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<mods:title id="5601C1D18EB9F70F345AE2F7B07AC855">Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species</mods:title>
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Key to the species of 
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Guignot
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1 Elytra immaculate, evenly colored (
<figureCitation id="134B00B1FFBCFFFF7708FAC2FDEB925C" box="[567,654,1373,1395]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 3, 4</figureCitation>
)................................................................ 2
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- Elytra with maculation of yellow to reddish brown bands or spots (
<figureCitation id="134B00B1FFBCFFFF765DFAE4FCD292BE" box="[866,951,1403,1425]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 5–9</figureCitation>
)...................................... 3
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2 Robust in (L/GW = 2.00–2.11) (
<figureCitation id="134B00B1FFBCFFFF7725FA07FD339281" box="[538,598,1432,1454]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 4</figureCitation>
); distance between eyes greater than 2/3 × greatest head width (HW/EW = 1.55–1.63) (
<figureCitation id="134B00B1FFBCFFFF75D8FA2AFE4392E4" box="[231,294,1461,1483]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 4</figureCitation>
, 
<figureCitation id="134B00B1FFBCFFFF740FFA2AFE2C92E4" box="[304,329,1461,1483]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="14" pageNumber="245">11</figureCitation>
); aedeagus as in 
<figureCitation id="134B00B1FFBCFFFF74D3FA2AFD5792E4" box="[492,562,1461,1483]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 17</figureCitation>
a–e, with left lateral lobe short, approximately 2/3 × length of median lobe (
<figureCitation id="134B00B1FFBCFFFF7034FA2AFA3592E4" box="[1291,1360,1461,1483]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 17</figureCitation>
d).............................................................................................. 
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<emphasis id="B904C026FFBCFFFF701FFA4CFAF992C6" bold="true" box="[1312,1436,1491,1513]" italics="true" pageId="14" pageNumber="245">L. uniformis</emphasis>
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- Slender and oval in (L/GW= 2.07–2.18) (
<figureCitation id="134B00B1FFBCFFFF77BBFA6FFDA49129" box="[644,705,1520,1542]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 3</figureCitation>
); distance between eyes less than 2/3 × greatest head width (EW/HW = 1.63–1.73) (
<figureCitation id="134B00B1FFBCFFFF746AF992FEF6910C" box="[341,403,1549,1571]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 3</figureCitation>
, 
<figureCitation id="134B00B1FFBCFFFF74A1F992FEDC910C" box="[414,441,1549,1571]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="14" pageNumber="245">10</figureCitation>
); aedeagus as in 
<figureCitation id="134B00B1FFBCFFFF7763F992FDC7910C" box="[604,674,1549,1571]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 16</figureCitation>
a–e, with left lateral lobe long, slender, nearly as long as median lobe, and with setal fringe not extending to midlength (
<figureCitation id="134B00B1FFBCFFFF775DF9B4FDC2916E" box="[610,679,1579,1601]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 16</figureCitation>
d)................................................... 
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<emphasis id="B904C026FFBCFFFF7017F9B4FAF9916E" bold="true" box="[1320,1436,1579,1601]" italics="true" pageId="14" pageNumber="245">L. angustus</emphasis>
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3 Elytra with maculation consisting of yellow to reddish-brown bands extending from lateral margins nearly to elytral suture, not, or very weakly, obliquely projected, rarely reduced to spots (
<figureCitation id="134B00B1FFBCFFFF7666F9FAFCD49154" box="[857,945,1637,1659]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 5, 6</figureCitation>
); robust, L/GW = 1.93–2.07, PNWpost/PNWant = 1.64–1.68........................................................................................... 4
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- Elytral maculation usually reduced to spots, most often in basal half; spots or bands obliquely oriented, more so in distal half (
<figureCitation id="134B00B1FFBCFFFF75D8F922FE5A91FC" box="[231,319,1725,1747]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 7–9</figureCitation>
); slender (
<figureCitation id="134B00B1FFBCFFFF749CF922FE9F91FC" box="[419,506,1725,1747]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Figs 7–9</figureCitation>
), L/GW = 2.00–2.29, PNWpost/PNWant = 1.53–1.62................................. 5
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4 Aedeagus as in 
<figureCitation id="134B00B1FFBCFFFF7448F944FED891DE" box="[375,445,1755,1777]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 18</figureCitation>
a–e, with median lobe distally expanded and bluntly rounded at apex (
<figureCitation id="134B00B1FFBCFFFF7163F944FBC091DE" box="[1116,1189,1755,1777]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 18</figureCitation>
a–c); left lateral lobe with setal fringe restricted to distal half (
<figureCitation id="134B00B1FFBCFFFF7703F967FDE79021" box="[572,642,1784,1806]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 18</figureCitation>
d); right lateral lobe triangular, with distal angle sharply rounded and not strongly projected (
<figureCitation id="134B00B1FFBCFFFF7478F88AFEE89004" box="[327,397,1813,1835]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 18</figureCitation>
e)............................................................................ 
<taxonomicName id="4C7067B7FFBCFFFF7003F889FAFE9003" box="[1340,1435,1814,1836]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B904C026FFBCFFFF7003F889FAFE9003" bold="true" box="[1340,1435,1814,1836]" italics="true" pageId="14" pageNumber="245">L. bicolor</emphasis>
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<paragraph id="8BCF1C34FFBCFFFF75A8F8ABFAFE90AB" blockId="14.[151,1436,1373,2011]" pageId="14" pageNumber="245">
<keyLead id="3081BF01FFBCFFFF75A8F8ABFAFE90AB" pageId="14" pageNumber="245">
- Aedeagus as in 
<figureCitation id="134B00B1FFBCFFFF7445F8ACFEA79066" box="[378,450,1843,1865]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 19</figureCitation>
a–e, with median lobe weakly expanded distally before attenuating to rounded apex (
<figureCitation id="134B00B1FFBCFFFF71C4F8ACFA209066" box="[1275,1349,1843,1865]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 19</figureCitation>
a–c); left lateral lobe with setal fringe extending along inner dorsal margin to midlength (
<figureCitation id="134B00B1FFBCFFFF76E8F8CFFB789049" box="[983,1053,1872,1894]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 19</figureCitation>
d); right lateral lobe subtriangular, moderately emarginated along length of dorsal margin, distal angle projecting with apex rounded (
<figureCitation id="134B00B1FFBCFFFF715AF8F2FBC990AC" box="[1125,1196,1901,1923]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 19</figureCitation>
e)........ 
<taxonomicName id="4C7067B7FFBCFFFF7023F8F1FAFE90AB" box="[1308,1435,1902,1924]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="armulatus">
<emphasis id="B904C026FFBCFFFF7023F8F1FAFE90AB" bold="true" box="[1308,1435,1902,1924]" italics="true" pageId="14" pageNumber="245">L. armulatus</emphasis>
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5 Eyes large (HW/EW = 1.60–1.67) (
<figureCitation id="134B00B1FFBCFFFF770AF814FD15908E" box="[565,624,1931,1953]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 7</figureCitation>
); elytral maculation with posterior bands or spots extending anterolaterally nearly to basal half of elytra (
<figureCitation id="134B00B1FFBCFFFF749AF837FE849091" box="[421,481,1960,1982]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 7</figureCitation>
); aedeagus as in 
<figureCitation id="134B00B1FFBCFFFF77B6F837FDB79091" box="[649,722,1960,1982]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 20</figureCitation>
a–e, with median lobe slender and not sinuate in dorsal aspect (
<figureCitation id="134B00B1FFBCFFFF7004F837FAE69091" box="[1339,1411,1960,1982]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 20</figureCitation>
b), weakly expanded before gradually attenuating to elongate apex (
<figureCitation id="134B00B1FFBCFFFF766BF85AFCC590F4" box="[852,928,1989,2011]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 20</figureCitation>
a, c); left lateral lobe slender (
<figureCitation id="134B00B1FFBCFFFF71FCF85AFA6E90F4" box="[1219,1291,1989,2011]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 20</figureCitation>
d); right lateral
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<paragraph id="8BCF1C34FFBDFFFE75DEFF08FAFE97C8" blockId="15.[151,1436,151,378]" pageId="15" pageNumber="246">
lobe triangular and distal angle not projecting (
<figureCitation id="134B00B1FFBDFFFE779DFF08FD8D9782" box="[674,744,151,173]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Fig. 20</figureCitation>
e)................................................ 
<taxonomicName id="4C7067B7FFBDFFFE707CFF08FAFE9782" box="[1347,1435,151,173]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="246" phylum="Arthropoda" rank="species" species="clayae">
<emphasis id="B904C026FFBDFFFE707CFF08FAFE9782" bold="true" box="[1347,1435,151,173]" italics="true" pageId="15" pageNumber="246">L. clayae</emphasis>
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- Eyes small (HW/EW = 1.47–1.55) (
<figureCitation id="134B00B1FFBDFFFE7703FF2BFDF197E5" box="[572,660,180,202]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="15" pageNumber="246">Figs 8, 9</figureCitation>
); posterior bands or spots extending anterodistally beyond middle of elytra onto basal half at lateral extreme............................................................................. 6
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6 Aedeagus as in 
<figureCitation id="134B00B1FFBDFFFE7443FF70FEA0962A" box="[380,453,239,261]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Figs 21</figureCitation>
a–e, with median lobe distally shortened and attenuate in lateral aspect (
<figureCitation id="134B00B1FFBDFFFE71BAFF70FBB5962A" box="[1157,1232,239,261]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Figs 21</figureCitation>
a, c); left lateral lobe broad (
<figureCitation id="134B00B1FFBDFFFE741AFE93FE0F960D" box="[293,362,268,290]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Fig. 21</figureCitation>
d); right lateral lobe subtriangular with distal angle strongly projecting (
<figureCitation id="134B00B1FFBDFFFE7125FE93FB04960D" box="[1050,1121,268,290]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Fig. 21</figureCitation>
e)................ 
<taxonomicName id="4C7067B7FFBDFFFE7006FE93FAFE960D" box="[1337,1435,268,290]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="246" phylum="Arthropoda" rank="species" species="nanops">
<emphasis id="B904C026FFBDFFFE7006FE93FAFE960D" bold="true" box="[1337,1435,268,290]" italics="true" pageId="15" pageNumber="246">L. nanops</emphasis>
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<paragraph id="8BCF1C34FFBDFFFE75A8FEB5FAFE9655" blockId="15.[151,1436,151,378]" pageId="15" pageNumber="246">
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- Aedeagus as in 
<figureCitation id="134B00B1FFBDFFFE7446FEB6FEA49610" box="[377,449,297,319]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Figs 22</figureCitation>
a–e, with median lobe distally lengthened and subparallel in lateral aspect 
<figureCitation id="134B00B1FFBDFFFE71B3FEB6FBB09610" box="[1164,1237,297,319]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Figs 22</figureCitation>
a, c); left lateral lobe less broad and distally attenuate (
<figureCitation id="134B00B1FFBDFFFE771FFED8FD0D9672" box="[544,616,327,349]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Fig. 22</figureCitation>
d); right lateral lobe triangular with distal angle acute, not projecting (
<figureCitation id="134B00B1FFBDFFFE703CFED8FA2E9672" box="[1283,1355,327,349]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="15" pageNumber="246">Fig. 22</figureCitation>
e).............................................................................................. 
<taxonomicName id="4C7067B7FFBDFFFE7037FEFBFAFE9655" box="[1288,1435,356,378]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="246" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="B904C026FFBDFFFE7037FEFBFAFE9655" bold="true" box="[1288,1435,356,378]" italics="true" pageId="15" pageNumber="246">L. octoguttatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>