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<document id="13E9494646C2004622479486C7F97626" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A6080615FDB9FE74FD2DE68B" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba waterstradti" docType="treatment" docVersion="11" lastPageNumber="64" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="62" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A6FCE45F508A454C44F19247E43412D7">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="0D516CB55C464630BEF36559EF2FE90D">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="6249D1B86BB3371F07CFF8F9AC76C5A0">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="CC4747E5065400F0C4521829184230F0">Coca-Abia, María Milagro</mods:namePart>
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<treatment id="03B56743A6080615FDB9FE74FD2DE68B" ID-DOI="http://doi.org/10.5281/zenodo.4773300" ID-GBIF-Taxon="182237254" ID-Zenodo-Dep="4773300" LSID="urn:lsid:plazi:treatment:03B56743A6080615FDB9FE74FD2DE68B" httpUri="http://treatment.plazi.org/id/03B56743A6080615FDB9FE74FD2DE68B" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62">
<subSubSection id="C30685DEA6080617FDB9FE74FC13E5C9" box="[578,1009,477,504]" pageId="61" pageNumber="62" type="nomenclature">
<paragraph id="8BA3D655A6080617FDB9FE74FC13E5C9" blockId="61.[578,1009,477,539]" box="[578,1009,477,504]" pageId="61" pageNumber="62">
<heading id="D0EB6139A6080617FDB9FE74FC13E5C9" box="[578,1009,477,504]" centered="true" fontSize="11" level="2" pageId="61" pageNumber="62" reason="2">
<taxonomicName id="4C1CADD6A6080617FDB9FE74FC13E5C9" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[578,1009,477,504]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FDB9FE74FCA1E5C9" bold="true" box="[578,835,477,504]" italics="true" pageId="61" pageNumber="62">Miridiba waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A6080617FCA9FE77FC0BE5C9" author="Moser J." box="[850,1001,478,504]" pageId="61" pageNumber="62" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30685DEA6080617FD23FDA8FCBEE62A" box="[728,860,513,539]" pageId="61" pageNumber="62" type="description">
<paragraph id="8BA3D655A6080617FD23FDA8FCBEE62A" blockId="61.[578,1009,477,539]" box="[728,860,513,539]" pageId="61" pageNumber="62">
<figureCitation id="1327CAD0A6080617FD23FDA8FCBEE62A" box="[728,860,513,539]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Figs 79–82</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6080617FF46FDEBFC0DE6AE" pageId="61" pageNumber="62" type="reference_group">
<paragraph id="8BA3D655A6080617FF46FDEBFC78E66C" blockId="61.[189,922,578,605]" box="[189,922,578,605]" pageId="61" pageNumber="62">
<treatmentCitationGroup id="AB0CF17BA6080617FF46FDEBFC78E66C" box="[189,922,578,605]" pageId="61" pageNumber="62">
<treatmentCitation id="0ABDF044A6080617FF46FDEBFD51E66D" author="Moser J." box="[189,691,578,605]" page="441" pageId="61" pageNumber="62" year="1912">
<taxonomicName id="4C1CADD6A6080617FF46FDEBFD51E66D" authority="Moser, 1912: 441" authorityName="Moser" authorityPageNumber="441" authorityYear="1912" box="[189,691,578,605]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FF46FDEBFE3EE66D" box="[189,476,578,604]" italics="true" pageId="61" pageNumber="62">Holotrichia waterstradti</emphasis>
<bibRefCitation id="EF8DABA4A6080617FE1FFDEAFD51E66D" author="Moser J." box="[484,691,578,605]" pageId="61" pageNumber="62" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912: 441</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A6080617FD39FDEBFD14E66D" box="[706,758,578,604]" pageId="61" pageNumber="62">type</typeStatus>
loc.:
<collectingCountry id="F30B96C5A6080617FCC3FDEBFC6FE66C" box="[824,909,578,605]" name="Brunei" pageId="61" pageNumber="62">Brunei</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A6080617FF46FD2DFC0DE6AE" blockId="61.[189,1007,644,671]" box="[189,1007,644,671]" pageId="61" pageNumber="62">
<treatmentCitationGroup id="AB0CF17BA6080617FF46FD2DFC0DE6AE" box="[189,1007,644,671]" pageId="61" pageNumber="62">
<taxonomicName id="4C1CADD6A6080617FF46FD2DFE5EE6AF" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[189,444,644,670]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FF46FD2DFE5EE6AF" box="[189,444,644,670]" italics="true" pageId="61" pageNumber="62">Miridiba waterstradti</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6080617FE22FD2DFD5DE6AF" author="Matsumoto T." box="[473,703,644,670]" page="3" pageId="61" pageNumber="62" year="2011">
<bibRefCitation id="EF8DABA4A6080617FE22FD2DFD5DE6AF" author="Matsumoto T." box="[473,703,644,670]" pageId="61" pageNumber="62" pagination="1 - 5" refId="ref45117" refString="Matsumoto T. 2011. Taxonomic notes on the genus Miridiba Reitter (Scarabaeidae, Melolonthinae, Melolonthini) from the Sunda Archipelago. Kogane, Tokyo 12: 1 - 5." type="journal article" year="2011">Matsumoto 2011: 3</bibRefCitation>
</treatmentCitation>
, figs 3, 14 (combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6080617FF46FD6CFE06E24F" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="8BA3D655A6080617FF46FD6CFED4E6EE" blockId="61.[189,310,709,735]" box="[189,310,709,735]" pageId="61" pageNumber="62">
<heading id="D0EB6139A6080617FF46FD6CFED4E6EE" bold="true" box="[189,310,709,735]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="B9680A47A6080617FF46FD6CFED4E6EE" bold="true" box="[189,310,709,735]" pageId="61" pageNumber="62">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6080617FF46FD5DFE06E24F" blockId="61.[188,1401,756,1662]" pageId="61" pageNumber="62">
External morphology of adult (
<figureCitation id="1327CAD0A6080617FDCBFD5DFD6BE73F" box="[560,649,756,782]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Fig. 79</figureCitation>
). Body size
<quantity id="4CE47BB0A6080617FCE4FD5DFC40E73F" box="[799,930,756,782]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="61" pageNumber="62" unit="mm" value="25.0" valueMax="26.0" valueMin="24.0">24–26 mm</quantity>
. Dorsal surface glabrous. Clypeus with conspicuous punctures, concave, shorter than frons, oblique laterally, anterior edge strongly emarginate at middle. Frons with punctures similar those of clypeus; frontal carina sharp, slightly bent backward and depressed at middle. Anterior pronotal margin glabrous, strongly flanged, narrowed at middle and lateral ends; posterior margin glabrous, finely flanged; lateral margins glabrous, smooth, moderately elevated at anterior half; anterior angles obtuse and rounded, not projected forward; posterior angles obtuse and marked. Prosternal process tongue-shaped. Scutellum without punctures at middle line. Elytral surface with punctures regularly distributed, each puncture with a tiny bulge; apical callus well definite; epipleuron glabrous. Foretibia with a moderately developed dorsal carina. Meso- and metatibia with a moderately developed transverse carina interrupted and complete, respectively; inner margin of dorsal surface with few small spines. Prepygidium with punctures irregularly distributed, more scattered at posterior margin; with pubescence short and decumbent, inconspicuous at posterior part. Pygidium irregularly punctate; surface without conspicuous pubescence, at most with tiny setae in each puncture hardly visible, apex with long pubescence. Ventrite 1 with posterior margin visible, enduring strong and short pubescence. Ventrite 2 with conspicuous short pubescence, decumbent and regularly distributed. Ventrites 3 and 4 without conspicuous pubescence. Ventrite 5 moderately depressed at posterior half; punctures and pubescence irregularly distributed; with a row of long pubescence at middle. Ventrite 6 moderately bulging anteriorly; conspicuous punctures and pubescence long and short irregularly distributed. Male genitalia: dorsal branch of parameres symmetrical, with sutural scar completely fused, cleft and a fan-shaped extension at distal end (
<figureCitation id="1327CAD0A6080617FD19FA39FC91E19A" box="[738,883,1424,1451]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Fig. 80A, C</figureCitation>
). Ventral branches close to each other, with elevations at outer margins little developed, apices upward, with a small membranous area close to phallobase (
<figureCitation id="1327CAD0A6080617FEB6FA7EFE5EE1C0" box="[333,444,1495,1521]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Fig. 80B</figureCitation>
). Phallobase shorter than paramere. Endophallus (
<figureCitation id="1327CAD0A6080617FBE2FA7EFB93E1C0" box="[1049,1137,1495,1521]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Fig. 81</figureCitation>
) strongly armed with small spines at middle of sac; distal end with two strong spines at lateral sides of two placoid raspulae; proximal end with numerous acicular soft setae. Apophysis of temones short with ends extend laterally surrounding endophallus partially (T-shaped) (
<figureCitation id="1327CAD0A6080617FD1EF9E9FCDCE26A" box="[741,830,1600,1627]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Fig. 81</figureCitation>
). Female genitalia as described for morphotype IV “
<emphasis id="B9680A47A6080617FF0BF9CDFE80E24F" box="[240,354,1636,1662]" italics="true" pageId="61" pageNumber="62">Bidentata</emphasis>
” (
<figureCitation id="1327CAD0A6080617FE84F9CDFE35E24F" box="[383,471,1636,1662]" captionStart="Figs 79–82" captionStartId="62.[189,241,1883,1909]" captionTargetBox="[278,1310,748,1831]" captionTargetId="figure-132@62.[264,1323,726,1844]" captionTargetPageId="62" captionText="Figs 79–82. Miridiba waterstradti (Moser, 1912). 79–81. Lectotype, ♂ (MFNB). 79. Habitus in dorsal view. 80. Parameres in dorsal (A), ventral (B) and lateral (C) views. 81. Endophallus and temones in dorsal view. 82. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="61" pageNumber="62">Fig. 82</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6080614FF46F90CFD62E5E5" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="materials_examined">
<paragraph id="8BA3D655A6080617FF46F90CFE48E28E" blockId="61.[189,426,1701,1727]" box="[189,426,1701,1727]" pageId="61" pageNumber="62">
<heading id="D0EB6139A6080617FF46F90CFE48E28E" bold="true" box="[189,426,1701,1727]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="B9680A47A6080617FF46F90CFE48E28E" bold="true" box="[189,426,1701,1727]" pageId="61" pageNumber="62">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6080617FF11F97DFC45E369" blockId="61.[189,1398,1748,1880]" pageId="61" pageNumber="62">
<materialsCitation id="3B74DC08A6080617FF11F97DFC45E369" ID-GBIF-Occurrence="3114121358" collectionCode="MFNB" collectorName="N. Borneo Brunei Waterstradt S." country="Brunei" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="BRUNEI" pageId="61" pageNumber="62" specimenCode="MFNB 2017" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B9680A47A6080617FF11F97DFE84E2DF" bold="true" box="[234,358,1748,1774]" pageId="61" pageNumber="62">
<typeStatus id="54A768F7A6080617FF11F97DFE84E2DF" box="[234,358,1748,1774]" pageId="61" pageNumber="62" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
<collectingCountry id="F30B96C5A6080617FF46F95EFEC8E320" box="[189,298,1783,1809]" name="Brunei" pageId="61" pageNumber="62">BRUNEI</collectingCountry>
•
<specimenCount id="9D1A1DDCA6080617FEBAF95EFEB8E320" box="[321,346,1783,1809]" pageId="61" pageNumber="62" type="male">♂</specimenCount>
; “
<collectorName id="26E9B383A6080617FE8AF95EFD0FE320" box="[369,749,1783,1809]" pageId="61" pageNumber="62">N. Borneo Brunei Waterstradt S.</collectorName>
”; “
<taxonomicName id="4C1CADD6A6080617FCEEF95EFBD1E320" authorityName="Moser" authorityYear="1912" box="[789,1075,1783,1809]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FCEEF95EFBD1E320" box="[789,1075,1783,1809]" italics="true" pageId="61" pageNumber="62">Holotrichia waterstradti</emphasis>
</taxonomicName>
Type ♂ Mos”; “SYNTYPE
<taxonomicName id="4C1CADD6A6080617FF46F8B3FD61E305" authority="Moser 1912" authorityName="Moser" authorityYear="1912" box="[189,643,1818,1845]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FF46F8B3FE00E305" box="[189,482,1818,1844]" italics="true" pageId="61" pageNumber="62">Holotrichia waterstradti</emphasis>
<bibRefCitation id="EF8DABA4A6080617FE0BF8B2FD61E305" author="Moser J." box="[496,643,1818,1845]" pageId="61" pageNumber="62" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser 1912</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA6080617FCDFF8B2FC27E305" box="[804,965,1818,1845]" pageId="61" pageNumber="62">MFNB 2017</specimenCode>
”; “Lectotype
<taxonomicName id="4C1CADD6A6080617FB8AF8B3FE81E366" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FB8AF8B3FA94E305" box="[1137,1398,1818,1844]" italics="true" pageId="61" pageNumber="62">Miridiba waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A6080617FF3EF894FEB9E366" author="Moser J." box="[197,347,1853,1879]" pageId="61" pageNumber="62" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A6080617FE90F894FDF5E366" box="[363,535,1853,1879]" pageId="61" pageNumber="62">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A6080617FDC7F894FD59E366" box="[572,699,1853,1879]" pageId="61" pageNumber="62">Coca-Abia</determinerName>
det. 2017”;
<collectionCode id="ED0D4E90A6080617FCB7F897FC41E369" box="[844,931,1854,1880]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="61" pageNumber="62">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A6080614FF11F8D6FD62E5E5" blockId="61.[189,1399,1919,2016]" lastBlockId="62.[189,1401,265,468]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<materialsCitation id="3B74DC08A6080614FF11F8D6FB7CE515" ID-GBIF-Occurrence="3114121388" collectionCode="MFNB" country="Brunei" determinerName="Chuan-bu Gao &amp; Coca-Abia" lastPageId="62" lastPageNumber="63" location="BRUNEI" pageId="61" pageNumber="62" specimenCode="MFNB 2017" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<emphasis id="B9680A47A6080617FF11F8D6FE40E3A8" bold="true" box="[234,418,1919,1945]" pageId="61" pageNumber="62">
<typeStatus id="54A768F7A6080617FF11F8D6FE40E3A8" box="[234,418,1919,1945]" pageId="61" pageNumber="62" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
(here indicated)
<collectingCountry id="F30B96C5A6080617FF46F80BFEC8E38D" box="[189,298,1954,1980]" name="Brunei" pageId="61" pageNumber="62">BRUNEI</collectingCountry>
•
<specimenCount id="9D1A1DDCA6080617FEAEF80BFE69E38D" box="[341,395,1954,1980]" pageId="61" pageNumber="62" type="female">1 ♀</specimenCount>
; “N. Borneo Brunei Waterstradt S.”; “
<taxonomicName id="4C1CADD6A6080617FC7FF80BFB4EE38D" authorityName="Moser" authorityYear="1912" box="[900,1196,1954,1980]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FC7FF80BFB4EE38D" box="[900,1196,1954,1980]" italics="true" pageId="61" pageNumber="62">Holotrichia waterstradti</emphasis>
</taxonomicName>
Type ♀ Mos”; “SYNTYPE
<taxonomicName id="4C1CADD6A6080617FEA7F86CFCC1E3EE" authority="Moser 1912" authorityName="Moser" authorityYear="1912" box="[348,803,1989,2016]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6080617FEA7F86CFD63E3EE" box="[348,641,1989,2015]" italics="true" pageId="61" pageNumber="62">Holotrichia waterstradti</emphasis>
<bibRefCitation id="EF8DABA4A6080617FD74F86FFCC1E3EE" author="Moser J." box="[655,803,1989,2016]" pageId="61" pageNumber="62" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser 1912</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA6080617FC3DF86FFB84E3EE" box="[966,1126,1989,2016]" pageId="61" pageNumber="62">MFNB 2017</specimenCode>
”; “♀”; “Paralectotype
<taxonomicName id="4C1CADD6A60B0614FF46FEA0FD85E512" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[189,615,265,291]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FF46FEA0FE59E512" box="[189,443,265,291]" italics="true" pageId="62" pageNumber="63">Miridiba waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A60B0614FE32FEA0FDBDE512" author="Moser J." box="[457,607,265,291]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A60B0614FD95FEA0FCFBE512" box="[622,793,265,291]" pageId="62" pageNumber="63">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A60B0614FCC6FEA0FC5EE512" box="[829,956,265,291]" pageId="62" pageNumber="63">Coca-Abia</determinerName>
det. 2017”;
<collectionCode id="ED0D4E90A60B0614FBB0FEA3FB7CE515" box="[1099,1182,266,292]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="62" pageNumber="63">MFNB</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B74DC08A60B0614FB4EFEA0FA94E55B" ID-GBIF-Occurrence="3114121348" collectionCode="MFNB" collectorName="N. Borneo Brunei Waterstradt S." determinerName="Chuan-bu Gao &amp; Coca-Abia" pageId="62" pageNumber="63" specimenCode="MFNB 2017" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1A1DDCA60B0614FB4EFEA0FB03E515" box="[1205,1249,265,292]" pageId="62" pageNumber="63" type="female">1 ♀</specimenCount>
; “N. Borneo Brunei Waterstradt S.”; “SYNTYPE
<taxonomicName id="4C1CADD6A60B0614FD81FE85FBD7E577" authority="Moser 1912" authorityName="Moser" authorityYear="1912" box="[634,1077,300,327]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FD81FE85FC79E577" box="[634,923,300,326]" italics="true" pageId="62" pageNumber="63">Holotrichia waterstradti</emphasis>
<bibRefCitation id="EF8DABA4A60B0614FC5DFE84FBD7E577" author="Moser J." box="[934,1077,300,327]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser 1912</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA60B0614FB37FE84FA8BE577" box="[1228,1385,300,327]" pageId="62" pageNumber="63">MFNB 2017</specimenCode>
”; “Paralectotype
<taxonomicName id="4C1CADD6A60B0614FE8FFEE6FCCAE55B" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[372,808,335,362]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FE8FFEE6FD97E558" box="[372,629,335,361]" italics="true" pageId="62" pageNumber="63">Miridiba waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A60B0614FD7CFEF9FCC2E55B" author="Moser J." box="[647,800,336,362]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A60B0614FCC9FEE6FC03E55B" box="[818,993,335,362]" pageId="62" pageNumber="63">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A60B0614FBF0FEE6FB68E558" box="[1035,1162,335,361]" pageId="62" pageNumber="63">Coca-Abia</determinerName>
det. 2017”;
<collectionCode id="ED0D4E90A60B0614FAD8FEF9FA94E55B" box="[1315,1398,336,362]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="62" pageNumber="63">MFNB</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B74DC08A60B0614FF2BFEDAFD62E5E5" ID-GBIF-Occurrence="3114121394" collectionCode="MFNB" determinerName="Chuan-bu Gao &amp; Coca-Abia" pageId="62" pageNumber="63" specimenCode="MFNB 2017" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1A1DDCA60B0614FF2BFEDAFF1CE5BC" box="[208,254,371,397]" pageId="62" pageNumber="63" type="female">1 ♀</specimenCount>
; “
<emphasis id="B9680A47A60B0614FEE0FEDBFE4FE5BD" box="[283,429,370,396]" italics="true" pageId="62" pageNumber="63">Waterstradti</emphasis>
Mos.”; “N. Borneo Brunei Waterstradt S.”; “SYNTYPE
<taxonomicName id="4C1CADD6A60B0614FBACFEDBFEAEE581" authority="Moser 1912" authorityName="Moser" authorityYear="1912" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FBACFEDBFA95E5BD" box="[1111,1399,370,396]" italics="true" pageId="62" pageNumber="63">Holotrichia waterstradti</emphasis>
<bibRefCitation id="EF8DABA4A60B0614FF46FE3FFEAEE581" author="Moser J." box="[189,332,406,432]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser 1912</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA60B0614FE24FE3FFD99E581" box="[479,635,406,432]" pageId="62" pageNumber="63">MFNB 2017</specimenCode>
”; “Paralectotype
<taxonomicName id="4C1CADD6A60B0614FCB1FE3FFB19E581" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[842,1275,406,432]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FCB1FE3FFBA8E581" box="[842,1098,406,432]" italics="true" pageId="62" pageNumber="63">Miridiba waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A60B0614FBA0FE3FFB11E581" author="Moser J." box="[1115,1267,406,432]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A60B0614FAFFFE3FFF0DE5E2" pageId="62" pageNumber="63">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A60B0614FEEEFE10FE76E5E2" box="[277,404,441,467]" pageId="62" pageNumber="63">Coca-Abia</determinerName>
det. 2017”;
<collectionCode id="ED0D4E90A60B0614FDDEFE13FD9EE5E5" box="[549,636,442,468]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="62" pageNumber="63">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA60B0615FF46FE52FCA3E558" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="discussion">
<paragraph id="8BA3D655A60B0614FF46FE52FECDE624" blockId="62.[189,303,507,533]" box="[189,303,507,533]" pageId="62" pageNumber="63">
<heading id="D0EB6139A60B0614FF46FE52FECDE624" bold="true" box="[189,303,507,533]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="B9680A47A60B0614FF46FE52FECDE624" bold="true" box="[189,303,507,533]" pageId="62" pageNumber="63">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A60B0615FF46FD80FCA3E558" blockId="62.[188,1399,553,650]" lastBlockId="63.[187,1399,265,361]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<bibRefCitation id="EF8DABA4A60B0614FF46FD80FE83E672" author="Moser J." box="[189,353,553,579]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser (1912)</bibRefCitation>
described
<taxonomicName id="4C1CADD6A60B0614FE12FD80FCEEE672" authorityName="Moser" authorityYear="1912" box="[489,780,553,579]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FE12FD80FCEEE672" box="[489,780,553,579]" italics="true" pageId="62" pageNumber="63">Holotrichia waterstradti</emphasis>
</taxonomicName>
from a series of specimens from Borneo (
<collectingCountry id="F30B96C5A60B0614FAE0FD80FA8CE675" box="[1307,1390,553,580]" name="Brunei" pageId="62" pageNumber="63">Brunei</collectingCountry>
) collected by Waterstradt.
<bibRefCitation id="EF8DABA4A60B0614FE1EFDE4FD5DE657" author="Matsumoto T." box="[485,703,588,615]" pageId="62" pageNumber="63" pagination="1 - 5" refId="ref45117" refString="Matsumoto T. 2011. Taxonomic notes on the genus Miridiba Reitter (Scarabaeidae, Melolonthinae, Melolonthini) from the Sunda Archipelago. Kogane, Tokyo 12: 1 - 5." type="journal article" year="2011">Matsumoto (2011)</bibRefCitation>
transferred this species based on a non-type male specimen from the westernmost part of Borneo Island. After studying
<specimenCount id="9D1A1DDCA60B0614FC68FDC6FBE3E6BB" box="[915,1025,623,650]" pageId="62" pageNumber="63" type="generic">four type</specimenCount>
specimens (MFNB) (
<specimenCount id="9D1A1DDCA60B0614FAFCFDC6FA94E6B8" box="[1287,1398,623,649]" pageId="62" pageNumber="63" type="male">one male</specimenCount>
and
<specimenCount id="9D1A1DDCA60A0615FF08FEA0FE7BE515" box="[243,409,265,292]" pageId="63" pageNumber="64" type="female">three females</specimenCount>
), we agree with the transfer of
<taxonomicName id="4C1CADD6A60A0615FCD9FEA3FC3BE512" authorityName="Moser" authorityYear="1912" box="[802,985,265,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60A0615FCD9FEA3FC3BE512" box="[802,985,265,291]" italics="true" pageId="63" pageNumber="64">H. waterstradti</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A60A0615FBFCFEA0FB8DE512" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[1031,1135,265,291]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A60A0615FBFCFEA0FB8DE512" box="[1031,1135,265,291]" italics="true" pageId="63" pageNumber="64">Miridiba</emphasis>
</taxonomicName>
. Moreover, male and female genitalia allow us to consider this species to belong to morphotype IV “
<emphasis id="B9680A47A60A0615FBB4FE85FB23E577" box="[1103,1217,300,326]" italics="true" pageId="63" pageNumber="64">Bidentata</emphasis>
”. We designate
<typeStatus id="54A768F7A60A0615FF40FEE6FEC5E558" box="[187,295,335,361]" pageId="63" pageNumber="64" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54A768F7A60A0615FE9BFEE6FDEBE558" box="[352,521,335,361]" pageId="63" pageNumber="64" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C1CADD6A60A0615FDC9FEF9FD0AE558" authorityName="Moser" authorityYear="1912" box="[562,744,335,361]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60A0615FDC9FEF9FD0AE558" box="[562,744,335,361]" italics="true" pageId="63" pageNumber="64">M. waterstradti</emphasis>
</taxonomicName>
herein.
</paragraph>
</subSubSection>
<caption id="DF6386DDA60B0614FF46F8F2FEFCE3D1" ID-DOI="http://doi.org/10.5281/zenodo.4770396" ID-Zenodo-Dep="4770396" httpUri="https://zenodo.org/record/4770396/files/figure.png" pageId="62" pageNumber="63" startId="62.[189,241,1883,1909]" targetBox="[278,1310,748,1831]" targetPageId="62">
<paragraph id="8BA3D655A60B0614FF46F8F2FEFCE3D1" blockId="62.[189,1399,1883,2016]" pageId="62" pageNumber="63">
<emphasis id="B9680A47A60B0614FF46F8F2FEABE344" bold="true" box="[189,329,1883,1909]" pageId="62" pageNumber="63">Figs 79–82.</emphasis>
<taxonomicName id="4C1CADD6A60B0614FEA9F8F2FCE6E347" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[338,772,1883,1910]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A60B0614FEA9F8F2FDB3E344" box="[338,593,1883,1909]" italics="true" pageId="62" pageNumber="63">Miridiba waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A60B0614FD9AF8F5FD19E347" author="Moser J." box="[609,763,1884,1910]" pageId="62" pageNumber="63" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B9680A47A60B0614FCEBF8F2FCBBE344" bold="true" box="[784,857,1883,1909]" pageId="62" pageNumber="63">79–81</emphasis>
. Lectotype, ♂ (MFNB).
<emphasis id="B9680A47A60B0614FB7BF8F2FB7FE344" bold="true" box="[1152,1181,1883,1909]" pageId="62" pageNumber="63">79</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A60B0614FEFEF8D6FEC0E3A8" bold="true" box="[261,290,1919,1945]" pageId="62" pageNumber="63">80</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A60B0614FC02F8D6FBF6E3A8" bold="true" box="[1017,1044,1919,1945]" pageId="62" pageNumber="63">81</emphasis>
. Endophallus and temones in dorsal view.
<emphasis id="B9680A47A60B0614FEAFF80BFE93E38D" bold="true" box="[340,369,1954,1980]" pageId="62" pageNumber="63">82</emphasis>
. Paralectotype, ♀ (MFNB), genitalia in dorsal view. Scale bars: 79 = 5.0 mm; 80–82 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA60A0615FF46FE39FDFCE5EB" box="[189,542,400,474]" pageId="63" pageNumber="64" type="distribution">
<paragraph id="8BA3D655A60A0615FF46FE39FEB4E59B" blockId="63.[189,342,400,426]" box="[189,342,400,426]" pageId="63" pageNumber="64">
<heading id="D0EB6139A60A0615FF46FE39FEB4E59B" bold="true" box="[189,342,400,426]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="B9680A47A60A0615FF46FE39FEB4E59B" bold="true" box="[189,342,400,426]" pageId="63" pageNumber="64">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A60A0615FF46FE69FDFCE5EB" blockId="63.[189,542,447,474]" box="[189,542,447,474]" pageId="63" pageNumber="64">
Borneo (
<collectingCountry id="F30B96C5A60A0615FED8FE16FE95E5E8" box="[291,375,447,473]" name="Brunei" pageId="63" pageNumber="64">Brunei</collectingCountry>
, Mt Bawang).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA60A0615FF46FDABFD2DE68B" pageId="63" pageNumber="64" type="description">
<paragraph id="8BA3D655A60A0615FF46FDABFCF4E62F" blockId="63.[189,790,514,543]" box="[189,790,514,543]" pageId="63" pageNumber="64">
<emphasis id="B9680A47A60A0615FF46FDABFCF4E62F" bold="true" box="[189,790,514,543]" italics="true" pageId="63" pageNumber="64">Species included in Morphotype V “Sinensis”</emphasis>
</paragraph>
<paragraph id="8BA3D655A60A0615FF46FD9FFD2DE68B" blockId="63.[187,1399,566,698]" pageId="63" pageNumber="64">The group is composed of four species, which are characterized by antenna 10-segmented, pronotal surface densely punctate (distance between punctures smaller than diameter of puncture), pronotal lateral margins glabrous and smooth except at middle where is moderately serrated with short pubescence, and male and female genitalia as specified above.</paragraph>
</subSubSection>
</treatment>
</document>
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