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<emphasis id="B91FEAECFFB4FF85FF4BFBB7FED8E676" bold="true" box="[151,365,1066,1092]" italics="true" pageId="22" pageNumber="23">Trionyx triunguis</emphasis>
(
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locality: “In Nilo rarior” (
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, 
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<treatmentCitationGroup id="AB7B11D0FFB4FF85FF4BFB10FDFEE697" box="[151,587,1165,1189]" pageId="22" pageNumber="23">
<treatmentCitation id="0ACA10EFFFB4FF85FF4BFB10FE5FE697" author="Suckow, G. A." box="[151,490,1165,1189]" page="37" pageId="22" pageNumber="23" year="1798">
<taxonomicName id="4C6B4D7DFFB4FF85FF4BFB10FE5FE697" authority="Suckow, 1798: 37" authorityName="Suckow" authorityPageNumber="37" authorityYear="1798" box="[151,490,1165,1189]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="striata">
<emphasis id="B91FEAECFFB4FF85FF4BFB10FE98E697" box="[151,301,1165,1189]" italics="true" pageId="22" pageNumber="23">Testudo striata</emphasis>
<bibRefCitation id="EFFA4B0FFFB4FF85FEE8FB10FE5FE697" author="Suckow, G. A." box="[308,490,1165,1189]" pageId="22" pageNumber="23" refId="ref90457" refString="Suckow, G. A. (1798) Anfangsgrunde der Theoretischen und Angewandten Naturgeschichte der Thiere. Dritter Theil. Von den Amphibien. Weidmannischen Buchhandlung, Leipzig, 298 pp. https: // doi. org / 10.5962 / bhl. title. 62582" type="book" year="1798">Suckow, 1798: 37</bibRefCitation>
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(partim).
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<taxonomicName id="4C6B4D7DFFB4FF85FF4BFB31FD9FE6F1" authority="Geoffroy, 1809: 116" authorityName="Geoffroy" authorityPageNumber="116" authorityYear="1809" box="[151,554,1195,1220]" class="Reptilia" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="aegytiacus">
<emphasis id="B91FEAECFFB4FF85FF4BFB31FEE2E6F1" box="[151,343,1196,1219]" italics="true" pageId="22" pageNumber="23">Trionyx aegytiacus</emphasis>
Geoffroy, 1809: 116
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(nomen substitutum pro 
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<emphasis id="B91FEAECFFB4FF85FCF5FB31FC6CE6F1" box="[809,985,1196,1220]" italics="true" pageId="22" pageNumber="23">Testudo triunguis</emphasis>
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).
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<treatmentCitationGroup id="AB7B11D0FFB4FF85FF4BFB56FC29E6D0" box="[151,924,1226,1250]" pageId="22" pageNumber="23">
<treatmentCitation id="0ACA10EFFFB4FF85FF4BFB56FE6AE6D0" author="Gray, J. E." box="[151,479,1226,1250]" page="46" pageId="22" pageNumber="23" year="1831">
<taxonomicName id="4C6B4D7DFFB4FF85FF4BFB56FE6AE6D0" ID-CoL="58WJY" authority="Gray, 1831: 46" authorityName="Gray" authorityPageNumber="46" authorityYear="1831" box="[151,479,1226,1250]" class="Reptilia" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="niloticus">
<emphasis id="B91FEAECFFB4FF85FF4BFB56FEF7E6D0" box="[151,322,1226,1250]" italics="true" pageId="22" pageNumber="23">Trionyx niloticus</emphasis>
<bibRefCitation id="EFFA4B0FFFB4FF85FE95FB57FE6AE6D0" author="Gray, J. E." box="[329,479,1226,1250]" pageId="22" pageNumber="23" pagination="7" refId="ref78140" refString="Gray, J. E. (1831) Description of a new species of chamaeleon discovered by Capt. Owen in Africa. Zoological Miscellany, 1, 7." type="journal article" year="1831">Gray, 1831: 46</bibRefCitation>
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(nomen substitutum pro 
<taxonomicName id="4C6B4D7DFFB4FF85FD02FB57FC3AE6D0" authorityName="Forskal" authorityYear="1775" box="[734,911,1226,1250]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="triunguis">
<emphasis id="B91FEAECFFB4FF85FD02FB57FC3AE6D0" box="[734,911,1226,1250]" italics="true" pageId="22" pageNumber="23">Testudo triunguis</emphasis>
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).
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<taxonomicName id="4C6B4D7DFFB4FF85FF4BFB77FDBFE733" authority="Cuvier, 1831: 18" authorityName="Cuvier" authorityPageNumber="18" authorityYear="1831" box="[151,522,1257,1281]" class="Reptilia" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="aegytiacus">
<emphasis id="B91FEAECFFB4FF85FF4BFB77FEE2E733" box="[151,343,1258,1281]" italics="true" pageId="22" pageNumber="23">Trionyx aegytiacus</emphasis>
Cuvier, 1831: 18
</taxonomicName>
. 
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locality: 
<collectingCountry id="F37C766EFFB4FF85FD7AFB74FD53E733" box="[678,742,1257,1281]" name="Egypt" pageId="22" pageNumber="23">Egypt</collectingCountry>
.
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<taxonomicName id="4C6B4D7DFFB4FF85FF4BFA95FE05E712" authority="Bell, 1837" authorityName="Bell" authorityYear="1837" box="[151,432,1288,1312]" class="Reptilia" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="labiatus">
<emphasis id="B91FEAECFFB4FF85FF4BFA95FE88E712" box="[151,317,1288,1312]" italics="true" pageId="22" pageNumber="23">Trionyx labiatus</emphasis>
Bell, 1837
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: unnumbered. 
<typeStatus id="54D0885CFFB4FF85FD99FA95FDC2E712" box="[581,631,1288,1312]" pageId="22" pageNumber="23">Type</typeStatus>
locality: 
<collectingCountry id="F37C766EFFB4FF85FD0BFA95FCE9E72D" box="[727,860,1288,1312]" name="Sierra Leone" pageId="22" pageNumber="23">Sierra Leone</collectingCountry>
.
</paragraph>
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<treatmentCitationGroup id="AB7B11D0FFB4FF85FF4BFABAFD48E70C" box="[151,765,1318,1343]" pageId="22" pageNumber="23">
<treatmentCitation id="0ACA10EFFFB4FF85FF4BFABAFDAEE70C" author="Hallowell, E." box="[151,539,1318,1342]" page="120" pageId="22" pageNumber="23" year="1844">
<taxonomicName id="4C6B4D7DFFB4FF85FF4BFABAFDAEE70C" ID-CoL="7CSZW" authority="Hallowell, 1844: 120" authorityName="Hallowell" authorityPageNumber="120" authorityYear="1844" box="[151,539,1318,1342]" class="Reptilia" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="mortoni">
<emphasis id="B91FEAECFFB4FF85FF4BFABAFE8EE70C" box="[151,315,1319,1342]" italics="true" pageId="22" pageNumber="23">Trionyx mortoni</emphasis>
<bibRefCitation id="EFFA4B0FFFB4FF85FE9DFABBFDAEE70C" author="Hallowell, E." box="[321,539,1318,1342]" pageId="22" pageNumber="23" pagination="118 - 120" refId="ref79559" refString="Hallowell, E. (1844) Description of new species of reptiles from Africa. Proceedings of the Academy of Natural Sciences of Philadelphia, 2, 118 - 120." type="journal article" year="1844">Hallowell, 1844: 120</bibRefCitation>
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. 
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locality: Africa.
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<treatmentCitationGroup id="AB7B11D0FFB4FF85FF4BFAD8FE24E74E" pageId="22" pageNumber="23">
<treatmentCitation id="0ACA10EFFFB4FF85FF4BFAD8FD99E76F" author="Cope, E. D." box="[151,556,1349,1373]" page="295" pageId="22" pageNumber="23" year="1860">
<taxonomicName id="4C6B4D7DFFB4FF85FF4BFAD8FD99E76F" ID-CoL="HH9M" authority="Cope, 1860 a: 295" authorityName="Cope" authorityPageNumber="295" authorityYear="1860" box="[151,556,1349,1373]" class="Reptilia" family="Trionychidae" genus="Aspidonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="aspilus">
<emphasis id="B91FEAECFFB4FF85FF4BFAD8FED9E76F" box="[151,364,1349,1373]" italics="true" pageId="22" pageNumber="23">Aspidonectes aspilus</emphasis>
<bibRefCitation id="EFFA4B0FFFB4FF85FEAFFAD8FD99E76F" author="Cope, E. D." box="[371,556,1349,1373]" pageId="22" pageNumber="23" pagination="294 - 297" refId="ref74428" refString="Cope, E. D. (1860 a) Notes and descriptions of foreign reptiles. Proceedings of the Academy of Natural Sciences of Philadelphia, 11, 294 - 297." type="journal article" year="1860">Cope, 1860a: 295</bibRefCitation>
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. 
<typeStatus id="54D0885CFFB4FF85FDEAFAD8FDDDE76F" box="[566,616,1349,1373]" pageId="22" pageNumber="23">Type</typeStatus>
locality: “Rembo and Ovenga rivers, …Fernando Vas, Equatorial West Africa”. 
<location id="8EB46025FFB4FF85FF1BFAF9FE89E749" LSID="urn:lsid:plazi:treatment:03C287E8FFB4FF85FF4BFBB7FB6DE513:8EB46025FFB4FF85FF1BFAF9FE89E749" box="[199,316,1380,1403]" country="Gabon" name="Fernan Vaz" pageId="22" pageNumber="23">Fernan Vaz</location>
, 
<collectingCountry id="F37C766EFFB4FF85FE9AFAF9FE38E74E" box="[326,397,1380,1404]" name="Gabon" pageId="22" pageNumber="23">Gabon</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFB4FF85FF4BFA1FFC71E7A8" blockId="22.[151,1437,1134,1465]" box="[151,964,1410,1435]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB7B11D0FFB4FF85FF4BFA1FFC71E7A8" box="[151,964,1410,1435]" pageId="22" pageNumber="23">
<treatmentCitation id="0ACA10EFFFB4FF85FF4BFA1FFE53E7A8" author="Gray, J. E." box="[151,486,1410,1434]" page="219" pageId="22" pageNumber="23" year="1869">
<taxonomicName id="4C6B4D7DFFB4FF85FF4BFA1FFE53E7A8" ID-CoL="6JHF5" authority="Gray, 1869: 219" authorityName="Gray" authorityPageNumber="219" authorityYear="1869" box="[151,486,1410,1434]" class="Reptilia" family="Trionychidae" genus="Fordia" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="africana">
<emphasis id="B91FEAECFFB4FF85FF4BFA1FFE8CE7A8" box="[151,313,1410,1434]" italics="true" pageId="22" pageNumber="23">Fordia africana</emphasis>
<bibRefCitation id="EFFA4B0FFFB4FF85FEE3FA1FFE53E7A8" author="Gray, J. E." box="[319,486,1410,1434]" pageId="22" pageNumber="23" pagination="165 - 225" refId="ref78421" refString="Gray, J. E. (1869) Notes on the families and genera of tortoises (Testudinata), and on the characters afforded by the study of their skulls. Proceedings of the Zoological Society of London, 1869, 165 - 225. https: // doi. org / 10.1111 / j. 1469 - 7998.1869. tb 07312. x" type="journal article" year="1869">Gray, 1869: 219</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="54D0885CFFB4FF85FE33FA1EFD94E7A9" box="[495,545,1411,1435]" pageId="22" pageNumber="23">Type</typeStatus>
locality: 
<location id="8EB46025FFB4FF85FD5DFA1FFD5EE7A8" LSID="urn:lsid:plazi:treatment:03C287E8FFB4FF85FF4BFBB7FB6DE513:8EB46025FFB4FF85FD5DFA1FFD5EE7A8" box="[641,747,1410,1434]" country="Sudan" name="Nile River" pageId="22" pageNumber="23" stateProvince="Al Khartoum">Nile River</location>
at 
<collectingRegion id="49AFF81CFFB4FF85FCD6FA1FFCC6E7A8" box="[778,883,1410,1434]" country="Sudan" name="Al Khartoum" pageId="22" pageNumber="23">Khartoum</collectingRegion>
, 
<collectingCountry id="F37C766EFFB4FF85FCA2FA1FFC74E7A8" box="[894,961,1410,1434]" name="Sudan" pageId="22" pageNumber="23">Sudan</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFB4FF85FF4BFA3FFB6CE78B" blockId="22.[151,1437,1134,1465]" box="[151,1241,1441,1465]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB7B11D0FFB4FF85FF4BFA3FFB6CE78B" box="[151,1241,1441,1465]" pageId="22" pageNumber="23">
<treatmentCitation id="0ACA10EFFFB4FF85FF4BFA3FFD0DE78B" author="Deraniyagala, P." box="[151,696,1441,1465]" page="30" pageId="22" pageNumber="23" year="1948">
<taxonomicName id="4C6B4D7DFFB4FF85FF4BFA3FFD0DE78B" authority="Deraniyagala, 1948: 30" authorityName="Deraniyagala" authorityPageNumber="30" authorityYear="1948" box="[151,696,1441,1465]" class="Reptilia" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Testudines" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="triunguis" subSpecies="rudolfianus">
<emphasis id="B91FEAECFFB4FF85FF4BFA3FFE77E78B" box="[151,450,1441,1465]" italics="true" pageId="22" pageNumber="23">Trionyx triunguis rudolfianus</emphasis>
<bibRefCitation id="EFFA4B0FFFB4FF85FE14FA3CFD0DE78B" author="Deraniyagala, P." box="[456,696,1441,1465]" pageId="22" pageNumber="23" pagination="1 - 42" refId="ref75253" refString="Deraniyagala, P. (1948) Some scientific results of two visits to Africa. Spolia Zeylanica, 25, 1 - 42." type="journal article" year="1948">Deraniyagala, 1948: 30</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="54D0885CFFB4FF85FD1DFA3CFD46E78B" box="[705,755,1441,1465]" pageId="22" pageNumber="23">Type</typeStatus>
locality: 
<location id="8EB46025FFB4FF85FC8FFA3CFC4CE78B" LSID="urn:lsid:plazi:treatment:03C287E8FFB4FF85FF4BFBB7FB6DE513:8EB46025FFB4FF85FC8FFA3CFC4CE78B" box="[851,1017,1441,1465]" country="Kenya" municipality="Lake Rudolf" name="Ferguson's Gulf" pageId="22" pageNumber="23">Ferguson’s Gulf</location>
, 
<collectingMunicipality id="6BB0AC84FFB4FF85FBDFFA3CFB30E78B" box="[1027,1157,1441,1465]" pageId="22" pageNumber="23">Lake Rudolf</collectingMunicipality>
, 
<collectingCountry id="F37C766EFFB4FF85FB53FA3CFB61E78B" box="[1167,1236,1441,1465]" name="Kenya" pageId="22" pageNumber="23">Kenya</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFB4FF85FF4BFA7AFF7DE487" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BD436FEFFB4FF85FF4BFA7AFF7DE487" blockId="22.[151,1437,1511,1826]" pageId="22" pageNumber="23">
<emphasis id="B91FEAECFFB4FF85FF4BFA7AFE9CE433" bold="true" box="[151,297,1511,1537]" pageId="22" pageNumber="23">Distribution</emphasis>
. It occurs both in the eastern Mediterranean and in African regions south of the Sahara Desert. In the Mediterranean it ranges along the eastern coastal areas, from 
<collectingCountry id="F37C766EFFB4FF85FC8BF996FC12E417" box="[855,935,1547,1573]" name="Turkey" pageId="22" pageNumber="23">Turkey</collectingCountry>
to 
<collectingCountry id="F37C766EFFB4FF85FC13F996FBA5E417" box="[975,1040,1547,1573]" name="Israel" pageId="22" pageNumber="23">Israel</collectingCountry>
, whereas in Africa it extends from West Africa to East Africa, southwards to the north of 
<collectingCountry id="F37C766EFFB4FF85FD28F9B2FCE1E47B" box="[756,852,1583,1609]" name="Namibia" pageId="22" pageNumber="23">Namibia</collectingCountry>
and northwards to 
<collectingCountry id="F37C766EFFB4FF85FBF7F9ADFBDBE478" box="[1067,1134,1584,1610]" name="Egypt" pageId="22" pageNumber="23">Egypt</collectingCountry>
(Nile River). In 
<collectingCountry id="F37C766EFFB4FF85FAF4F9B2FF5DE45F" name="Equatorial Guinea" pageId="22" pageNumber="23">Equatorial Guinea</collectingCountry>
it has been recorded from 
<collectingCountry id="F37C766EFFB4FF85FDD3F9CEFDCFE45F" box="[527,634,1619,1645]" name="Equatorial Guinea" pageId="22" pageNumber="23">Río Muni</collectingCountry>
at Monte Mitra in Monte Alén National Park (
<bibRefCitation id="EFFA4B0FFFB4FF85FBA7F9CEFA20E45C" author="Gonwouo, N. L. &amp; Nsang, A." box="[1147,1429,1619,1646]" pageId="22" pageNumber="23" pagination="32 - 49" refId="ref78060" refString="Gonwouo, N. L. &amp; Nsang, A. (2005) An assessment of the reptiles and amphibians of the Monte Mitra forest, Monte Alen National Park. In: Sunderland, T. C. H. (Ed.), A Biodiversity Assessment of the Monte Mitra Forest, Monte Alen National Park, Equatorial Guinea. Smithsonian Institution, Washington, D. C., pp. 32 - 49." type="book chapter" year="2005">Gonwouo &amp; Nsang 2005</bibRefCitation>
) and also it is expected to be found close to the Cameroonian border (Campo River) (
<bibRefCitation id="EFFA4B0FFFB4FF85FBAEF9EAFAFBE4A3" author="Rhodin, A. G. J. &amp; Iverson, J. B. &amp; Bour, R. &amp; Fritz, U. &amp; Georges, A. &amp; Shaffer, H. B." box="[1138,1358,1655,1681]" pageId="22" pageNumber="23" pagination="9 - 14" refId="ref87744" refString="Rhodin, A. G. J., Iverson, J. B., Bour, R., Fritz, U., Georges, A. &amp; Shaffer, H. B. (2017) Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. In: Rhodin, A. G. J., Iverson, J. B., van Dijk, P. P., Stanford, C. B., Goode, E. V., Buhlmann, K. A. &amp; Mittermeier, R. A. (Eds.), Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN / SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs, 8, 9 - 14." type="journal article" year="2017">
Rhodin 
<emphasis id="B91FEAECFFB4FF85FB0CF9E5FABEE4A3" box="[1232,1291,1655,1681]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2017
</bibRefCitation>
) (
<figureCitation id="13502A7BFFB4FF85FABBF9EAFF0EE487" captionStart="MAPS 3" captionStartId="25.[151,223,1943,1967]" captionTargetBox="[203,1383,192,1904]" captionTargetId="figure-18@25.[161,1427,181,1919]" captionTargetPageId="25" captionText="MAPS 3A–C. Distribution maps for Equatorial Guinean records of (A) Cyclanorbis elegans; (B) Trionyx triunguis; (C) Agama lebretoni." figureDoi="http://doi.org/10.5281/zenodo.7304631" httpUri="https://zenodo.org/record/7304631/files/figure.png" pageId="22" pageNumber="23">Map 3B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFB4FF85FF1BF922FB6DE513" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BD436FEFFB4FF85FF1BF922FEFEE4CC" blockId="22.[151,1437,1511,1826]" pageId="22" pageNumber="23">
<emphasis id="B91FEAECFFB4FF85FF1BF922FEFAE4EB" bold="true" box="[199,335,1727,1753]" pageId="22" pageNumber="23">Comments.</emphasis>
The current known record of this species is based on a single specimen from Monte Mitra (
<bibRefCitation id="EFFA4B0FFFB4FF85FAF0F922FEF5E4CC" author="Gonwouo, N. L. &amp; Nsang, A." pageId="22" pageNumber="23" pagination="32 - 49" refId="ref78060" refString="Gonwouo, N. L. &amp; Nsang, A. (2005) An assessment of the reptiles and amphibians of the Monte Mitra forest, Monte Alen National Park. In: Sunderland, T. C. H. (Ed.), A Biodiversity Assessment of the Monte Mitra Forest, Monte Alen National Park, Equatorial Guinea. Smithsonian Institution, Washington, D. C., pp. 32 - 49." type="book chapter" year="2005">Gonwouo &amp; Nsang 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8BD436FEFFB4FF85FF1BF89AFB6DE513" blockId="22.[151,1437,1511,1826]" box="[199,1240,1799,1826]" pageId="22" pageNumber="23">
<emphasis id="B91FEAECFFB4FF85FF1BF89AFE73E513" bold="true" box="[199,454,1799,1825]" pageId="22" pageNumber="23">Specimens examined.</emphasis>
No specimens of this species were found in the examined collections.
</paragraph>
</subSubSection>
</treatment>
</document>