treatments-xml/data/BD/66/2E/BD662E6FFF91FFDAFC5D80DF828AF8A5.xml

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<document id="0852741FA11286F0E61650BDB95E2966" ID-ISSN="1536-9307" ID-ZooBank="D9F2DCCC-E054-4DB8-9695-5715677C8CDC" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638249738338" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Plíšková, Jana &amp; Šťáhlavský, František" docDate="2016" docId="BD662E6FFF91FFDAFC5D80DF828AF8A5" docLanguage="en" docName="Euscorpius.226.1-21.pdf" docOrigin="Euscorpius 226" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Compsobuthus somalilandus Kovarik 2012" docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="415F5617FF9DFFCAFFDD854C8625FFCB" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n." masterLastPageNumber="21" masterPageNumber="1" pageNumber="11" updateTime="1699435298193" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="642DFBB19E9130D8E17BEFE09FF3C663">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n.</mods:title>
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<paragraph id="35709F79FF91FFC6FC5D80DF837BFA61" blockId="12.[896,1374,1427,1481]" box="[896,1374,1427,1450]" pageId="12" pageNumber="11">
<heading id="6E382815FF91FFC6FC5D80DF837BFA61" box="[896,1374,1427,1450]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="11" reason="4">
<taxonomicName id="F2CFE4FAFF91FFC6FC5D80DF837BFA61" ID-CoL="XHGG" authority="Kovarik, 2012" authorityName="Kovarik" authorityYear="2012" box="[896,1374,1427,1450]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="07BB436BFF91FFC6FC5D80DF829BFA61" bold="true" box="[896,1214,1427,1450]" italics="true" pageId="12" pageNumber="11">Compsobuthus somalilandus</emphasis>
Kovařík, 2012
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</paragraph>
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<paragraph id="35709F79FF91FFC6FC1E80FE833EFA02" blockId="12.[896,1374,1427,1481]" box="[963,1307,1458,1481]" pageId="12" pageNumber="11">
(
<figureCitation id="ADF483FCFF91FFC6FC1180FE8268FA02" box="[972,1101,1458,1481]" captionStart="Figures 918" captionStartId="7.[192,278,1590,1613]" captionTargetBox="[191,1437,215,1561]" captionTargetId="figure-6@7.[192,1437,216,1560]" captionTargetPageId="7" captionText="Figures 918: Figures 914. Compsobuthus eritreaensis sp. n. Figures 9, 11, 13. Male holotype, carapace and tergites IIII (9), coxosternal area and sternites (11), and lateral view of telson (13). Figures 10, 12, 14. Female paratype, carapace and tergites IIII (10), coxosternal area and sternites (12), and lateral view of telson (14). Figures 1516. C. abyssinicus, lateral view of telson of male (15) and female (16) from locality 11EW. Figures 1718. C. somalilandus, lateral view of telson of paratype male (17) and female (18) from type locality." pageId="12" pageNumber="11">Figs. 1718</figureCitation>
,
<figureCitation id="ADF483FCFF91FFC6FB8A80FE82BFFA02" box="[1111,1178,1458,1481]" captionStart="Figures 1938" captionStartId="8.[192,278,1710,1733]" captionTargetBox="[200,1430,225,1668]" captionTargetId="figure-12@8.[192,1439,213,1680]" captionTargetPageId="8" captionText="Figures 1938: Figures 1922. Compsobuthus abyssinicus, from locality 11EA. Figures 1920. Female, pedipalp chela dorsal (19) and external (20). Figures 2122. Male, pedipalp chela dorsal (21) and external (22). Figures 2326. C. somalilandus, from type locality. Figures 2324. Female paratype, pedipalp chela dorsal (23) and external (24). Figures 2526. Male holotype, pedipalp chela dorsal (25) and external (26). Figures 2738. Compsobuthus eritreaensis sp. n. Figures 2728. Female paratype from type locality, pedipalp chela dorsal (27) and external (28). Figures 2936. Male holotype from type locality, pedipalp segments. Chela dorsal (29), external (30), and ventral (31). Patella dorsal (32), external (33) and ventral (34). Femur dorsal (35). Movable finger dentition (36). The trichobothrial pattern is indicated in Figures 3033 and 35. Figures 3738. Male holotype, right legs III (37) and IV (38), retrolateral aspect." pageId="12" pageNumber="11">2326</figureCitation>
,
<figureCitation id="ADF483FCFF91FFC6FB7A80FE82CFFA02" box="[1191,1258,1458,1481]" captionStart="Figure 6162" captionStartId="16.[192,268,1757,1780]" captionTargetBox="[191,1440,215,1722]" captionTargetId="figure-12@16.[192,1439,216,1721]" captionTargetPageId="16" captionText="Figure 6162: Compsobuthus somalilandus, male paratype in vivo habitus at type locality (61) and the type locality (62)." pageId="12" pageNumber="11">6162</figureCitation>
,
<figureCitation id="ADF483FCFF91FFC6FB2A80FE8337FA02" box="[1271,1298,1458,1481]" captionStart="Figure 77" captionStartId="19.[192,268,1203,1226]" captionTargetBox="[201,1431,152,1168]" captionTargetId="figure-6@19.[192,1437,144,1173]" captionTargetPageId="19" captionText="Figure 77: Map showing the distribution of Compsobuthus in Eritrea, Ethiopia and Somaliland. Marks indicate sites checked during 20112016 expeditions (except for C. abyssinicus from Djibouti which is verified record)." pageId="12" pageNumber="11">77</figureCitation>
)
</paragraph>
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<taxonomicName id="F2CFE4FAFF91FFC6FCE380A3837AF9CD" ID-CoL="XHGG" authority="Kovarik, 2012: 7 - 8" authorityName="Kovarik" authorityPageNumber="7 - 8" authorityYear="2012" box="[830,1375,1519,1542]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="07BB436BFF91FFC6FCE380A38252F9CD" box="[830,1143,1519,1542]" italics="true" pageId="12" pageNumber="11">Compsobuthus somalilandus</emphasis>
Kovařík, 2012: 78
</taxonomicName>
, figs. 819, 3742; Kovařík &amp; Ojanguren, 2013: 157, figs. 783794, 926931.
</paragraph>
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<collectionCode id="53DE07BCFF91FFC6FC5A8321820AF94B" box="[903,1071,1645,1664]" pageId="12" pageNumber="11">LOCALITY AND</collectionCode>
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<collectionCode id="53DE07BCFF91FFC6FB588321832DF94B" box="[1157,1288,1645,1664]" pageId="12" pageNumber="11">REPOSITORY</collectionCode>
. Somaliland, near Berbera,
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a.s.l.
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; first authors collection
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(
<collectionCode id="53DE07BCFF91FFC6FB9B83EB82AAF975" box="[1094,1167,1703,1726]" pageId="12" pageNumber="11">FKCP</collectionCode>
).
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<collectionCode id="53DE07BCFF91FFC6FCE383A88249F930" box="[830,1132,1764,1787]" pageId="12" pageNumber="11">TYPE MATERIAL EXAMINED</collectionCode>
. Somaliland, near Berbera,
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,
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(
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),
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<collectingDate id="51354051FF91FFC6FABD824F8556F8F2" pageId="12" pageNumber="11" value="2011-07-09">9.VII. 2011</collectingDate>
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,
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<specimenCount id="23C954F0FF91FFC6FC7A826E85EDF8F2" box="[935,968,1826,1849]" pageId="12" pageNumber="11" type="female">4♀</specimenCount>
(
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and
<typeStatus id="EA7421DBFF91FFC6FBB6826E82F1F8F2" box="[1131,1236,1826,1849]" pageId="12" pageNumber="11" type="paratype">paratypes</typeStatus>
,
<figureCitation id="ADF483FCFF91FFC6FB39826E834DF8F2" box="[1252,1384,1826,1849]" captionStart="Figures 918" captionStartId="7.[192,278,1590,1613]" captionTargetBox="[191,1437,215,1561]" captionTargetId="figure-6@7.[192,1437,216,1560]" captionTargetPageId="7" captionText="Figures 918: Figures 914. Compsobuthus eritreaensis sp. n. Figures 9, 11, 13. Male holotype, carapace and tergites IIII (9), coxosternal area and sternites (11), and lateral view of telson (13). Figures 10, 12, 14. Female paratype, carapace and tergites IIII (10), coxosternal area and sternites (12), and lateral view of telson (14). Figures 1516. C. abyssinicus, lateral view of telson of male (15) and female (16) from locality 11EW. Figures 1718. C. somalilandus, lateral view of telson of paratype male (17) and female (18) from type locality." pageId="12" pageNumber="11">Figs. 1718</figureCitation>
,
<figureCitation id="ADF483FCFF91FFC6FAA5826E857CF89C" captionStart="Figures 1938" captionStartId="8.[192,278,1710,1733]" captionTargetBox="[200,1430,225,1668]" captionTargetId="figure-12@8.[192,1439,213,1680]" captionTargetPageId="8" captionText="Figures 1938: Figures 1922. Compsobuthus abyssinicus, from locality 11EA. Figures 1920. Female, pedipalp chela dorsal (19) and external (20). Figures 2122. Male, pedipalp chela dorsal (21) and external (22). Figures 2326. C. somalilandus, from type locality. Figures 2324. Female paratype, pedipalp chela dorsal (23) and external (24). Figures 2526. Male holotype, pedipalp chela dorsal (25) and external (26). Figures 2738. Compsobuthus eritreaensis sp. n. Figures 2728. Female paratype from type locality, pedipalp chela dorsal (27) and external (28). Figures 2936. Male holotype from type locality, pedipalp segments. Chela dorsal (29), external (30), and ventral (31). Patella dorsal (32), external (33) and ventral (34). Femur dorsal (35). Movable finger dentition (36). The trichobothrial pattern is indicated in Figures 3033 and 35. Figures 3738. Male holotype, right legs III (37) and IV (38), retrolateral aspect." pageId="12" pageNumber="11">23 26</figureCitation>
,
<figureCitation id="ADF483FCFF91FFC6FCB0820C85ADF89C" box="[877,904,1856,1879]" captionStart="Figure 6162" captionStartId="16.[192,268,1757,1780]" captionTargetBox="[191,1440,215,1722]" captionTargetId="figure-12@16.[192,1439,216,1721]" captionTargetPageId="16" captionText="Figure 6162: Compsobuthus somalilandus, male paratype in vivo habitus at type locality (61) and the type locality (62)." pageId="12" pageNumber="11">61</figureCitation>
), leg.
<collectorName id="983AFAAFFF91FFC6FC07820C8270F89C" box="[986,1109,1856,1879]" pageId="12" pageNumber="11">F. Kovařík</collectorName>
; near Berbera,
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,
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a.s.l.
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,
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,
<specimenCount id="23C954F0FF91FFC6FAC682138319F8BD" box="[1307,1340,1887,1910]" pageId="12" pageNumber="11" type="female">1♀</specimenCount>
, leg.
<collectorName id="983AFAAFFF91FFC4FA588213873DF8D0" lastPageId="14" lastPageNumber="13" pageId="12" pageNumber="11">F. Kovařík</collectorName>
; near Sheikh, foothills of
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,
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<geoCoordinate id="50FBF9BEFF93FFC4FED7826E87B4F8F2" box="[266,401,1826,1849]" degrees="45" direction="east" minutes="09.762" orientation="longitude" pageId="14" pageNumber="13" precision="1" value="45.1627">45°09.762'E</geoCoordinate>
,
<elevation id="BEE2784AFF93FFC4FE43826E8404F8F2" box="[414,545,1826,1849]" metricMagnitude="2" metricUnit="m" metricValue="8.959999999999999" pageId="14" pageNumber="13" unit="m" value="896.0">
<quantity id="F237329CFF93FFC4FE43826E87C0F8F2" box="[414,485,1826,1849]" metricMagnitude="2" metricUnit="m" metricValue="8.959999999999999" pageId="14" pageNumber="13" unit="m" value="896.0">896 m</quantity>
a.s.l.
</elevation>
,
<specimenCount id="23C954F0FF93FFC4FDED826E8477F8F2" box="[560,594,1826,1849]" pageId="14" pageNumber="13" type="female">2♀</specimenCount>
(
<typeStatus id="EA7421DBFF93FFC4FD84826E8494F8F2" box="[601,689,1826,1849]" pageId="14" pageNumber="13" type="allotype">allotype</typeStatus>
and
<typeStatus id="EA7421DBFF93FFC4FD36826E86CBF893" pageId="14" pageNumber="13" type="paratype">paratype</typeStatus>
),
<date id="4171B9B9FF93FFC4FED5820D8744F893" box="[264,353,1857,1880]" pageId="14" pageNumber="13" value="2010-11">
<collectingDate id="51354051FF93FFC4FED5820D8744F893" box="[264,353,1857,1880]" pageId="14" pageNumber="13" value="2010-11">XI.2010</collectingDate>
</date>
, leg
</materialsCitation>
.
<materialsCitation id="85A79524FF93FFC4FE40820D843FF8BC" collectionCode="T" collectorName="P. Novak. All" pageId="14" pageNumber="13" specimenCount="1" typeStatus="holotype">
<collectionCode id="53DE07BCFF93FFC4FE40820D8795F893" box="[413,432,1857,1880]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="13">T</collectionCode>
. Mazuch and
<collectorName id="983AFAAFFF93FFC4FD97820D84C5F893" box="[586,736,1857,1880]" pageId="14" pageNumber="13">P. Novák. All</collectorName>
types are in the first authors collection
</materialsCitation>
(
<collectionCode id="53DE07BCFF93FFC4FDF5822C8454F8BC" box="[552,625,1888,1911]" pageId="14" pageNumber="13">FKCP</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="61B0CFF1FF90FFC7FF1D824285AFF8F5" pageId="13" pageNumber="12" startId="13.[192,278,1806,1829]" targetBox="[191,1440,214,1777]" targetPageId="13">
<paragraph id="35709F79FF90FFC7FF1D824285AFF8F5" blockId="13.[192,1440,1806,1854]" pageId="13" pageNumber="12">
<emphasis id="07BB436BFF90FFC7FF1D8242874DF8EE" bold="true" box="[192,360,1806,1829]" pageId="13" pageNumber="12">Figures 5556:</emphasis>
<taxonomicName id="F2CFE4FAFF90FFC7FEAD825C8453F8EE" authority="Kovařík &amp; Lowe &amp; Plíšková &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Plíšková &amp; Šťáhlavský" authorityYear="2016" box="[368,630,1808,1829]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="eritreaensis" status="sp. nov.">
<emphasis id="07BB436BFF90FFC7FEAD825C8453F8EE" box="[368,630,1808,1829]" italics="true" pageId="13" pageNumber="12">Compsobuthus eritreaensis</emphasis>
</taxonomicName>
<emphasis id="07BB436BFF90FFC7FDA3825C8491F8EE" bold="true" box="[638,692,1808,1829]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="1C88FE10FF90FFC7FDA3825C8491F8EE" box="[638,692,1808,1829]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female paratype with newborns from type locality (55) and the type locality 15EI, Eritrea, near Massawa, 15°36'58.7&quot;N 39°22'32.8&quot;E, 74 m a.s.l. (56).
</paragraph>
</caption>
<caption id="61B0CFF1FF93FFC4FF1D83E28226F915" pageId="14" pageNumber="13" startId="14.[192,278,1710,1733]" targetBox="[191,1440,192,1681]" targetPageId="14">
<paragraph id="35709F79FF93FFC4FF1D83E28226F915" blockId="14.[192,1440,1710,1758]" pageId="14" pageNumber="13">
<emphasis id="07BB436BFF93FFC4FF1D83E28427F90E" bold="true" box="[192,514,1710,1733]" pageId="14" pageNumber="13">Figures 5759: Figures 5758</emphasis>
.
<taxonomicName id="F2CFE4FAFF93FFC4FDD283FC8530F90E" authority="Kovařík &amp; Lowe &amp; Plíšková &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Plíšková &amp; Šťáhlavský" authorityYear="2016" box="[527,789,1712,1733]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="eritreaensis" status="sp. nov.">
<emphasis id="07BB436BFF93FFC4FDD283FC8530F90E" box="[527,789,1712,1733]" italics="true" pageId="14" pageNumber="13">Compsobuthus eritreaensis</emphasis>
</taxonomicName>
<emphasis id="07BB436BFF93FFC4FCC383FC8571F90E" bold="true" box="[798,852,1712,1733]" pageId="14" pageNumber="13">
<taxonomicNameLabel id="1C88FE10FF93FFC4FCC383FC8571F90E" box="[798,852,1712,1733]" pageId="14" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratypes in vivo habitus at locality 15EJ. Female (57) and male (58).
<emphasis id="07BB436BFF93FFC4FEF4838587AEF915" bold="true" box="[297,395,1737,1758]" pageId="14" pageNumber="13">Figure 59</emphasis>
. Locality 15EJ, Eritrea, Dese Island, 15°26'39.2&quot;N 39°45'32.7&quot;E.
</paragraph>
</caption>
<subSubSection id="7DD5CCF2FF93FFDAFCE38248828AF8A5" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13" type="diagnosis">
<paragraph id="35709F79FF93FFDAFCE38248828AF8A5" blockId="14.[830,1441,1796,1911]" lastBlockId="16.[830,1441,1818,1902]" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
DIAGNOSIS. Total length
<quantity id="F237329CFF93FFC4FB90824882E5F8D0" box="[1101,1216,1796,1819]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="14" pageNumber="13" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">2832 mm</quantity>
. Sexual dimorphism minor, adult males with chela of pedipalps broader and fingers of pedipalps flexed proximally; there is no difference in length and width of metasomal segments. Base color uniformly yellow to yellowish brown, with dark spots. Movable finger of pedipalp bears 10 rows of granules, all without external and with internal accessory granules (
<taxonomicName id="F2CFE4FAFF92FFC5FEEB826287F0F88E" authorityName=": Sissom" authorityYear="1994" box="[310,469,1838,1861]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="07BB436BFF92FFC5FEEB826287F0F88E" box="[310,469,1838,1861]" italics="true" pageId="15" pageNumber="14">acutecarinatus</emphasis>
</taxonomicName>
group of Levy &amp; Amitai, 1980). Pedipalp chela length/width ratio 4.0 in males and
<quantity id="F237329CFF92FFC5FCAD839B8597F925" box="[880,946,1751,1774]" metricMagnitude="-1" metricUnit="m" metricValue="1.2192" pageId="15" pageNumber="14" unit="in" value="4.8">4.8 in</quantity>
females. Manus of chela shorter than fixed finger. Trochanter of pedipalps with one to twelve spinules and without setae. Anterior margin of carapace bears 8 symmetrically distributed setae. First to third metasomal segments bear 10 carinae, fourth bears 8 or 10 carinae. All metasomal segments longer than wide. Pectinal teeth number
<quantity id="F237329CFF8DFFDAFE1B82758409F89B" box="[454,556,1849,1872]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.334" metricValueMin="4.572" pageId="16" pageNumber="15" unit="in" value="19.5" valueMax="21.0" valueMin="18.0">1821 in</quantity>
males and
<quantity id="F237329CFF8DFFDAFD6182758507F89B" box="[700,802,1849,1872]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="16" pageNumber="15" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">1518 in</quantity>
females. Sternites and ventral surface of metasoma granulated. Seventh sternite bears four crenulate carinae. Telson bulbous, aculeus shorter than vesicle. Subaculear tubercle present, long and spinoid.
</paragraph>
</subSubSection>
</treatment>
</document>