treatments-xml/data/1A/37/87/1A3787F69D369277FDC0FEB6CB932ABC.xml

<document id="3D311C02E3E44AB1EF443C9DE8637AB3" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D369277FDC0FEB6CB932ABC" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius dendrolagus Telnov 2020, sp. nov." docType="treatment" docUuid="1B512828-B4EA-44C2-94CE-3C778DBB5C4C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="45" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="43" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0A6263FFBE62A5EBAEEB8E62A3D5AE94">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="5A8D21CBACEDC109DF83AB13D9CF62BB">Telnov, Dmitry</mods:namePart>
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<mods:title id="3BEFCBAE1E9EE54C5A3E5B92A5B45D7E">European Journal of Taxonomy</mods:title>
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<mods:date id="B3AC97847C526B4DC3D14971D1BE536B">2020</mods:date>
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<heading id="C969818C9D369271FDC0FEB6CBB62941" box="[595,992,264,291]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="43" reason="2">
<taxonomicName id="559E4D639D369271FDC0FEB6CB292941" ID-CoL="7X3WN" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[595,895,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="dendrolagus" status="sp. nov.">
<emphasis id="A0EAEAF29D369271FDC0FEB6CB292941" bold="true" box="[595,895,264,291]" italics="true" pageId="42" pageNumber="43">Maechidius dendrolagus</emphasis>
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<figureCitation id="0AA52A659D369271FD96FEEFCA082909" box="[517,606,336,363]" captionStart="Figs 35–37" captionStartId="43.[189,241,1928,1954]" captionTargetBox="[357,1227,273,1901]" captionTargetId="figure-54@43.[349,1238,265,1915]" captionTargetPageId="43" captionText="Figs 35–37. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 35. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 36. M. esau Heller, 1914 (lectotype ♂ of M. setosus Moser, 1920; ZMHB). 37. M. esau, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122140" httpUri="https://zenodo.org/record/4122140/files/figure.png" pageId="42" pageNumber="43">Figs 35</figureCitation>
, 
<figureCitation id="0AA52A659D369271FDF9FEEFCACE2909" box="[618,664,337,363]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="42" pageNumber="43">116</figureCitation>
, 
<figureCitation id="0AA52A659D369271FD37FEEFCA852909" box="[676,723,337,363]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="42" pageNumber="43">212</figureCitation>
, 
<figureCitation id="0AA52A659D369271FD4CFEEFCB582909" box="[735,782,337,363]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="42" pageNumber="43">302</figureCitation>
, 375, 469, 543, 611–613
</paragraph>
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<emphasis id="A0EAEAF29D369271FF2EFE2DC99F29CF" bold="true" box="[189,457,403,429]" pageId="42" pageNumber="43">Differential diagnosis</emphasis>
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<paragraph id="922136E09D369271FF2EFE7CCCD529BE" blockId="42.[189,1155,450,477]" box="[189,1155,450,477]" pageId="42" pageNumber="43">The new species differs from all congeners primarily in the shape of the aedeagus.</paragraph>
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<subSubSection id="DA84656B9D369271FF2EFDBACC3F2A13" pageId="42" pageNumber="43" type="etymology">
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<emphasis id="A0EAEAF29D369271FF2EFDBAC9122A7C" bold="true" box="[189,324,516,542]" pageId="42" pageNumber="43">Etymology</emphasis>
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The species is named after 
<emphasis id="A0EAEAF29D369271FD8FFD8DCAE22A2F" box="[540,692,563,589]" italics="true" pageId="42" pageNumber="43">Dendrolagus</emphasis>
S. Müller, 1840 (tree-kangaroo), an enigmatic genus of marsupial endemic to New 
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, adjacent islands and Australian Queensland.
</paragraph>
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<emphasis id="A0EAEAF29D369271FF2EFD27C93A2AD1" bold="true" box="[189,364,665,691]" pageId="42" pageNumber="43">Type material</emphasis>
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<materialsCitation id="22F63CBD9D369271FF79FD77CCE52B49" ID-GBIF-Occurrence="2898878317" collectingDate="2000-12-16" collectionCode="NHMB" collectorName="L. Cizek" country="Papua New Guinea" elevation="0" latitude="-4.7" location="Salemben village" longLatPrecision="1306" longitude="145.4" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Madang Prov." typeStatus="holotype">
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<collectingCountry id="EA8976709D369271FF2EFD53C9832B65" box="[189,469,749,775]" name="Papua New Guinea" pageId="42" pageNumber="43">PAPUA NEW GUINEA</collectingCountry>
• 
<specimenCount id="8498FD699D369271FE79FD53CA542B65" box="[490,514,749,775]" pageId="42" pageNumber="43" type="male">♂</specimenCount>
; “Collection Naturhistorisches Museum Basel // Papua New Guinea 
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<collectorName id="3F6B53369D369271FE92FCAFC9302B48" box="[257,358,784,811]" pageId="42" pageNumber="43">L. Cizek</collectorName>
lgt.// 
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<date id="E62010209D369271FCDCFCAECBB72B48" box="[847,993,784,810]" pageId="42" pageNumber="43" value="2000-12-16">
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16.XII. 
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, 
<elevation id="19B3D1D39D369271FC7EFCAECC6D2B48" box="[1005,1083,784,811]" pageId="42" pageNumber="43">750 m</elevation>
”; 
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.
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<materialsCitation id="22F63CBD9D369271FF2EFCC8CB8E2BF3" ID-GBIF-Occurrence="2898878518" box="[189,984,886,913]" collectingDate="2000-12-16" collectionCode="NHMB" collectorName="L. Cizek" country="Papua New Guinea" elevation="0" latitude="-4.7" location="PAPUA NEW GUINEA" longLatPrecision="1306" longitude="145.4" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Madang Prov." typeStatus="paratype">
<collectingCountry id="EA8976709D369271FF2EFCC8C98F2BF2" box="[189,473,886,913]" name="Papua New Guinea" pageId="42" pageNumber="43">PAPUA NEW GUINEA</collectingCountry>
• 
<specimenCount id="8498FD699D369271FE61FCC8CA762BF3" box="[498,544,886,913]" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
; same labels as for holotype; 
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.
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<emphasis id="A0EAEAF29D369271FF2EFC06C9182BB0" bold="true" box="[189,334,952,978]" pageId="42" pageNumber="43">Description</emphasis>
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MEASUREMENTS. Male 
<typeStatus id="4D2588429D369271FE52FC59CA7D2C63" box="[449,555,999,1025]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
, total body length 
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. Head 
<quantity id="55669B059D369271FC5AFC56CC652C60" box="[969,1075,1000,1026]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="42" pageNumber="43" unit="mm" value="1.7">1.70 mm</quantity>
long, across eyes 
<quantity id="55669B059D369271FA9EFC59CD212C60" box="[1293,1399,999,1026]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="42" pageNumber="43" unit="mm" value="2.05">2.05 mm</quantity>
wide. Pronotum 
<quantity id="55669B059D369271FEEDFBB5C9B02C44" box="[382,486,1035,1062]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="42" pageNumber="43" unit="mm" value="1.75">1.75 mm</quantity>
long, maximum width 
<quantity id="55669B059D369271FD66FBB2CB092C44" box="[757,863,1036,1062]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="42" pageNumber="43" unit="mm" value="2.8">2.80 mm</quantity>
. Elytral length 5.00 mm, maximum combined width 
<quantity id="55669B059D369271FE94FB91C9252C28" box="[263,371,1071,1098]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="42" pageNumber="43" unit="mm" value="3.8">3.80 mm</quantity>
. 
<typeStatus id="4D2588429D369271FEEDFB8EC9B32C28" box="[382,485,1072,1098]" pageId="42" pageNumber="43" type="paratype">Paratype</typeStatus>
is 8.00 mm long.
</paragraph>
<paragraph id="922136E09D369277FF2EFBCFC8A729EF" blockId="42.[189,1399,1137,2025]" lastBlockId="44.[189,1398,265,397]" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
With general features of the 
<taxonomicName id="559E4D639D369271FE6CFBCFCA3D2CE9" authorityName="Heller" authorityYear="1910" box="[511,619,1137,1163]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D369271FE6CFBCFCA3D2CE9" box="[511,619,1137,1163]" italics="true" pageId="42" pageNumber="43">pauxillus</emphasis>
</taxonomicName>
group. Dorsal surface opaque except on anterior and lateral margins of labroclypeus. Frons with paired hump. Male labroclypeus (
<figureCitation id="0AA52A659D369271FC02FB2BCBA22CD2" box="[913,1012,1173,1200]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="42" pageNumber="43">Fig. 116</figureCitation>
) deeply U-shaped emarginate on anterior margin. Anterolateral angles of labroclypeus acute, strongly protruding, raised up at angle of nearly 90° to frons in lateral view. Lateral margins of labroclypeus sinuous in both dorsal and lateral views. Head dorsum except on margins of labroclypeus covered with microscopic velvety pubescence. Head punctures irregularly shaped, very deep, moderately dense. Head with moderately long, appressed elongate scale-like setae. Setae becoming distinctly longer, suberect to erect and stronger clavate on humps of frons (
<figureCitation id="0AA52A659D369271FEEAFAD2C98B2DE5" box="[377,477,1388,1415]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="42" pageNumber="43">Fig. 116</figureCitation>
). Antenna 9-segmented, club 3-lamellate. Pronotum transverse, slightly sinuous on anterior margin with protruding anterolateral angles. Basal margin of pronotum broadly rounded to slightly sinuous, posterolateral angles almost right-angled. Lateral margin of pronotum in dorsal view gradually widened towards middle, deeply emarginate postmedially, crenulate all along (
<figureCitation id="0AA52A659D369271FA90FA66CD3C2D90" box="[1283,1386,1496,1522]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="42" pageNumber="43">Fig. 212</figureCitation>
). Lateral margin of pronotum sinuous in lateral view. Two flat humps on anterior margin opposite to those on frons (
<figureCitation id="0AA52A659D369271FEA0F99EC9CD2E59" box="[307,411,1568,1595]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="42" pageNumber="43">Fig. 116</figureCitation>
). Pronotal punctures ovoid, variably large, very deep, dense. Pronotal dorsum and hypomeron covered with microscopic velvety pubescence. Pronotal setae minute, rising from anterior margin of each puncture, generally not surpassing midlength of corresponding puncture. Few slightly longer scale-like setae scattered over pronotal disc, not surpassing length of corresponding punctures. Posterolateral angles of pronotum with long scale-like clavate setae. Setae on pronotal humps also long, erect, scale-like, clavate. Hypomeron flange-like produced, slightly emarginate and very long setose opposite to compound eye. Antennal pocket deep. Scutellar shield elongate, narrowly rounded apically. Elytron with glabrous elevated broadly interrupted tracks of two (in 
<typeStatus id="4D2588429D369271FC68F8A5CC322F54" box="[1019,1124,1819,1846]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
) or three (in 
<typeStatus id="4D2588429D369271FA9BF8A5CD392F57" box="[1288,1391,1819,1845]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
) longitudinal carinae, including sutural one. Elytral punctures irregularly shaped, deep and dense (
<figureCitation id="0AA52A659D369271FF55F8DDC97D2F1F" box="[198,299,1891,1917]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="42" pageNumber="43">Fig. 302</figureCitation>
) and generally smaller than those on pronotum. Setae minute, arising from anterior margin of each puncture, not surpassing midlength of corresponding punctures. Much longer suberect scale-like clavate setae arranged in irregular longitudinal rows across elytra. Male pygidium flattened dorsally, dense and deep punctate (
<figureCitation id="0AA52A659D369271FE7BF870CA192F8B" box="[488,591,1998,2025]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="42" pageNumber="43">Fig. 469</figureCitation>
). Intervening spaces covered with microscopical velvety pubescence. Setae of pygidium minute on anterior half, becoming longer, suberect and scale-like in apical half. Male protibia with inconspicuous longitudinal dorsal carina, with two acute distal teeth (Fig. 375). Male protibial terminal spur strongly curved. Spiculum gastrale as in Fig. 543. Male aedeagus as in 
<figureCitation id="0AA52A659D309277FA97FEEEC8BB29EF" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="44" pageNumber="45">Figs 611– 613</figureCitation>
.
</paragraph>
<caption id="C6E166689D379270FF2EF836CABB2F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122140" ID-Zenodo-Dep="4122140" httpUri="https://zenodo.org/record/4122140/files/figure.png" pageId="43" pageNumber="44" startId="43.[189,241,1928,1954]" targetBox="[357,1227,273,1901]" targetPageId="43">
<paragraph id="922136E09D379270FF2EF836CABB2F8A" blockId="43.[189,1399,1928,2025]" pageId="43" pageNumber="44">
<emphasis id="A0EAEAF29D379270FF2EF836C91B2FC0" bold="true" box="[189,333,1928,1954]" pageId="43" pageNumber="44">Figs 35–37.</emphasis>
Papuan 
<taxonomicName id="559E4D639D379270FE28F836CB542FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[443,770,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D379270FE28F836CA122FC0" box="[443,580,1928,1954]" italics="true" pageId="43" pageNumber="44">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D379270FDDCF836CB542FC0" author="Macleay W. S." box="[591,770,1928,1954]" pageId="43" pageNumber="44" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view. 
<emphasis id="A0EAEAF29D379270FB8BF836CC632FC0" bold="true" box="[1048,1077,1928,1954]" pageId="43" pageNumber="44">35</emphasis>
. 
<taxonomicName id="559E4D639D379270FBDBF837CD532FC0" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1096,1285,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="dendrolagus" status="sp. nov.">
<emphasis id="A0EAEAF29D379270FBDBF837CD532FC0" box="[1096,1285,1928,1954]" italics="true" pageId="43" pageNumber="44">M. dendrolagus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D379270FA82F836CD242FC0" box="[1297,1394,1928,1954]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMB). 
<emphasis id="A0EAEAF29D379270FE45F815C9A52FA7" bold="true" box="[470,499,1963,1989]" pageId="43" pageNumber="44">36</emphasis>
. 
<taxonomicName id="559E4D639D379270FD95F812CB5F2FA7" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[518,777,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="esau">
<emphasis id="A0EAEAF29D379270FD95F812CA3E2FA7" box="[518,616,1964,1989]" italics="true" pageId="43" pageNumber="44">M. esau</emphasis>
<bibRefCitation id="F60F4B119D379270FDE0F815CB5F2FA7" author="Heller K. M." box="[627,777,1963,1989]" pageId="43" pageNumber="44" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
(lectotype ♂ of 
<taxonomicName id="559E4D639D379270FC49F812CD562FA7" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[986,1280,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="A0EAEAF29D379270FC49F812CC0C2FA7" box="[986,1114,1964,1989]" italics="true" pageId="43" pageNumber="44">M. setosus</emphasis>
<bibRefCitation id="F60F4B119D379270FBF5F812CD562FA7" author="Moser J." box="[1126,1280,1963,1990]" pageId="43" pageNumber="44" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
; ZMHB). 
<emphasis id="A0EAEAF29D379270FF2EF870C88C2F8A" bold="true" box="[189,218,1998,2024]" pageId="43" pageNumber="44">37</emphasis>
. 
<taxonomicName id="559E4D639D379270FF7AF871C9102F8A" authorityName="Heller" authorityYear="1914" box="[233,326,1999,2024]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="esau">
<emphasis id="A0EAEAF29D379270FF7AF871C9102F8A" box="[233,326,1999,2024]" italics="true" pageId="43" pageNumber="44">M. esau</emphasis>
</taxonomicName>
, lectotype, ♂ (SNSD). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D309277FF2EFE0AC9E629AC" blockId="44.[189,432,436,462]" box="[189,432,436,462]" pageId="44" pageNumber="45">
<heading id="C969818C9D309277FF2EFE0AC9E629AC" bold="true" box="[189,432,436,462]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A0EAEAF29D309277FF2EFE0AC9E629AC" bold="true" box="[189,432,436,462]" pageId="44" pageNumber="45">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D309277FF2EFE5DC9FD299F" blockId="44.[189,427,483,509]" box="[189,427,483,509]" pageId="44" pageNumber="45">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D309277FF2EFD9BCB632A0C" box="[189,821,549,623]" pageId="44" pageNumber="45" type="biology_ecology">
<paragraph id="922136E09D309277FF2EFD9BC9762A5D" blockId="44.[189,288,549,575]" box="[189,288,549,575]" pageId="44" pageNumber="45">
<heading id="C969818C9D309277FF2EFD9BC9762A5D" bold="true" box="[189,288,549,575]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A0EAEAF29D309277FF2EFD9BC9762A5D" bold="true" box="[189,288,549,575]" pageId="44" pageNumber="45">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D309277FF2EFDEACB632A0C" blockId="44.[189,821,596,623]" box="[189,821,596,623]" pageId="44" pageNumber="45">
Occurs in lowland rainforests at about 
<quantity id="55669B059D309277FD17FDEACA982A0C" box="[644,718,596,623]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="44" pageNumber="45" unit="m" value="750.0">750 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D309277FF2EFD2BCB932ABC" box="[189,965,661,735]" pageId="44" pageNumber="45" type="distribution">
<paragraph id="922136E09D309277FF2EFD2BC9002ACD" blockId="44.[189,342,661,687]" box="[189,342,661,687]" pageId="44" pageNumber="45">
<heading id="C969818C9D309277FF2EFD2BC9002ACD" bold="true" box="[189,342,661,687]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A0EAEAF29D309277FF2EFD2BC9002ACD" bold="true" box="[189,342,661,687]" pageId="44" pageNumber="45">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D309277FF2EFD7ACB932ABC" blockId="44.[189,965,708,735]" box="[189,965,708,735]" pageId="44" pageNumber="45">
Hitherto only known from Madang Province of East New 
<collectingCountry id="EA8976709D309277FCFAFD7ACB942ABC" box="[873,962,708,734]" name="Guinea-Bissau" pageId="44" pageNumber="45">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>