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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3690.1.1" ID-GBIF-Dataset="c8670273-a22c-4b4a-972d-38a43cd66f86" ID-ISSN="1175-5326" ID-Zenodo-Dep="6329811" ID-ZooBank="0874D336-BA8C-4266-AA50-633167C816F3" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1629874133667" checkinUser="felipe" docAuthor="PLANT, ADRIAN R." docDate="2013" docId="03D787D0FFF3FFE7FF05FF22FBEBFE0F" docLanguage="en" docName="zt03690p098.pdf" docOrigin="Zootaxa 3690 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3690.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.10:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="10" docTitle="Hybos serratus Yang &amp; Yang, 1992: 1089" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="FFEEFFA8FFCBFFDDFF92FFB5FF99FF88" masterDocTitle="The genus Hybos Meigen (Diptera: Empidoidea: Hybotidae) in Thailand" masterLastPageNumber="98" masterPageNumber="1" pageNumber="57" updateTime="1679570708792" updateUser="felipe">
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<mods:titleInfo>
<mods:title>The genus Hybos Meigen (Diptera: Empidoidea: Hybotidae) in Thailand</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>PLANT, ADRIAN R.</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2013</mods:date>
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<mods:number>2013-07-17</mods:number>
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<mods:detail type="volume">
<mods:number>3690</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:end>98</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.3690.1.1</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3690.1.1</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6339120" ID-GBIF-Taxon="193889223" ID-Zenodo-Dep="6339120" LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F" httpUri="http://treatment.plazi.org/id/03D787D0FFF3FFE7FF05FF22FBEBFE0F" lastPageId="58" lastPageNumber="58" pageId="56" pageNumber="57">
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<taxonomicName ID-CoL="3MYNW" authority="Yang &amp; Yang" authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[151,508,151,177]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="serratus">
<emphasis bold="true" box="[151,508,151,177]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[151,333,151,177]" italics="true" pageId="56" pageNumber="57">Hybos serratus</emphasis>
Yang &amp; Yang
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<paragraph blockId="56.[151,508,151,210]" box="[151,463,186,210]" pageId="56" pageNumber="57">
(
<figureCitation box="[159,231,186,210]" captionStart="FIGURES 1–4" captionStartId="4.[151,263,1949,1972]" captionTargetBox="[159,1427,658,1918]" captionTargetId="figure-314@4.[151,1436,652,1928]" captionTargetPageId="4" captionText="FIGURES 1–4. Morphology of Hybos spp. 1. H. bispinatus sp. nov. ♀ habitus. 2. H. khamfui sp. nov. ♂ habitus. 3. H. serratus Yang&amp; Yang♂ right posterior femur, ventral view. 4. H. lannaensis sp. nov. ♀ postpedicel and stylus. aps = anterior proclinate setae, pvf = posteroventral fringe, vs = ventral spines." figureDoi="http://doi.org/10.5281/zenodo.6329813" httpUri="https://zenodo.org/record/6329813/files/figure.png" pageId="56" pageNumber="57">Figs 3</figureCitation>
,
<figureCitation box="[243,344,186,210]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="56" pageNumber="57">141–150</figureCitation>
,
<figureCitation box="[355,399,186,210]" captionStart="FIGURES 269–277" captionStartId="89.[151,264,1945,1968]" captionTargetBox="[244,1325,201,1898]" captionTargetId="figure-80@89.[219,1367,165,1924]" captionTargetPageId="89" captionText="FIGURES 269–277. Distribution of Hybos spp. in Thailand plotted at a resolution of 1° of latitude and longitude. 269. H. serratus Yang &amp; Yang. 270. H. shamshevi sp. nov. 271. H. sinclairi sp. nov. 272. H. songbai sp. nov. 273. H. steatopygus sp. nov. 274. H. stigmaticus sp. nov. 275. H. subapicalis sp. nov. 276. H. tetricus sp. nov. 277. H. thaosaeo sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6329911" httpUri="https://zenodo.org/record/6329911/files/figure.png" pageId="56" pageNumber="57">269</figureCitation>
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<figureCitation box="[411,455,186,210]" captionStart="FIGURES 291–298" captionStartId="93.[151,263,1912,1935]" captionTargetBox="[229,1367,185,1880]" captionTargetId="figure-80@93.[189,1403,163,1912]" captionTargetPageId="93" captionText="FIGURES 291–298. Phenology of Hybos spp. monthly abundance (as % of total number of individuals captured) is plotted against month of capture. 291. H. inthanonensis sp. nov. 292. H. kaluang sp. nov. 293. H. khamfui sp. nov. 294. H. particularis Yang, Yang &amp; Hu. 295. H. pisadaanus sp. nov. 296. H. saenmueangmai sp. nov. 297. H. serratus Yang &amp; Yang. 298. H. songbai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6329917" httpUri="https://zenodo.org/record/6329917/files/figure.png" pageId="56" pageNumber="57">297</figureCitation>
)
</paragraph>
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<paragraph blockId="56.[151,582,258,280]" box="[151,582,258,280]" pageId="56" pageNumber="57">
<treatmentCitationGroup box="[151,582,258,280]" pageId="56" pageNumber="57">
<treatmentCitation author="Yang, D. &amp; Yang, C. K." box="[151,579,258,280]" page="1089" pageId="56" pageNumber="57" year="1992">
<taxonomicName ID-CoL="3MYNW" authority="Yang &amp; Yang, 1992: 1089" authorityName="Yang &amp; Yang, 1992:" authorityPageNumber="1089" authorityYear="1089" box="[151,579,258,280]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[151,305,258,280]" italics="true" pageId="56" pageNumber="57">Hybos serratus</emphasis>
<bibRefCitation author="Yang, D. &amp; Yang, C. K." box="[311,579,258,280]" pageId="56" pageNumber="57" pagination="1089 - 1097" refId="ref82013" refString="Yang, D. &amp; Yang, C. K. (1992) Diptera: Empididae. In: Chen, S. (Ed.), Insects of the Hengduan Mountains Region 2. Science Press, Beijing, pp. 1089 - 1097." type="book chapter" year="1992">Yang &amp; Yang, 1992: 1089</bibRefCitation>
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.
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</paragraph>
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<subSubSection pageId="56" pageNumber="57" type="materials_examined">
<paragraph blockId="56.[151,1436,325,2030]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[151,390,325,350]" pageId="56" pageNumber="57">Material examined.</emphasis>
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<specimenCount box="[404,467,325,349]" pageId="56" pageNumber="57" type="male">231♂</specimenCount>
,
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,
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<collectingRegion box="[736,991,325,349]" country="Thailand" name="Chiang Mai" pageId="56" pageNumber="57">Chiang Mai Province</collectingRegion>
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<collectingMunicipality box="[691,795,361,385]" pageId="56" pageNumber="57">Kiewlom</collectingMunicipality>
2/
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<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FF05FE38FE98FE2D" box="[151,257,397,421]" country="Thailand" latitude="20.05915" longLatPrecision="1" longitude="99.14253" name="Kiewlom" pageId="56" pageNumber="57">Kiewlom</location>
1/montane forest,
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<collectingMunicipality box="[818,1133,433,458]" pageId="56" pageNumber="57">Thung Buatong View Point</collectingMunicipality>
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:
<collectingRegion box="[588,856,469,493]" country="Thailand" name="Chiang Mai" pageId="56" pageNumber="57">Chiang Mai Province</collectingRegion>
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<collectorName box="[880,1067,469,493]" pageId="56" pageNumber="57">Doi Chiangdao</collectorName>
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HRD Pakea Stn (evergreen forest and coffee plantation),
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nr. rd. from
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,
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:
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;
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<geoCoordinate box="[907,1005,577,601]" degrees="18.82" direction="north" orientation="latitude" pageId="56" pageNumber="57" precision="555" value="18.82">18.82°N</geoCoordinate>
,
<geoCoordinate box="[1020,1114,577,601]" degrees="98.89" direction="east" orientation="longitude" pageId="56" pageNumber="57" precision="555" value="98.89">98.89°E</geoCoordinate>
,
<quantity box="[1129,1221,577,601]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="56" pageNumber="57" unit="m" value="1550.0">1550 m</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642424" pageId="56" pageNumber="57">
<geoCoordinate box="[1236,1334,577,601]" degrees="18.80" direction="north" orientation="latitude" pageId="56" pageNumber="57" precision="555" value="18.8">18.80°N</geoCoordinate>
,
<geoCoordinate box="[1349,1431,577,601]" degrees="98.90" direction="east" orientation="longitude" pageId="56" pageNumber="57" precision="555" value="98.9">98.90E</geoCoordinate>
,
<quantity box="[151,240,613,637]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="56" pageNumber="57" unit="m" value="1250.0">1250 m</quantity>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642417" pageId="56" pageNumber="57">
<collectingRegion box="[252,495,613,637]" country="Thailand" name="Chiang Mai" pageId="56" pageNumber="57">Chiang Mai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FE69FDD0FCAAFDF5" box="[507,819,613,637]" country="Thailand" latitude="19.5" longLatPrecision="5580" longitude="97.95" name="Doi Inthanon National Park" pageId="56" pageNumber="57" stateProvince="Chiang Mai">Doi Inthanon National Park</location>
(data lost):
<collectingRegion box="[956,1236,613,638]" country="Thailand" name="Mae Hong Son" pageId="56" pageNumber="57">Mae Hong Son Province</collectingRegion>
,
<locationDeviation location="Mae Hong Son" pageId="56" pageNumber="57">
<quantity box="[1248,1320,613,637]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="56" pageNumber="57" unit="km" value="22.0">22 km</quantity>
N of
<collectingRegion country="Thailand" name="Mae Hong Son" pageId="56" pageNumber="57">Mae Hong Son</collectingRegion>
</locationDeviation>
,
<geoCoordinate box="[295,377,649,673]" degrees="19.5" direction="north" orientation="latitude" pageId="56" pageNumber="57" precision="5555" value="19.5">19.5°N</geoCoordinate>
,
<geoCoordinate box="[398,492,649,673]" degrees="97.95" direction="east" orientation="longitude" pageId="56" pageNumber="57" precision="555" value="97.95">97.95°E</geoCoordinate>
,
<quantity box="[512,595,649,673]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="56" pageNumber="57" unit="m" value="930.0">
<elevation box="[512,595,649,673]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="56" pageNumber="57" unit="m" value="930.0">930 m</elevation>
</quantity>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642473" pageId="56" pageNumber="57">
<collectingRegion box="[614,779,649,673]" country="Thailand" name="Nan" pageId="56" pageNumber="57">Nan Province</collectingRegion>
,
<collectingMunicipality box="[799,1145,649,673]" pageId="56" pageNumber="57">Doi Phu Kha National Park</collectingMunicipality>
;
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FB1EFD3CFB4AFD29" box="[1164,1235,649,673]" country="Thailand" latitude="19.205816" longLatPrecision="1" longitude="101.07695" municipality="Doi Phu Kha National Park" name="Office" pageId="56" pageNumber="57" stateProvince="Nan">Office</location>
6,
<geoCoordinate box="[1287,1431,649,673]" degrees="19" direction="north" minutes="12.349" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="19.205816">19°12.349'N</geoCoordinate>
,
<geoCoordinate box="[151,292,685,709]" degrees="101" direction="east" minutes="4.617" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.07695">101°4.617'E</geoCoordinate>
,
<quantity box="[303,390,685,709]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="56" pageNumber="57" unit="m" value="1360.0">
<elevation box="[303,390,685,709]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="56" pageNumber="57" unit="m" value="1360.0">1360 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642423" box="[402,914,685,709]" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FE00FD18FE40FD4D" box="[402,473,685,709]" country="Thailand" latitude="19.207634" longLatPrecision="1" longitude="101.0811" name="Office" pageId="56" pageNumber="57">Office</location>
11,
<geoCoordinate box="[520,664,685,709]" degrees="19" direction="north" minutes="12.458" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="19.207634">19°12.458'N</geoCoordinate>
,
<geoCoordinate box="[675,815,685,709]" degrees="101" direction="east" minutes="4.866" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.0811">101°4.866'E</geoCoordinate>
,
<quantity box="[827,914,685,709]" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="56" pageNumber="57" unit="m" value="1359.0">
<elevation box="[827,914,685,709]" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="56" pageNumber="57" unit="m" value="1359.0">1359 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642480" box="[925,1436,685,709]" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FC0FFD18FC7DFD4D" box="[925,996,685,709]" country="Thailand" latitude="19.2023" longLatPrecision="1" longitude="101.078514" name="Office" pageId="56" pageNumber="57">Office</location>
12,
<geoCoordinate box="[1045,1186,685,709]" degrees="19" direction="north" minutes="12.138" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="19.2023">19°12.138'N</geoCoordinate>
<geoCoordinate box="[1193,1333,685,709]" degrees="101" direction="east" minutes="4.711" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.078514">101°4.711'E</geoCoordinate>
,
<quantity box="[1344,1432,685,709]" metricMagnitude="3" metricUnit="m" metricValue="1.331" pageId="56" pageNumber="57" unit="m" value="1331.0">
<elevation box="[1344,1432,685,709]" metricMagnitude="3" metricUnit="m" metricValue="1.331" pageId="56" pageNumber="57" unit="m" value="1331.0">1331 m</elevation>
</quantity>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642474" box="[151,1315,721,746]" pageId="56" pageNumber="57">
<collectingRegion box="[151,401,721,745]" country="Thailand" name="Phitsanulok" pageId="56" pageNumber="57">Phitsanulok Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FE37FD64FCFAFD61" box="[421,867,721,746]" country="Thailand" latitude="16.844017" longLatPrecision="1" longitude="100.88157" name="Thung Salaeng Luang National Park" pageId="56" pageNumber="57" stateProvince="Phitsanulok">Thung Salaeng Luang National Park</location>
,
<geoCoordinate box="[887,1032,721,745]" degrees="16" direction="north" minutes="50.641" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="16.844017">16°50.641'N</geoCoordinate>
,
<geoCoordinate box="[1052,1207,721,745]" degrees="100" direction="east" minutes="52.894" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="100.88157">100°52.894'E</geoCoordinate>
,
<quantity box="[1227,1311,721,745]" metricMagnitude="2" metricUnit="m" metricValue="5.57" pageId="56" pageNumber="57" unit="m" value="557.0">
<elevation box="[1227,1311,721,745]" metricMagnitude="2" metricUnit="m" metricValue="5.57" pageId="56" pageNumber="57" unit="m" value="557.0">557 m</elevation>
</quantity>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642477" pageId="56" pageNumber="57">
<collectingRegion country="Thailand" name="Lampang" pageId="56" pageNumber="57">Lampang Province</collectingRegion>
,
<collectingMunicipality box="[272,567,757,781]" pageId="56" pageNumber="57">Chae Son National Park</collectingMunicipality>
;
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FDDBFD40FD20FC85" box="[585,697,757,781]" country="Thailand" latitude="18.863583" longLatPrecision="1" longitude="99.3687" municipality="Chae Son National Park" name="Doi Laan" pageId="56" pageNumber="57" stateProvince="Lampang">Doi Laan</location>
unit-2,
<geoCoordinate box="[799,943,757,781]" degrees="18" direction="north" minutes="51.815" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="18.863583">18°51.815'N</geoCoordinate>
,
<geoCoordinate box="[962,1103,757,781]" degrees="99" direction="east" minutes="22.122" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="99.3687">99°22.122'E</geoCoordinate>
,
<quantity box="[1122,1217,757,781]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="56" pageNumber="57" unit="m" value="1413.0">
<elevation box="[1122,1217,757,781]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="56" pageNumber="57" unit="m" value="1413.0">1413 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642496" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FB46FD40FADDFC85" box="[1236,1348,757,781]" country="Thailand" latitude="18.8471" longLatPrecision="9" longitude="99.36183" name="Doi Laan" pageId="56" pageNumber="57">Doi Laan</location>
unit-1,
<geoCoordinate box="[151,295,793,817]" degrees="18" direction="north" minutes="50.826" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="18.8471">18°50.826'N</geoCoordinate>
,
<geoCoordinate box="[313,441,793,817]" degrees="99" direction="east" minutes="21.71" orientation="longitude" pageId="56" pageNumber="57" precision="9" value="99.36183">99°21.71'E</geoCoordinate>
,
<quantity box="[459,554,793,817]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="56" pageNumber="57" unit="m" value="1413.0">
<elevation box="[459,554,793,817]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="56" pageNumber="57" unit="m" value="1413.0">1413 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642469" box="[572,1234,793,817]" pageId="56" pageNumber="57">
campground/lavatory,
<geoCoordinate box="[831,975,793,817]" degrees="18" direction="north" minutes="49.894" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="18.831566">18°49.894'N</geoCoordinate>
,
<geoCoordinate box="[993,1134,793,817]" degrees="99" direction="east" minutes="28.354" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="99.472565">99°28.354'E</geoCoordinate>
,
<quantity box="[1153,1234,793,817]" metricMagnitude="2" metricUnit="m" metricValue="4.67" pageId="56" pageNumber="57" unit="m" value="467.0">467 m</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642363" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FB76FCAFFAC3FCB9" box="[1252,1370,794,817]" country="Thailand" latitude="18.8274" longLatPrecision="1" longitude="99.41185" name="Mae Paan" pageId="56" pageNumber="57">Mae Paan</location>
unit;
<geoCoordinate box="[151,295,829,853]" degrees="18" direction="north" minutes="49.644" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="18.8274">18°49.644'N</geoCoordinate>
,
<geoCoordinate box="[307,447,829,853]" degrees="99" direction="east" minutes="24.711" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="99.41185">99°24.711'E</geoCoordinate>
,
<quantity box="[459,534,829,853]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="56" pageNumber="57" unit="m" value="815.0">
<elevation box="[459,534,829,853]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="56" pageNumber="57" unit="m" value="815.0">815 m</elevation>
</quantity>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642493" pageId="56" pageNumber="57">
<collectingRegion box="[545,848,829,853]" country="Thailand" name="Kamphaeng Phet" pageId="56" pageNumber="57">Kamphaeng Phet Province</collectingRegion>
,
<collectingMunicipality box="[860,1149,829,854]" pageId="56" pageNumber="57">Mae Wong National Park</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FB1BFC88FA93FCDD" box="[1161,1290,829,853]" country="Thailand" latitude="16.086866" longLatPrecision="1" longitude="99.1096" municipality="Mae Wong National Park" name="Chong Yen" pageId="56" pageNumber="57" stateProvince="Kamphaeng Phet">Chong Yen</location>
,
<geoCoordinate box="[1301,1432,829,853]" degrees="16" direction="north" minutes="5.212" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="16.086866">16°5.212'N</geoCoordinate>
,
<geoCoordinate box="[151,277,865,889]" degrees="99" direction="east" minutes="6.576" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="99.1096">99°6.576'E</geoCoordinate>
,
<quantity box="[289,378,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.3059999999999998" pageId="56" pageNumber="57" unit="m" value="1306.0">
<elevation box="[289,378,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.3059999999999998" pageId="56" pageNumber="57" unit="m" value="1306.0">1306 m</elevation>
</quantity>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642377" box="[389,1382,865,889]" pageId="56" pageNumber="57">
<collectingRegion box="[389,550,865,889]" country="Thailand" name="Loei" pageId="56" pageNumber="57">Loei Province</collectingRegion>
,
<collectingMunicipality box="[562,836,865,889]" pageId="56" pageNumber="57">Phu Ruea National Park</collectingMunicipality>
;
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FCDDFCD4FC26FCF1" box="[847,959,865,889]" country="Thailand" latitude="17.51225" longLatPrecision="1" longitude="101.34335" municipality="Phu Ruea National Park" name="Sa Sawan" pageId="56" pageNumber="57" stateProvince="Loei">Sa Sawan</location>
,
<geoCoordinate box="[971,1116,865,889]" degrees="17" direction="north" minutes="30.735" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="17.51225">17°30.735'N</geoCoordinate>
,
<geoCoordinate box="[1127,1283,865,889]" degrees="101" direction="east" minutes="20.601" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.34335">101°20.601'E</geoCoordinate>
,
<quantity box="[1294,1382,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.3519999999999999" pageId="56" pageNumber="57" unit="m" value="1352.0">
<elevation box="[1294,1382,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.3519999999999999" pageId="56" pageNumber="57" unit="m" value="1352.0">1352 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642368" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FAE0FCD4FF68FC16" country="Thailand" latitude="17.508368" longLatPrecision="1" longitude="101.3478" name="Pah Lo Noy" pageId="56" pageNumber="57">Pah Lo Noy</location>
,
<geoCoordinate box="[256,400,901,925]" degrees="17" direction="north" minutes="30.502" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="17.508368">17°30.502'N</geoCoordinate>
,
<geoCoordinate box="[415,570,901,925]" degrees="101" direction="east" minutes="20.868" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.3478">101°20.868'E</geoCoordinate>
,
<quantity box="[585,666,901,925]" metricMagnitude="3" metricUnit="m" metricValue="1.343" pageId="56" pageNumber="57" unit="m" value="1343.0">
<elevation box="[585,666,901,925]" metricMagnitude="3" metricUnit="m" metricValue="1.343" pageId="56" pageNumber="57" unit="m" value="1343.0">1343m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642366" box="[681,1223,901,925]" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FD3BFC33FD6CFC15" box="[681,757,902,925]" country="Thailand" latitude="17.512333" longLatPrecision="13" longitude="101.34417" name="Nature" pageId="56" pageNumber="57">Nature</location>
trail,
<geoCoordinate box="[830,960,901,925]" degrees="17" direction="north" minutes="30.74" orientation="latitude" pageId="56" pageNumber="57" precision="9" value="17.512333">17°30.74'N</geoCoordinate>
,
<geoCoordinate box="[975,1116,901,925]" degrees="101" direction="east" minutes="20.65" orientation="longitude" pageId="56" pageNumber="57" precision="9" value="101.34417">101°20.65'E</geoCoordinate>
,
<quantity box="[1131,1223,901,925]" metricMagnitude="3" metricUnit="m" metricValue="1.353" pageId="56" pageNumber="57" unit="m" value="1353.0">
<elevation box="[1131,1223,901,925]" metricMagnitude="3" metricUnit="m" metricValue="1.353" pageId="56" pageNumber="57" unit="m" value="1353.0">1353 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642373" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FB44FC33FACAFC16" box="[1238,1363,902,926]" country="Thailand" latitude="17.502132" longLatPrecision="1" longitude="101.33898" name="Huay Taey" pageId="56" pageNumber="57">Huay Taey</location>
ditch,
<geoCoordinate box="[151,293,937,961]" degrees="17" direction="north" minutes="30.128" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="17.502132">17°30.128'N</geoCoordinate>
<geoCoordinate box="[300,454,937,961]" degrees="101" direction="east" minutes="20.339" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.33898">101°20.339'E</geoCoordinate>
,
<quantity box="[466,554,937,961]" metricMagnitude="3" metricUnit="m" metricValue="1.233" pageId="56" pageNumber="57" unit="m" value="1233.0">
<elevation box="[466,554,937,961]" metricMagnitude="3" metricUnit="m" metricValue="1.233" pageId="56" pageNumber="57" unit="m" value="1233.0">1233 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642462" pageId="56" pageNumber="57">
<location LSID="urn:lsid:plazi:treatment:03D787D0FFF3FFE7FF05FF22FBEBFE0F:8EA1601DFFF3FFE5FDA4FC1FFC85FC49" box="[566,796,937,961]" country="Thailand" latitude="17.5007" longLatPrecision="1" longitude="101.34123" name="Pan Hin Khan Maak" pageId="56" pageNumber="57">Pan Hin Khan Maak</location>
ditch,
<geoCoordinate box="[874,1014,937,961]" degrees="17" direction="north" minutes="30.042" orientation="latitude" pageId="56" pageNumber="57" precision="1" value="17.5007">17°30.042'N</geoCoordinate>
<geoCoordinate box="[1022,1177,937,961]" degrees="101" direction="east" minutes="20.474" orientation="longitude" pageId="56" pageNumber="57" precision="1" value="101.34123">101°20.474'E</geoCoordinate>
,
<quantity box="[1189,1278,937,961]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="56" pageNumber="57" unit="m" value="1219.0">
<elevation box="[1189,1278,937,961]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="56" pageNumber="57" unit="m" value="1219.0">1219 m</elevation>
</quantity>
: various dates 2006–2011 (
<collectionCode box="[290,366,973,997]" pageId="56" pageNumber="57">QSBG</collectionCode>
and
<collectionCode box="[421,518,973,997]" pageId="56" pageNumber="57">NMGW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="56" pageNumber="57" type="diagnosis">
<paragraph blockId="56.[151,1436,325,2030]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[199,321,1009,1034]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
A yellow legged species with tip (and occasionally dorsum) of hind femur blackish. The front tibia is conspicuously darker than the mid tibia and the hind tibia yellowish or suffused blackish at least near base and tip. The antennal stylus is distinctly subplumose with a bare tip. Both left and right surstyli are narrowly elongate but the right surstylus is minutely serrate apically. The hypandrium has two broad apical lobes. Tergite 8 of females encircles the abdomen and sternite 8 has dark posteroapical lateral extensions, appearing broadly T-shaped in posterior view.
</paragraph>
</subSubSection>
<subSubSection lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="description">
<paragraph blockId="56.[151,1436,325,2030]" lastBlockId="57.[151,1436,151,503]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<emphasis bold="true" box="[199,344,1225,1250]" pageId="56" pageNumber="57">Description.</emphasis>
<emphasis box="[352,409,1225,1249]" italics="true" pageId="56" pageNumber="57">Male</emphasis>
: length
<quantity box="[504,643,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="4.2" metricValueMin="3.1" pageId="56" pageNumber="57" unit="mm" value="3.65" valueMax="4.2" valueMin="3.1">3.1–4.2 mm</quantity>
.
<emphasis box="[655,716,1225,1249]" italics="true" pageId="56" pageNumber="57">Head</emphasis>
subshining black; face dark, blackish yellow below. Postocular setae black, a few paler hairs on lower occiput. Antenna black, postpedicel ovate in lateral view, ~2.5X long as wide; stylus ~5X long as postpedicel, black and distinctly subplumose on basal 0.8–0.85, bare and somewhat whitish in some views distally. Mouthparts blackish, palpus very narrow, with fine apical seta.
<emphasis box="[1212,1293,1333,1357]" italics="true" pageId="56" pageNumber="57">Thorax</emphasis>
with ground colour black, postalar callus, scutellum basally and pleura somewhat paler; scutum (especially at sides and about prescutellar area) and pleura dusted; acr biserial, dc uniserial, small and fine; posterior dc and acr, upper npl (2X long as lower npl), pa and 2 sct distinct, other setae small, hair-like.
<emphasis box="[931,984,1442,1465]" italics="true" pageId="56" pageNumber="57">Legs</emphasis>
mostly yellow with F
<subScript attach="left" box="[1235,1243,1456,1470]" fontSize="6" pageId="56" pageNumber="57">3</subScript>
on apical 0.1, F
<subScript attach="right" box="[1428,1436,1456,1470]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
and F
<subScript attach="left" box="[214,222,1495,1509]" fontSize="6" pageId="56" pageNumber="57">2</subScript>
very narrowly at tip blackish; T
<subScript attach="left" box="[582,590,1495,1509]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
, T
<subScript attach="left" box="[621,629,1495,1509]" fontSize="6" pageId="56" pageNumber="57">3</subScript>
, sometimes F
<subScript attach="left" box="[786,794,1495,1509]" fontSize="6" pageId="56" pageNumber="57">3</subScript>
vaguely on dorsum, and posterior tarsi somewhat darker. Coxae with pale hairs and bristles. F
<subScript attach="left" box="[563,571,1534,1548]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
with erect pv hairs not longer than limb is deep. F
<subScript attach="left" box="[1139,1147,1534,1548]" fontSize="6" pageId="56" pageNumber="57">2</subScript>
slightly distorted, weakly concave in front about 0.3 from base where ad series of fine bristles almost as long as limb is deep. F
<subScript attach="left" box="[1336,1344,1573,1587]" fontSize="6" pageId="56" pageNumber="57">3</subScript>
(
<figureCitation box="[1357,1426,1557,1581]" captionStart="FIGURES 1–4" captionStartId="4.[151,263,1949,1972]" captionTargetBox="[159,1427,658,1918]" captionTargetId="figure-314@4.[151,1436,652,1928]" captionTargetPageId="4" captionText="FIGURES 1–4. Morphology of Hybos spp. 1. H. bispinatus sp. nov. ♀ habitus. 2. H. khamfui sp. nov. ♂ habitus. 3. H. serratus Yang&amp; Yang♂ right posterior femur, ventral view. 4. H. lannaensis sp. nov. ♀ postpedicel and stylus. aps = anterior proclinate setae, pvf = posteroventral fringe, vs = ventral spines." figureDoi="http://doi.org/10.5281/zenodo.6329813" httpUri="https://zenodo.org/record/6329813/files/figure.png" pageId="56" pageNumber="57">Fig. 3</figureCitation>
) moderately inflated, widest 0.6–0.7 from base; viewed laterally upper margin convex, lower margin linear; ventral spines black, comprising ~7–8 strong bristles (some of which on proximal 0.7 are slightly longer than limb is deep) behind which are ~10–12 mostly shorter and more evenly sized bristles, occasionally a few extra bristles between the rows especially near base; pv fringe with ~3 strong black bristles on distal 0.3, otherwise weakly developed and yellowish; 3–4 black curved anterior proclinate bristles on distal 0.5. T
<subScript attach="left" box="[949,957,1756,1770]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
usually with fine but distinct hair dorsally at 0.6 and 1 somewhat longer seta anteroapically in apical circlet, otherwise short-haired. T
<subScript attach="left" box="[1200,1208,1794,1808]" fontSize="6" pageId="56" pageNumber="57">2</subScript>
with strong bristles dorsally at ~0.2 and ~0.4–0.5 from base and 1 rather longer bristle ventrally at ~0.5; apical circlet with 1 av bristle much longer than others. T
<subScript attach="left" box="[452,460,1869,1883]" fontSize="6" pageId="56" pageNumber="57">3</subScript>
somewhat inflated distally, ~1.5–2.0X wide at tip than at base; 1 strong bristle dorsally ~0.5–0.6 from base; apical circlet with 3 distinct bristles. MT
<subScript attach="left" box="[843,851,1908,1922]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
and MT
<subScript attach="left" box="[948,956,1908,1922]" fontSize="6" pageId="56" pageNumber="57">2</subScript>
with distinct ventrobasal bristle; also MT
<subScript attach="left" box="[1428,1436,1908,1922]" fontSize="6" pageId="56" pageNumber="57">2</subScript>
with distinct ventral bristle near middle.
<emphasis box="[622,680,1932,1955]" italics="true" pageId="56" pageNumber="57">Wing</emphasis>
membrane tinged yellowish brown, veins brown; stigma distinct, reaching costa at ≥0.9 distance between end of R
<subScript attach="left" box="[732,740,1983,1997]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
and R
<subScript attach="left" box="[821,845,1983,1997]" fontSize="6" pageId="56" pageNumber="57">2+3</subScript>
. Squamae with pale hairs. Halter white.
<emphasis box="[1330,1435,1968,1992]" italics="true" pageId="56" pageNumber="57">Abdomen</emphasis>
subshining brownish black, tergites with faintly bronze reflections, tergites 2 and 3 with long pale to brownish bristles on posterior margin, distal segments with shorter hairs. Terminalia (
<figureCitation box="[1001,1156,151,175]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Figs 141–147</figureCitation>
) black, with black setae; left epandrial lamella (
<figureCitation box="[405,495,188,212]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Fig 142</figureCitation>
) with strong bristles; left surstylus (
<figureCitation box="[901,1057,188,212]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Figs 145–147</figureCitation>
) long, narrow, wider subapically; right surstylus long, narrow, serrate apically (
<figureCitation box="[663,762,223,247]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Fig. 144</figureCitation>
); hypandrium (
<figureCitation box="[934,1030,223,247]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Fig. 143</figureCitation>
) with 2 broad apical lobes.
<emphasis box="[1345,1429,224,248]" italics="true" pageId="57" pageNumber="58">Female</emphasis>
. Similar to male but F
<subScript attach="left" box="[405,413,275,289]" fontSize="6" pageId="57" pageNumber="58">3</subScript>
much less strongly inflated, widest ~0.8–0.9 from base; ventral spines comprising 7–8 distinct bristles arranged in single row; pv fringe weaker; usually only 1 distinct proclinate bristle anteriorly near tip. Abdomen with much shorter hairs. Terminalia (
<figureCitation box="[748,907,335,359]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Figs 148–150</figureCitation>
) black, tergite 8 tubular, encircling abdomen; sternite 8 with usually more strongly sclerotized posteroapical lateral extensions, appearing broadly T-shaped in posterior view (
<figureCitation box="[329,428,407,431]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Fig. 149</figureCitation>
), extending dorsally in lateral view (
<figureCitation box="[845,944,407,431]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58">Fig. 150</figureCitation>
); sometimes posteroapical lateral processes not easily discernable, being weakly sclerotized or with area between them and median part of sternite 8 equally strongly sclerotized.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6329870" ID-Zenodo-Dep="6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,264,1386,1409]" targetBox="[160,1422,578,1348]" targetPageId="57">
<paragraph blockId="57.[151,1437,1386,1501]" pageId="57" pageNumber="58">
<emphasis bold="true" box="[151,370,1386,1409]" pageId="57" pageNumber="58">FIGURES 141–150.</emphasis>
<taxonomicName authority="Yang &amp; Yang." authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[381,706,1386,1409]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[381,540,1387,1409]" italics="true" pageId="57" pageNumber="58">Hybos serratus</emphasis>
Yang &amp; Yang.
</taxonomicName>
<emphasis bold="true" box="[716,761,1386,1409]" pageId="57" pageNumber="58">141.</emphasis>
♂ terminalia, dorsal view.
<emphasis bold="true" box="[1062,1106,1386,1409]" pageId="57" pageNumber="58">142.</emphasis>
left epandrial lamella and left surstylus, lateral view.
<emphasis bold="true" box="[396,440,1417,1440]" pageId="57" pageNumber="58">143.</emphasis>
hypandrium, ventral view.
<emphasis bold="true" box="[735,780,1417,1440]" pageId="57" pageNumber="58">144.</emphasis>
apex of right surstylus.
<emphasis bold="true" box="[1045,1140,1417,1440]" pageId="57" pageNumber="58">145–147.</emphasis>
apex of left surstylus.
<emphasis bold="true" box="[1392,1436,1417,1440]" pageId="57" pageNumber="58">145.</emphasis>
specimen from Doi Chiang Dao.
<emphasis bold="true" box="[490,534,1447,1470]" pageId="57" pageNumber="58">146.</emphasis>
specimen from Doi Phahompok.
<emphasis bold="true" box="[878,923,1447,1470]" pageId="57" pageNumber="58">147.</emphasis>
specimen from Phu Ruea.
<emphasis bold="true" box="[1199,1295,1447,1470]" pageId="57" pageNumber="58">148–150.</emphasis>
♀ terminalia.
<emphasis bold="true" box="[151,195,1478,1501]" pageId="57" pageNumber="58">148.</emphasis>
ventral view.
<emphasis bold="true" box="[338,383,1478,1501]" pageId="57" pageNumber="58">149.</emphasis>
posterior view.
<emphasis bold="true" box="[545,590,1478,1501]" pageId="57" pageNumber="58">150.</emphasis>
lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8.
</paragraph>
</caption>
<subSubSection lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="discussion">
<paragraph blockId="57.[151,1436,1539,2033]" pageId="57" pageNumber="58">
<emphasis bold="true" box="[199,324,1539,1564]" pageId="57" pageNumber="58">Comment.</emphasis>
<taxonomicName authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[331,501,1540,1564]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[331,501,1540,1564]" italics="true" pageId="57" pageNumber="58">Hybos serratus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[704,867,1540,1564]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aceriformis" status="sp. nov.">
<emphasis box="[704,867,1540,1564]" italics="true" pageId="57" pageNumber="58">H. aceriformis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[874,964,1540,1564]" pageId="57" pageNumber="58">
<taxonomicNameLabel box="[874,964,1540,1564]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from which is is best distinguished by the form of the male and female terminalia. It might also be confused with
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[949,1135,1577,1600]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="anisoserratus" status="sp. nov.">
<emphasis box="[949,1135,1577,1600]" italics="true" pageId="57" pageNumber="58">H. anisoserratus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1142,1232,1576,1600]" pageId="57" pageNumber="58">
<taxonomicNameLabel box="[1142,1232,1576,1600]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
which is however a smaller species, usually with more extensively yellow legs.
</paragraph>
<paragraph blockId="57.[151,1436,1539,2033]" pageId="57" pageNumber="58">
Although described from
<collectingRegion box="[493,692,1648,1672]" country="China" name="Sichuan" pageId="57" pageNumber="58">Sichuan Province</collectingRegion>
<collectingCountry box="[701,768,1648,1672]" name="China" pageId="57" pageNumber="58">China</collectingCountry>
(
<bibRefCitation author="Yang, D. &amp; Yang, C. K." box="[784,1007,1648,1673]" pageId="57" pageNumber="58" pagination="1089 - 1097" refId="ref82013" refString="Yang, D. &amp; Yang, C. K. (1992) Diptera: Empididae. In: Chen, S. (Ed.), Insects of the Hengduan Mountains Region 2. Science Press, Beijing, pp. 1089 - 1097." type="book chapter" year="1992">Yang &amp; Yang 1992</bibRefCitation>
)
<taxonomicName authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[1025,1153,1649,1672]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[1025,1153,1649,1672]" italics="true" pageId="57" pageNumber="58">H. serratus</emphasis>
</taxonomicName>
is widespread in eastern
<collectingCountry box="[151,221,1684,1708]" name="China" pageId="57" pageNumber="58">China</collectingCountry>
, including
<collectingRegion box="[349,425,1684,1707]" country="China" name="Henan" pageId="57" pageNumber="58">Henan</collectingRegion>
,
<collectingRegion box="[438,537,1684,1708]" country="China" name="Guizhou" pageId="57" pageNumber="58">Guizhou</collectingRegion>
,
<collectingRegion box="[550,649,1684,1708]" country="China" name="Guangxi" pageId="57" pageNumber="58">Guangxi</collectingRegion>
,
<collectingRegion box="[663,744,1684,1708]" country="China" name="Hainan" pageId="57" pageNumber="58">Hainan</collectingRegion>
and also occurs in
<collectingCountry box="[970,1065,1684,1708]" name="Vietnam" pageId="57" pageNumber="58">Vietnam</collectingCountry>
(
<bibRefCitation author="Yang, D. &amp; Yang, C." box="[1081,1307,1684,1708]" pageId="57" pageNumber="58" pagination="1 - 335" refId="ref82158" refString="Yang, D. &amp; Yang, C. (2004) Diptera Empididae: Hemerodromiinae, Hybotinae. Fauna Sinica, Insecta, 34, i - x, 1 - 335." type="journal article" year="2004">Yang &amp; Yang 2004</bibRefCitation>
;
<bibRefCitation author="Huo, S. &amp; Grootaert, P. &amp; Yang, D." pageId="57" pageNumber="58" pagination="47 - 55" refId="ref80575" refString="Huo, S., Grootaert, P. &amp; Yang, D. (2010) Two new yellow-legged species of the genus Hybos from Vietnam (Diptera: Empidoidea: Hybotinae). Zootaxa, 2512, 47 - 55." type="journal article" year="2010">
Huo
<emphasis box="[1378,1429,1684,1708]" italics="true" pageId="57" pageNumber="58">et al</emphasis>
. 2010
</bibRefCitation>
;
<bibRefCitation author="Yang, D. &amp; Wang, M. &amp; Zhu, Y. &amp; Zhang, L." box="[222,412,1720,1745]" pageId="57" pageNumber="58" refId="ref81809" refString="Yang, D., Wang, M., Zhu, Y. &amp; Zhang, L. (2010) Insect Fauna of Henan, Diptera: Empidoidea. Science Press, Beijing, 418 pp." type="book" year="2010">
Yang
<emphasis box="[287,337,1720,1744]" italics="true" pageId="57" pageNumber="58">et al</emphasis>
. 2010
</bibRefCitation>
). The species is here recorded from
<collectingCountry box="[819,918,1720,1744]" name="Thailand" pageId="57" pageNumber="58">Thailand</collectingCountry>
for the first time where it is widespread north of ~
<geoCoordinate box="[204,266,1756,1780]" degrees="16" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="55555" value="16.0">16°N</geoCoordinate>
, occurring as far south as the northern Tenasserim Hills and Petchabun Range (
<figureCitation box="[1163,1259,1756,1780]" captionStart="FIGURES 269–277" captionStartId="89.[151,264,1945,1968]" captionTargetBox="[244,1325,201,1898]" captionTargetId="figure-80@89.[219,1367,165,1924]" captionTargetPageId="89" captionText="FIGURES 269–277. Distribution of Hybos spp. in Thailand plotted at a resolution of 1° of latitude and longitude. 269. H. serratus Yang &amp; Yang. 270. H. shamshevi sp. nov. 271. H. sinclairi sp. nov. 272. H. songbai sp. nov. 273. H. steatopygus sp. nov. 274. H. stigmaticus sp. nov. 275. H. subapicalis sp. nov. 276. H. tetricus sp. nov. 277. H. thaosaeo sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6329911" httpUri="https://zenodo.org/record/6329911/files/figure.png" pageId="57" pageNumber="58">Fig. 269</figureCitation>
) but apparently absent from middle and southern parts of the country. In
<collectingCountry box="[785,887,1792,1816]" name="Thailand" pageId="57" pageNumber="58">Thailand</collectingCountry>
,
<taxonomicName authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[899,1024,1793,1816]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[899,1024,1793,1816]" italics="true" pageId="57" pageNumber="58">H. serratus</emphasis>
</taxonomicName>
occurs at elevations of
<quantity box="[1288,1436,1792,1816]" metricMagnitude="3" metricUnit="m" metricValue="1.3205000000000002" metricValueMax="2.1740000000000004" metricValueMin="0.46699999999999997" pageId="57" pageNumber="58" unit="m" value="1320.5" valueMax="2174.0" valueMin="467.0">467–2,174 m</quantity>
but is most abundant in evergreen and other moist forest
<typeStatus box="[818,877,1829,1853]" pageId="57" pageNumber="58">types</typeStatus>
above ~
<quantity box="[978,1077,1828,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="57" pageNumber="58" unit="m" value="1200.0">1,200 m</quantity>
. Adults have been found in all months except January and March (
<figureCitation box="[551,646,1864,1888]" captionStart="FIGURES 291–298" captionStartId="93.[151,263,1912,1935]" captionTargetBox="[229,1367,185,1880]" captionTargetId="figure-80@93.[189,1403,163,1912]" captionTargetPageId="93" captionText="FIGURES 291–298. Phenology of Hybos spp. monthly abundance (as % of total number of individuals captured) is plotted against month of capture. 291. H. inthanonensis sp. nov. 292. H. kaluang sp. nov. 293. H. khamfui sp. nov. 294. H. particularis Yang, Yang &amp; Hu. 295. H. pisadaanus sp. nov. 296. H. saenmueangmai sp. nov. 297. H. serratus Yang &amp; Yang. 298. H. songbai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6329917" httpUri="https://zenodo.org/record/6329917/files/figure.png" pageId="57" pageNumber="58">Fig. 297</figureCitation>
) but adult numbers are clearly correlated with the wet season between late May and December becoming maximal in December and October.
</paragraph>
<paragraph blockId="57.[151,1436,1539,2033]" lastBlockId="58.[151,1436,151,392]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<collectingCountry box="[199,298,1936,1960]" name="Thailand" pageId="57" pageNumber="58">Thailand</collectingCountry>
specimens apparently differ slightly from examples from
<collectingCountry box="[984,1051,1936,1960]" name="China" pageId="57" pageNumber="58">China</collectingCountry>
and
<collectingCountry box="[1114,1209,1936,1960]" name="Vietnam" pageId="57" pageNumber="58">Vietnam</collectingCountry>
in having the hind femur only at most vaguely darkened dorsally but with a blackish tip. Also both left and right surstyli of
<collectingCountry box="[1337,1436,1972,1996]" name="Thailand" pageId="57" pageNumber="58">Thailand</collectingCountry>
specimens are apparently somewhat narrower than ‘typical’ (cf.
<bibRefCitation author="Yang, D. &amp; Yang, C." box="[898,1126,2008,2033]" pageId="57" pageNumber="58" pagination="1 - 335" refId="ref82158" refString="Yang, D. &amp; Yang, C. (2004) Diptera Empididae: Hemerodromiinae, Hybotinae. Fauna Sinica, Insecta, 34, i - x, 1 - 335." type="journal article" year="2004">Yang &amp; Yang 2004</bibRefCitation>
, figs 372–373;
<bibRefCitation author="Yang, D. &amp; Wang, M. &amp; Zhu, Y. &amp; Zhang, L." lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" refId="ref81809" refString="Yang, D., Wang, M., Zhu, Y. &amp; Zhang, L. (2010) Insect Fauna of Henan, Diptera: Empidoidea. Science Press, Beijing, 418 pp." type="book" year="2010">
Yang
<emphasis box="[1377,1429,2008,2032]" italics="true" pageId="57" pageNumber="58">et al</emphasis>
. 2010
</bibRefCitation>
, fig. 200) but I have little doubt that they are
<taxonomicName authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[725,851,153,176]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[725,851,153,176]" italics="true" pageId="58" pageNumber="59">H. serratus</emphasis>
</taxonomicName>
. Indeed, there are small differences in characters of the male terminalia within
<collectingCountry box="[470,569,188,212]" name="Thailand" pageId="58" pageNumber="59">Thailand</collectingCountry>
and furthermore, these differences are consistently present in populations present on disjunct mountain ranges in the country. For example, the apex of the left surstylus is abruptly narrowed apically in specimens from Doi Chiang Dao (
<figureCitation box="[677,775,260,284]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="58" pageNumber="59">Fig. 145</figureCitation>
) but of different form in examples from Doi Phahompok (
<figureCitation box="[159,257,295,319]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="58" pageNumber="59">Fig. 146</figureCitation>
) and from Phu Reua (
<figureCitation box="[516,617,295,319]" captionStart="FIGURES 141–150" captionStartId="57.[151,264,1386,1409]" captionTargetBox="[160,1422,578,1348]" captionTargetId="figure-482@57.[151,1435,556,1365]" captionTargetPageId="57" captionText="FIGURES 141–150. Hybos serratus Yang &amp; Yang. 141. ♂ terminalia, dorsal view. 142. left epandrial lamella and left surstylus, lateral view. 143. hypandrium, ventral view. 144. apex of right surstylus. 145–147. apex of left surstylus. 145. specimen from Doi Chiang Dao. 146. specimen from Doi Phahompok. 147. specimen from Phu Ruea. 148–150. ♀ terminalia. 148. ventral view. 149. posterior view. 150. lateral view. RS = right surstylus, S8= sternite 8, T8 = tergite 8." figureDoi="http://doi.org/10.5281/zenodo.6329870" httpUri="https://zenodo.org/record/6329870/files/figure.png" pageId="58" pageNumber="59">Fig. 147</figureCitation>
). It is hypothesised that these differences in morphology of the surstyli reflect recent and ongoing active infra-specific radiation in isolated populations.
<taxonomicName authorityName="Yang &amp; Yang, 1992:" authorityYear="1089" box="[1065,1236,332,356]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="serratus">
<emphasis box="[1065,1236,332,356]" italics="true" pageId="58" pageNumber="59">Hybos serratus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[151,337,368,392]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="anisoserratus" status="sp. nov.">
<emphasis box="[151,337,368,392]" italics="true" pageId="58" pageNumber="59">H. anisoserratus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[344,434,368,392]" pageId="58" pageNumber="59">
<taxonomicNameLabel box="[344,434,368,392]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in which species account the differences are further discussed.
</paragraph>
</subSubSection>
</treatment>
</document>
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