,1547]" captionTargetId="figure-245@3.[151,1435,777,1547]" captionTargetPageId="3" captionText="FIGURE 2. Head in ventral view and number of antennal segments. (A) Hathor spinosa Deschodt, new genus and species with eight segments; (B) Onthophagus suffuses Klug, 1855 with nine segments." figureDoi="http://doi.org/10.5281/zenodo.10184373" httpUri="https://zenodo.org/record/10184373/files/figure.png" pageId="3" pageNumber="282">2A</figureCitation>
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,
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<figureCitation id="919C3BACFFC2FFE6FEA5FF29D683AEBE" box="[373,387,187,213]" captionStart="FIGURE 3" captionStartId="4.[152,255,1100,1124]" captionTargetBox="[151,1436,181,1076]" captionTargetId="figure-25@4.[151,1436,181,1076]" captionTargetPageId="4" captionText="FIGURE 3. Map of South Africa showing the type locality of Hathor spinosa Deschodt, new genus and species (red dot west of Pretoria)." figureDoi="http://doi.org/10.5281/zenodo.10184375" httpUri="https://zenodo.org/record/10184375/files/figure.png" pageId="3" pageNumber="282">3</figureCitation>
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</paragraph>
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<paragraph id="09182729FFC2FFE6FF47FE96D3FCAF76" blockId="3.[151,1437,259,754]" box="[151,1276,259,286]" pageId="3" pageNumber="282">
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Type material.
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<materialsCitation id="B9CF2D74FFC2FFE6FE90FE91D3FCAF76" box="[320,1276,259,286]" collectingDate="2023-01-07" collectionCode="TMSA" collectorName="Schurveberg & C. Deschodt" pageId="3" pageNumber="282" specimenCount="1" typeStatus="holotype">
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<typeStatus id="D61C998BFFC2FFE6FE90FE91D6ABAF76" box="[320,427,259,285]" pageId="3" pageNumber="282" type="holotype">Holotype</typeStatus>
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, female:
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<collectorName id="A45242FFFFC2FFE6FDC1FE91D59EAF76" box="[529,670,259,285]" pageId="3" pageNumber="282">Schurveberg</collectorName>
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| -25.768° 27.964° |
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<date id="7D1901E9FFC2FFE6FC56FE91D4EFAF76" box="[902,1007,259,285]" pageId="3" pageNumber="282" value="2023-01-07">
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<collectingDate id="6D5DF801FFC2FFE6FC56FE91D4EFAF76" box="[902,1007,259,285]" pageId="3" pageNumber="282" value="2023-01-07">07.i.2023</collectingDate>
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</date>
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<collectorName id="A45242FFFFC2FFE6FBD3FE91D38EAF76" box="[1027,1166,259,285]" pageId="3" pageNumber="282">C. Deschodt</collectorName>
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(
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<collectionCode id="6FB6BFECFFC2FFE6FB4CFE91D3F0AF76" box="[1180,1264,259,285]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="282">TMSA</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="09182729FFC2FFE6FF17FEB5D530AC99" blockId="3.[151,1437,259,754]" pageId="3" pageNumber="282">
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Description of the
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<typeStatus id="D61C998BFFC2FFE6FE74FEB5D506AF2A" box="[420,518,295,321]" pageId="3" pageNumber="282" type="holotype">holotype</typeStatus>
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(female):
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<emphasis id="3BD3FB3BFFC2FFE6FD52FEB5D5BFAF2A" box="[642,703,295,321]" italics="true" pageId="3" pageNumber="282">Head</emphasis>
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: (
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<figureCitation id="919C3BACFFC2FFE6FD0AFEB5D4ABAF29" box="[730,939,295,322]" captionStart="FIGURE 1" captionStartId="2.[152,255,1563,1587]" captionTargetBox="[158,1435,195,1540]" captionTargetId="figure-25@2.[151,1435,181,1540]" captionTargetPageId="2" captionText="FIGURE 1. Hathor spinosa Deschodt, new genus and species. (A) habitus in dorsal view showing the reduced fourth denticle of the protibiae; (B) habitus in ventral view; (C) habitus in lateral view showing the spine on the pronotum and; (D) frontal view showing the characteristic horns on the vertex and the triangular spine-like process situated anteromedially on the prothorax." figureDoi="http://doi.org/10.5281/zenodo.10184369" httpUri="https://zenodo.org/record/10184369/files/figure.png" pageId="3" pageNumber="282">Fig. 1A, C and D</figureCitation>
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and
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<figureCitation id="919C3BACFFC2FFE6FC39FEB5D30DAF2A" box="[1001,1037,295,321]" captionStart="FIGURE 2" captionStartId="3.[152,255,1570,1594]" captionTargetBox="[179,1417,777,1547]" captionTargetId="figure-245@3.[151,1435,777,1547]" captionTargetPageId="3" captionText="FIGURE 2. Head in ventral view and number of antennal segments. (A) Hathor spinosa Deschodt, new genus and species with eight segments; (B) Onthophagus suffuses Klug, 1855 with nine segments." figureDoi="http://doi.org/10.5281/zenodo.10184373" httpUri="https://zenodo.org/record/10184373/files/figure.png" pageId="3" pageNumber="282">2A</figureCitation>
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) Clypeus extended and narrowing anteriorly, upturned at end, shallowly sinuate; anterior surface of frontoclypeal suture smooth and punctate with associated setae; frontoclypeal suture distinct medially, effaced at the sides, curving anteriorly; vertex with distinct and setate horns.
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<emphasis id="3BD3FB3BFFC2FFE6FE88FE01D6CAAFC6" box="[344,458,403,429]" italics="true" pageId="3" pageNumber="282">Prothorax</emphasis>
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: Dorsal surface punctate with short associated setae; lateral surface with small granules and short associated setae; anterior vertical surface smooth, prothoracic carina straight.; triangular spine-like process present medially.
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<emphasis id="3BD3FB3BFFC2FFE6FEB0FE4ED695AF9E" box="[352,405,476,501]" italics="true" pageId="3" pageNumber="282">Legs</emphasis>
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: Protibiae with four teeth on outside, fourth tooth reduced, small denticle present between second and third teeth and between third and fourth teeth; tibiae ending at a slanted angle; first article of the protarsus longer than the next three combined, last protarsus almost as long as the previous three segments combined; meso and metalegs widening distally.
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<emphasis id="3BD3FB3BFFC2FFE6FD98FDDAD590AC0A" box="[584,656,584,609]" italics="true" pageId="3" pageNumber="282">Venter</emphasis>
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. (
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<figureCitation id="919C3BACFFC2FFE6FD77FDD5D5FDAC0A" box="[679,765,583,609]" captionStart="FIGURE 1" captionStartId="2.[152,255,1563,1587]" captionTargetBox="[158,1435,195,1540]" captionTargetId="figure-25@2.[151,1435,181,1540]" captionTargetPageId="2" captionText="FIGURE 1. Hathor spinosa Deschodt, new genus and species. (A) habitus in dorsal view showing the reduced fourth denticle of the protibiae; (B) habitus in ventral view; (C) habitus in lateral view showing the spine on the pronotum and; (D) frontal view showing the characteristic horns on the vertex and the triangular spine-like process situated anteromedially on the prothorax." figureDoi="http://doi.org/10.5281/zenodo.10184369" httpUri="https://zenodo.org/record/10184369/files/figure.png" pageId="3" pageNumber="282">Fig. 1B</figureCitation>
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) Mesometasternal suture distinct and very slightly sinuate; mesosternum narrow, punctate with small punctures that are closer together than their diameters; metasternum surface smooth with small punctures that are about three times their diameters apart.
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<emphasis id="3BD3FB3BFFC2FFE6FB9DFD1DD3B7ACC2" box="[1101,1207,655,681]" italics="true" pageId="3" pageNumber="282">Abdomen</emphasis>
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: surface of ventrites smooth with short setae.
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<taxonomicName id="CEA75CAAFFC2FFE6FE6AFD21D525ACA6" box="[442,549,691,717]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="282" phylum="Chordata" rank="genus">
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<emphasis id="3BD3FB3BFFC2FFE6FE6AFD21D525ACA6" box="[442,549,691,717]" italics="true" pageId="3" pageNumber="282">Pygidium</emphasis>
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</taxonomicName>
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: Surface smooth with medium punctures about one puncture diameter apart; punctures with thin associated setae.
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</paragraph>
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<caption id="5DD877A1FFC2FFE6FF48F9B0D48DA834" ID-DOI="http://doi.org/10.5281/zenodo.10184373" ID-Zenodo-Dep="10184373" httpUri="https://zenodo.org/record/10184373/files/figure.png" pageId="3" pageNumber="282" startId="3.[152,255,1570,1594]" targetBox="[179,1417,777,1547]" targetPageId="3" targetType="figure">
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<paragraph id="09182729FFC2FFE6FF48F9B0D48DA834" blockId="3.[151,1436,1570,1631]" pageId="3" pageNumber="282">
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<emphasis id="3BD3FB3BFFC2FFE6FF48F9B0D617A851" bold="true" box="[152,279,1570,1594]" pageId="3" pageNumber="282">FIGURE 2.</emphasis>
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Head in ventral view and number of antennal segments. (A)
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<taxonomicName id="CEA75CAAFFC2FFE6FCA9F9B0D37DA851" authority="Deschodt" authorityName="Deschodt" authorityYear="2023" box="[889,1149,1570,1594]" class="Insecta" genus="Hathor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="spinosa" status="new genus and species">
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<emphasis id="3BD3FB3BFFC2FFE6FCA9F9B0D315A851" box="[889,1045,1570,1594]" italics="true" pageId="3" pageNumber="282">Hathor spinosa</emphasis>
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Deschodt
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</taxonomicName>
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,
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<taxonomicNameLabel id="20E04640FFC2FFE6FB56F9B1D26AA851" box="[1158,1386,1570,1595]" pageId="3" pageNumber="282" rank="species">new genus and species</taxonomicNameLabel>
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with eight segments; (B)
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<taxonomicName id="CEA75CAAFFC2FFE6FEB4F9D4D5BCA835" authority="Klug, 1855" authorityName="Klug" authorityYear="1855" box="[356,700,1606,1631]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="suffuses">
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<emphasis id="3BD3FB3BFFC2FFE6FEB4F9D4D543A834" box="[356,579,1606,1631]" italics="true" pageId="3" pageNumber="282">Onthophagus suffuses</emphasis>
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Klug, 1855
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</taxonomicName>
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with nine segments.
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</paragraph>
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</caption>
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<paragraph id="09182729FFC2FFE6FF17F923D6DDA884" blockId="3.[151,1436,1713,1883]" pageId="3" pageNumber="282">Etymology. The specific epithet is feminine in gender and refers to the triangular spine-like process that is seen on the front of the prothorax.</paragraph>
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<paragraph id="09182729FFC2FFE6FF17F96BD5EAA930" blockId="3.[151,1436,1713,1883]" pageId="3" pageNumber="282">
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Remarks. As it was found in putative association with the very widespread and commonly found ant species
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<taxonomicName id="CEA75CAAFFC2FFE6FF47F88FD69DA95C" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[151,413,1821,1847]" class="Insecta" family="Formicidae" genus="Anoplolepis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="custodiens">
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<emphasis id="3BD3FB3BFFC2FFE6FF47F88FD69DA95C" box="[151,413,1821,1847]" italics="true" pageId="3" pageNumber="282">Anoplolepis custodiens</emphasis>
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</taxonomicName>
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, the potential distribution of
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<taxonomicName id="CEA75CAAFFC2FFE6FD22F88FD319A95C" authority="Deschodt" authorityName="Deschodt" authorityYear="2023" box="[754,1049,1821,1847]" class="Insecta" genus="Hathor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="spinosa" status="new genus and species">
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<emphasis id="3BD3FB3BFFC2FFE6FD22F88FD4A2A95D" box="[754,930,1821,1847]" italics="true" pageId="3" pageNumber="282">Hathor spinosa</emphasis>
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Deschodt
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</taxonomicName>
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,
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<taxonomicNameLabel id="20E04640FFC2FFE6FBF6F88FD22DA95C" box="[1062,1325,1821,1847]" pageId="3" pageNumber="282" rank="species">new genus and species</taxonomicNameLabel>
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might be most of Southern Africa and further to the north east.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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