137 lines
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137 lines
15 KiB
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<document id="77000E28730EBE8DE4643ADFA93BDD73" ID-CLB-Dataset="28443" ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.59.24461" ID-GBIF-Dataset="cd5bfd83-dd3d-4792-8e58-b4db2ab59601" ID-Pensoft-Pub="2305-2562-1-55" ID-ZooBank="BCA6F985BE5347C5B64D005DA0DD69BE" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2305-2562-1-55" ModsDocOrigin="African Invertebrates 59 (1)" ModsDocTitle="A new species of Ischiodon Sack (Diptera, Syrphidae) from Madagascar" checkinTime="1552155266034" checkinUser="pensoft" docAuthor="Mengual, Ximo" docDate="2018" docId="8A66212B13DF0CB8C976E6049D7C1933" docLanguage="en" docName="AfrInvert 59(1): 55-73" docOrigin="African Invertebrates 59 (1)" docSource="http://dx.doi.org/10.3897/afrinvertebr.59.24461" docTitle="Ischiodon astales Mengual, 2018, sp. n." docType="treatment" docUuid="B32ED255-02F5-4465-8CBC-7B7EA64404CC" docUuidSource="ZooBank" docVersion="7" lastPageNumber="58" masterDocId="FFE0FFA3FFC2FFA4FFB2217A7F79FFAC" masterDocTitle="A new species of Ischiodon Sack (Diptera, Syrphidae) from Madagascar" masterLastPageNumber="73" masterPageNumber="55" pageNumber="56" updateTime="1701380832606" updateUser="ExternalLinkService">
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<mods:mods id="CA81694F63A51631B53A68F6D72A1D5C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3EBD3A3382F015226624DEEA72DFDFF8">
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<mods:title id="B60A704433A1684CC4A6B8A35DD23FE1">A new species of Ischiodon Sack (Diptera, Syrphidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="6052FC411C760F31388F0DA45A835F7A" type="personal">
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<mods:roleTerm id="64CE639ECE41A2F96F6C11068E0BF316">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C02C21D43102962D286ADF1C99F986CD">Mengual, Ximo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0E8A754812DE36BF5065DEBCB767034F">text</mods:typeOfResource>
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<mods:titleInfo id="294EE4FF516C4074523BEE15A1F48F0E">
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<mods:title id="84B3CB2D8B063013A3B8A0D4DDC04763">African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part id="F8047790A16E53A68DBF9659313A7032">
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<mods:date id="317CA89BDDF3350EF0ABF880EC8E27E9">2018</mods:date>
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<mods:detail id="A8E7C34B62C3DAD09987FA98CA132CBD" type="volume">
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<mods:number id="172DF21024DFA5CB7AEB8865A30E7293">59</mods:number>
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</mods:detail>
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<mods:detail id="793943B5FAB9F8F60972C7F08F36A18E" type="issue">
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<mods:number id="80BDA0CB143C0DB4134FE3630DEA2F80">1</mods:number>
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</mods:detail>
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<mods:start id="CE031C3239E483BE246E84D40C81C9AB">55</mods:start>
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<mods:end id="E32BEF249982ACFDD6DFA3635961753A">73</mods:end>
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<mods:location id="359461663D6B87B048EDBED4CA497E7A">
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<mods:url id="110BAD2947F8CAC4F2AA14AF5D62EAD3">http://dx.doi.org/10.3897/afrinvertebr.59.24461</mods:url>
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</mods:location>
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<mods:classification id="FDA76F871917154E450B047897140C5A">journal article</mods:classification>
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<mods:identifier id="AADE03A52A80BCDE7885C3295C6CDEAF" type="DOI">http://dx.doi.org/10.3897/afrinvertebr.59.24461</mods:identifier>
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<mods:identifier id="F4FB8DADBC760A4FBC7074FD9F418D7F" type="Pensoft-Pub">2305-2562-1-55</mods:identifier>
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<mods:identifier id="05B6ED6D8FF2AACAE0E657FCA72D2E3A" type="ZooBank">BCA6F985BE5347C5B64D005DA0DD69BE</mods:identifier>
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</mods:mods>
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<treatment id="8A66212B13DF0CB8C976E6049D7C1933" ID-GBIF-Taxon="154366241" LSID="urn:lsid:zoobank.org:act:B32ED255-02F5-4465-8CBC-7B7EA64404CC" httpUri="http://treatment.plazi.org/id/8A66212B13DF0CB8C976E6049D7C1933" lastPageId="3" lastPageNumber="58" pageId="1" pageNumber="56">
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<subSubSection id="EDED05656412D74664AC76AF6648E126" pageId="1" pageNumber="56" type="nomenclature">
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<paragraph id="11E8D09B95C3FAFD8C400EFF9F824C2E" pageId="1" pageNumber="56">
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<taxonomicName id="0CF7DB8E1A010136BBEEB2B74F57D804" ID-CoL="9HNJM" LSID="http://zoobank.org/B32ED255-02F5-4465-8CBC-7B7EA64404CC" class="Insecta" family="Syrphidae" genus="Ischiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischiodon astales" order="Diptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="astales">Ischiodon astales</taxonomicName>
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<taxonomicNameLabel id="C41BC2662118AC670C878E3B113ECA02" pageId="1" pageNumber="56">sp. n.</taxonomicNameLabel>
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Figures 2, 3
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<normalizedToken id="DD183383FD67F6A66D7173E3F61BEAAF" originalValue="A–D">A-D</normalizedToken>
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, 4
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</paragraph>
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</subSubSection>
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<subSubSection id="EA730A042028033EE85C457505111DB4" pageId="2" pageNumber="57" type="differential diagnosis">
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<paragraph id="DE88E99843BDD8D26A4E620B2FFDF7E3" pageId="2" pageNumber="57">
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<pageBreakToken id="8C345DA72C2A58BF928116BAC7E11FDC" pageId="2" pageNumber="57" start="start">Differential</pageBreakToken>
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diagnosis.
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</paragraph>
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<paragraph id="CF17EE55116991CCD8B5293A81836981" pageId="2" pageNumber="57">
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Females and males of this new species do not have a calcar on the metatrochanter and they are easy to distinguish from other species since the two broad, yellow maculae on tergite 2 do not reach the lateral margins (Figs 2A, C, 3A, C). Moreover, males of
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<taxonomicName id="40F70868669F59CDB9AE0A6499B5EFEF" lsidName="I. astales" pageId="2" pageNumber="57" rank="species" species="astales">I. astales</taxonomicName>
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sp. n. have symmetrical claws on the fore leg, the same as males of
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<taxonomicName id="4156B9EA861DD59BB4D88E0728EED44B" lsidName="I. scutellaris" pageId="2" pageNumber="57" rank="species" species="scutellaris">I. scutellaris</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="051A800A1178805BE1F8E1B57E69EC4B" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57" type="description">
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<paragraph id="D2E2C3B8C802914EB777FAE4B8ED44AE" pageId="2" pageNumber="57">Description.</paragraph>
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<paragraph id="ABB678740C06E529E0C311CC85E99DBF" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">
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Male: Head (Fig. 2A, B): Face with distinct facial tubercle, entirely yellow, yellow pilose; gena yellow with two small black macula at eye margin; lunule yellow; frons yellow, yellow pilose; vertical triangle black; antenna yellow, black pilose, basoflagellomere orange, darker dorsally; arista orange-yellow, bare; eye bare; occiput
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<pageBreakToken id="007479F97132D93444D14CB2DE2ED6A0" pageId="3" pageNumber="58" start="start">black</pageBreakToken>
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, white pollinose and white pilose on ventral 1/2, yellow pollinose and yellow pilose on dorsal 1/2.
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</paragraph>
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<paragraph id="B5DE95AEA400CE83313EF28B26613F31" pageId="3" pageNumber="58">Thorax (Fig. 2A, C): Scutum shiny black, with long yellow pile; postpronotum paler, brownish, bare; notopleuron and supra-alar area yellow; postalar callus darker, brownish; scutellum yellow with medial area brownish, yellow pilose, subscutellar fringe yellow. Pleuron black, except posterior anepisternum yellow on posterior 1/2 and katepisternum with a dorsal yellow macula, yellow pilose; katepisternal pile patches separated; metaepisternum bare; metasternum bare; postmetacoxal bridge incomplete; calypter yellow, yellow pilose; plumula long, pale; halter pedicel and capitulum yellow; posterior spiracular fringes yellow. Wing: hyaline, bare on basal 1/2 and with sparse and irregularly scattered microtrichia on apical 1/2; alula mostly bare, with a few microtrichia posteriorly. Legs: fore and mid legs yellow with femora slightly more orange, coxae black and basal 1/2 of trochanters dark, entirely yellow pilose; hind leg with black coxae and basal part of trochanter dark; yellow metafemur with an almost imperceptible brown area on the apical 1/2, and metatibia slightly darker (dark yellow) on apical 1/2, black and yellow pilose.</paragraph>
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<paragraph id="F89848A7508B46A184D20B930AF9E6E9" pageId="3" pageNumber="58">Abdomen (Fig. 2A, C): Parallel-sided, with lateral margin from apical 1/2 of tergite 2 to tergite 5. Tergite 1 black, except yellow on lateral margins, yellow pilose; tergite 2 black, paler medially on posterior margin, with two large, rounded yellow maculae that do not reach lateral margins, yellow pilose, with short black pile on posterior margin; tergite 3 black on basal 1/2 becoming orange or brownish on posterior 1/2, with a medial yellow fascia that does not reach the lateral margins, with scattered, short, black pile; tergite 4 brownish-orange with a medial yellow fascia, black pilose except on the yellow fascia; tergite 5 brownish-orange, black pilose. Sternite 1 yellow; sternite 2 yellow with a medial, round, black macula; sterna 3 and 4 yellow becoming darker (brownish orange) towards genitalia; all sterna with yellow pile. Male genitalia: large, brownish-orange, as in Fig. 2E, F.</paragraph>
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<caption id="45753C16AEFBB1E5C4DE9ADAA05B9FEC" pageId="3" pageNumber="58">
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<paragraph id="631605384286C47E7A6095C7F3FE2B51" pageId="3" pageNumber="58">
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Figure 2.
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<taxonomicName id="49E6EDA698026734740FFE8C7BAE439B" class="Insecta" family="Syrphidae" genus="Ischiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischiodon astales" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="astales">Ischiodon astales</taxonomicName>
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sp. n., male holotype CASENT 2210258: A Habitus, lateral view B Frontal view C Habitus, dorsal view D Labels E Male genitalia, lateral view F Hypandrium, ventral view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<paragraph id="40CF24C9DF3BD849E7E4141100079E13" pageId="3" pageNumber="58">Female: Similar to male except for normal sexual dimorphism and as follows: frons black on dorsal 1/3, yellow on ventral 2/3 with a variable medial brownish orange vitta.</paragraph>
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<paragraph id="C14F640DB804CCEA0FC1B171A6141DAC" pageId="3" pageNumber="58">
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Variation: Due to the small number of specimens, one must expect variability in the coloration of the abdomen, legs and female frons, as in other
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<taxonomicName id="B786EEA0FA0A460DF8DFEA4DFEE2330B" class="Insecta" family="Syrphidae" genus="Ischiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischiodon" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">Ischiodon</taxonomicName>
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species.
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</paragraph>
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<paragraph id="6EBF305C8E19217FEB25C2FCD29B1574" pageId="3" pageNumber="58">Length (N=4): Body, 10.0 mm (9.0-12.0 mm); wing, 7.7 mm (7.5-8.0 mm).</paragraph>
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</subSubSection>
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<subSubSection id="1FF5EBC2C52179DA2C562AE08953F0E6" pageId="3" pageNumber="58" type="distribution">
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<paragraph id="17CCB05AC260604841F07A80C534DA5C" pageId="3" pageNumber="58">Geographical distribution.</paragraph>
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<paragraph id="05B8D9E316E87A8C870383E3BDB8868F" pageId="3" pageNumber="58">Species known from the southern regions of Atsimo-Andrefana and Anosy, part of the former Toliara Province, Madagascar (Fig. 4; see Suppl. material 1: Table S1).</paragraph>
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</subSubSection>
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<subSubSection id="1A369854B7EDD5AF170261CE217AD64E" pageId="3" pageNumber="58" type="etymology">
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<paragraph id="30B723A62E4A89319FB5FBA82468716C" pageId="3" pageNumber="58">Etymology.</paragraph>
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<paragraph id="80D8B910EE99C6E7D0FC8D55B4FF30D6" pageId="3" pageNumber="58">
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The specific epithet is derived from the Greek astales meaning unarmed, unclad (
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<bibRefCitation id="53F0A6C326DEEC9B876C1803C5A68CEF" author="Brown, RW" journalOrPublisher="Published by the author, Baltimore" pageId="6" pageNumber="61" title="Composition of scientific words a manual of methods and a lexicon of materials for the practice of logotechnics." url="https://archive.org/details/compositionofsci00brow" year="1956">Brown 1956</bibRefCitation>
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: 122). Species epithet is to be treated as an adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="7BF6EB068EC86C520EB7E0EBD713850E" pageId="3" pageNumber="58" type="type locality">
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<paragraph id="88B52BDC0783A05218F1630530CEB53C" pageId="3" pageNumber="58">Type locality.</paragraph>
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<paragraph id="F85A0C0B1820DFD047A258B474BF8F25" pageId="3" pageNumber="58">
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Madagascar: Atsimo-Andrefana Region, Mikea forest, NW of Manombo,
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<geoCoordinate id="0A0D078CC9060BFC963A0355EAE44298" direction="south" orientation="latitude" precision="9" value="-22.903667">22°54.22'S</geoCoordinate>
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,
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<geoCoordinate id="71C5801399DE6521C2C3F35D55D8C170" direction="east" orientation="longitude" precision="9" value="43.4755">43°28.53'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CB7019FCF543F806A1170F10A1BF66EC" pageId="3" pageNumber="58" type="materials_examined">
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<paragraph id="CA90BB0D412718178B87CBDE01BD4579" pageId="3" pageNumber="58">Material examined.</paragraph>
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<paragraph id="F458EA65748990AC78BA7F812E7CC761" pageId="3" pageNumber="58">
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Type material. Holotype, male, pinned with genitalia in microvial, deposited in the CAS and labelled: "MADAGASCAR, Tulear // Province, Mikea Forest, // NW of Manombo, el 30 m, // 17 - 27 July 2003" "
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<geoCoordinate id="561627F28CF6CB970F519A0CC6565292" direction="south" orientation="latitude" precision="9" value="-22.903667">22°54.22'S</geoCoordinate>
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,
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<geoCoordinate id="70044C93F5110DC36186C8D3763E6581" direction="east" orientation="longitude" precision="9" value="43.4755">43°28.53'E</geoCoordinate>
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// coll: M. Irwin, R.
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<normalizedToken id="1E4FD72191849DEED03C186601718E9F" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
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// California Acad of Sciencies // malaise trap - in deciduous // dry forest MA-02-18A-62" "CASENT // 2210258" "HOLOTYPE // Ischiodon // astales // Det. X. Mengual 2017" [red]. Paratypes: Madagascar: Atsimo-Andrefana Region, Mikea Forest, NW of Manombo, 37 m., -22.91333°, 43.48217°, 12-20.xi.2001, M. Irwin, R.
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<normalizedToken id="DB6A433718CF53B9F1A090E54E67DD78" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
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[1♀, ZFMK; CASENT 2210121]; Atsimo-Andrefana Region, Zombitse-Vohibasia National Park, near ANGAP Office, 840 m., -22.8865°, 44.69217°, 31.
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<normalizedToken id="4B94CB8F17365CA32AB7E88721338A83" originalValue="x–">x-</normalizedToken>
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16.xi.2002, R.
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<normalizedToken id="51EF9F36499A5FFC2DBBF451C955CE6F" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
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[1♀, CSCA; ZFMK-DIP-00026782]; Anosy Region, Berenty Special Reserve, 8 km NW Amboasary, 85 m., -25.00667°, 46.30333°, 26.
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<normalizedToken id="515D714208C1E2368F11CC7168957413" originalValue="x–">x-</normalizedToken>
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2.xi.2002, M. Irwin, F. Parker, R.
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<normalizedToken id="DAB8F2568AD62E0EF78CB2794C20EF94" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
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[1♀, ZFMK; CASENT 2210162]; Anosy Region, Andohahela National Park, Ihazofotsy Parcelle III, 80 m., -24.83083°, 46.53617°, 23-30.xi.2003, M. Irwin, F. Parker, R.
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<normalizedToken id="2616C55F3E94249654C72304A44AFC9B" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
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[1♀, CAS; CASENT 2210150].
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</paragraph>
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</subSubSection>
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<subSubSection id="7E572254998E18600EA3DAB31EDF447C" pageId="3" pageNumber="58" type="remarks">
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<paragraph id="2D687910F048C5CCC43BD7743A46D6D0" pageId="3" pageNumber="58">Remarks.</paragraph>
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<paragraph id="B6E8A23F947F674A6F3E1B58A55874B6" pageId="3" pageNumber="58">All specimens of this new species were collected using Malaise traps in different habitats. The male holotype was collected in a deciduous dry forest, while females CASENT 2210150 and CASENT 2210121 were collected in dry spiny forests and the female CASENT 2210162 in a gallery forest. On the other hand, the female ZFMK-DIP-00026782 was collected at a relative high altitude compared with the other specimens.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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