treatments-xml/data/F6/CA/D1/F6CAD151C84054CF8014305006360C57.xml

<document id="A6830EAF206155EBED701D137A3B3665" ID-DOI="10.3897/phytokeys.251.141836" ID-publisher-id="141836" URI-arpha="F9731662-1F94-53C7-A46D-DB49A469819F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1738164160160" checkinUser="pensoft" docAuthor="Lu, Zhao-Cen, Chang, Shi-Li, Mo, Ming-Lin, Wu, You-Dong &amp; Xu, Wei-Bin" docDate="2025" docId="F6CAD151C84054CF8014305006360C57" docLanguage="en" docName="PhytoKeys 251: 233-240" docOrigin="PhytoKeys 251" docSource="https://phytokeys.pensoft.net/article/141836/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Tilia saxatilis Z. C. Lu &amp; W. B. Xu 2025, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="240" masterDocId="F97316621F9453C7A46DDB49A469819F" masterDocTitle="Tilia saxatilis (Malvaceae), a new species from limestone areas of Guangxi, China" masterLastPageNumber="240" masterPageNumber="233" pageNumber="233" updateTime="1738169050610" updateUser="admin">
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<mods:title id="0357E59C70CC2E62F8416DC536E03FC0">Tilia saxatilis (Malvaceae), a new species from limestone areas of Guangxi, China</mods:title>
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<mods:namePart id="46B6452151E44D7A7E0F87E035E60676">Lu, Zhao-Cen</mods:namePart>
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<mods:affiliation id="27E9EDED720840EC8D6185C6663D35A6">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain / Nonggang Karst Ecosystem Observation and Research Station of Guangxi, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:namePart id="6568A210437E1146BF69E87C6CC0BFF3">Chang, Shi-Li</mods:namePart>
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<mods:affiliation id="482312C10C5A32E070BC259B8713A204">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain / Nonggang Karst Ecosystem Observation and Research Station of Guangxi, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China &amp; College of Life Sciences, Guangxi Normal University, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:namePart id="6DCDE411DCB14ECA22E9AADDFB24F9A6">Mo, Ming-Lin</mods:namePart>
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<mods:affiliation id="CF57A1BDB3AC73B4D9540090ACCD8F91">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain / Nonggang Karst Ecosystem Observation and Research Station of Guangxi, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China &amp; College of Life Sciences, Guangxi Normal University, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:namePart id="64587E61BB22BB7BE6B686D6697390B5">Wu, You-Dong</mods:namePart>
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<mods:affiliation id="7D345C2DBF2E35276093CDE47ECC5027">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain / Nonggang Karst Ecosystem Observation and Research Station of Guangxi, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China &amp; College of Life Sciences, Guangxi Normal University, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:namePart id="51672E99F8B1C3CAF0C185431417539B">Xu, Wei-Bin</mods:namePart>
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<mods:affiliation id="18E4C05FFEFDE54ACC85AAA0561CF8E3">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain / Nonggang Karst Ecosystem Observation and Research Station of Guangxi, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:part id="0063050EB2D7049EDE241421B989E31B">
<mods:date id="D6CFBACAFF0A867D92F40C7B18B12777">2025</mods:date>
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<mods:number id="83BCCBAA8B627FB0D2605065A21535B6">2025-01-29</mods:number>
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<mods:number id="B418BA100A05DDE3D010829ADA216412">251</mods:number>
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<mods:identifier id="0E2614092412DF07AC334C09BB9538F2" type="DOI">10.3897/phytokeys.251.141836</mods:identifier>
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<treatment id="F6CAD151C84054CF8014305006360C57" ID-DOI="http://doi.org/10.5281/zenodo.14765780" ID-Zenodo-Dep="14765780" ID-arpha="F6CAD151-C840-54CF-8014-305006360C57" ID-ipni="urn:lsid:ipni.org:names:77355914-1" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57" LSID-IPNI="urn:lsid:ipni.org:names:77355914-1" httpUri="http://treatment.plazi.org/id/F6CAD151C84054CF8014305006360C57" scope_family="Malvaceae">
<subSubSection id="C8AF746576F64FF5C50988F470199380" type="nomenclature">
<paragraph id="653740A93D9C842E0F4340362888E8C6">
<taxonomicName id="A8925E78656D479822932CDB7283AD98" authority="Z. C. Lu &amp; W. B. Xu" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis" status="sp. nov.">
<emphasis id="52204905D8E9FB5FD494ED06D1BCB1B6" italics="true">Tilia saxatilis</emphasis>
Z. C. Lu &amp; W. B. Xu
</taxonomicName>
<taxonomicNameLabel id="D6067F8E291783BA5765EC4282FD4A4D" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C2DE04DFDF89828154DC32673574184E" type="description">
<paragraph id="E4075D0555D20F6F76CF6AB0CCBA95BC">
<figureCitation id="23604FCA2B4E7EB1283E177811FF6B74" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tilia saxatilis sp. nov. A habit B winter buds C flowering branches D trunk E fruiting branches F habitat (White circle shows where the new species grow)." figureDoi="10.3897/phytokeys.251.141836.figure1" httpUri="https://binary.pensoft.net/fig/1238086">Figs 1</figureCitation>
, 
<figureCitation id="E354FF6688C0A6298E9E6014821DD968" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tilia saxatilis sp. nov. A leaf in abaxial view (Red circle shows tuft domatia in vein axils) B bract in abaxial view C bract and cyme D bract and infructescence E flower in frontal view F flower in lateral view G flower in dorsal view H dissection of flower I fruits J seeds." figureDoi="10.3897/phytokeys.251.141836.figure2" httpUri="https://binary.pensoft.net/fig/1238087">2</figureCitation>
, 
<figureCitation id="88D86D07809019CAFC48C860437CBDE9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype sheet of Tilia saxatilis (IBK)." figureDoi="10.3897/phytokeys.251.141836.figure3" httpUri="https://binary.pensoft.net/fig/1238088">3 Chinese name: shí shān duàn (石山椴</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBHAC" type="diagnosis">
<paragraph id="3230622CBFDC52918CDA45BBD7E97B56">
<heading id="23483B05FE2D757F9C850FB86A661D89" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="3386C7A5ED088EE58C22E47F5B8EFFDE">
This new species is similar to 
<taxonomicName id="0B80D4F4300DB937EAF3FE6E412E8409" authority="Szyszyl." authorityName="Szyszyl." class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="tuan">
<emphasis id="1FE0F27ECDC23F80B81E82AF617286BD" italics="true">Tilia tuan</emphasis>
Szyszyl.
</taxonomicName>
, but differs in having leaf blades that are oblong or ovate-oblong (vs. narrowly ovate or ovate-oblong to ovate-orbicular), margins entire (vs. entire or with a few minute teeth near apex or prominently dentate); fruit ellipsoid (vs. globose or obovoid-globose), 5 - angled (vs. not ridged), apex acute (vs. rounded).
</paragraph>
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<paragraph id="C51DE88726B8195392E91415F610C263">
<label id="1A53DA9575BE55DF83D54EFF13C21DCA">Figure 1.</label>
</paragraph>
<paragraph id="896001782286378A98C2FD0D2CDAF63F">
<taxonomicName id="A5412C59E51B1335691A5E38D0BE66D9" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis" status="sp. nov.">
<emphasis id="7D2552FEAF985F22B9AE402E94D8E668" italics="true">Tilia saxatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4921B460153AF4D178B41894473A6F39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F30899F7B7AFF53FB2A8516835439E5B" bold="true">A</emphasis>
habit 
<emphasis id="63D91D2E45E9F463961C4E7896518E70" bold="true">B</emphasis>
winter buds 
<emphasis id="A04CF8C1F237BE176D2DD0ACAFA394A4" bold="true">C</emphasis>
flowering branches 
<emphasis id="8DC64681622B51991381F1CBBFF8997A" bold="true">D</emphasis>
trunk 
<emphasis id="84188F633CDA06D7B634FA872BC0C40F" bold="true">E</emphasis>
fruiting branches 
<emphasis id="B498FE09D5477EAC926A1B3E964C1CDE" bold="true">F</emphasis>
habitat (White circle shows where the new species grow).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EWIAC" type="material">
<paragraph id="A630DAC7734372ED5903874097CA0676">
<heading id="FD43143B124B9B1F000BFCBC98ACB0A7" reason="title">Type.</heading>
</paragraph>
<paragraph id="C1F4D9AAD1658CC34B3830DF8C94F86A">
<materialsCitation id="8C21FF1A537DC471CC0E4ADB41E25D29" collectedFrom="limestone slope" collectingDate="2023-09-28" collectionCode="IBSC, PE" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="769" latitude="24.079994" location="Shangzhen Village" longLatPrecision="1" longitude="107.82876" municipality="Hechi City" specimenCode="IBK 00464802, IBK 00464803, IBK 00464804" specimenCount="2" stateProvince="Guangxi" typeStatus="holotype">
<collectingCountry id="FE8202C3B39346FA83EE21C19D40F950" name="China">China</collectingCountry>
• 
<collectingRegion id="005387688B6AA6ED114069C7B84C3FF3" country="China" name="Guangxi">Guangxi</collectingRegion>
: 
<collectingMunicipality id="A1F87A933C63017C7D577F3416917488">Hechi City</collectingMunicipality>
, 
<collectingCounty id="02FE2C0A3CBCB22C34953426391E24EB">Du’an County</collectingCounty>
, 
<collectingMunicipality id="A17A48FB967D14F95B2CEA4C700BAC6D">Bao’an Town</collectingMunicipality>
, 
<location id="FB51AA27C72FDC9EAD481513C20D0141" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:FB51AA27C72FDC9EAD481513C20D0141" country="China" county="Du'an County" latitude="24.079994" longLatPrecision="1" longitude="107.82876" municipality="Hechi City" name="Shangzhen Village" stateProvince="Guangxi">Shangzhen Village</location>
, 
<location id="BC0AEE29F64B84BAE7863B792287B49E" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:BC0AEE29F64B84BAE7863B792287B49E" country="China" county="Du'an County" latitude="24.079994" longLatPrecision="1" longitude="107.82876" municipality="Hechi City" name="Nongwen" stateProvince="Guangxi">Nongwen</location>
, around the point 
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<geoCoordinate id="6EE55A81D9C3697A79075D3CAFE37267" degrees="24.07999417" direction="north" orientation="latitude" precision="1" value="24.079994">24.07999417 ° N</geoCoordinate>
, 
<geoCoordinate id="365512D987D31E763899D56A04C92A5B" degrees="107.82876" direction="east" orientation="longitude" precision="1" value="107.82876">107.82876 ° E</geoCoordinate>
</named-content>
, 
<collectedFrom id="FE0BAAA7D2E99F7D4D89730E8AF56630">limestone slope</collectedFrom>
, alt. 
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<elevation id="A42F583DBD18970068DD6673FD67FBE7" metricMagnitude="2" metricUnit="m" metricValue="7.69" unit="m" value="769.0">769 m</elevation>
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, 
<date id="DCF11ACDD055EFDA412D374A857FE686" value="2023-09-28">
<collectingDate id="0002D8850F0C78D6469EFF49A7020132" value="2023-09-28">28 September 2023</collectingDate>
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, 
<emphasis id="E84C745732FFB03E24FA784BFF894DEB" italics="true">
<collectorName id="99136F1787B9E155DD72EBCA262F6C47">W. B. Xu</collectorName>
, 
<collectorName id="7EA8D181C61D43AC95D91151F16411B2">Z. C. Lu</collectorName>
, 
<collectorName id="67E6CA84DAAAA2FB939AFDD72DF070B0">M. L Mo</collectorName>
, 
<collectorName id="CACB102798125D836E47233319C76CD5">S. L. Chang</collectorName>
&amp; 
<collectorName id="9287896A4F4514E90DF03A97D763975C">J. Q. Huang</collectorName>
17323
</emphasis>
(
<typeStatus id="B3357C57075A3D48503459AF6DC234B5" type="holotype">holotype</typeStatus>
: 
<specimenCode id="124D1026A8749FDBE697707DE7BDE980" collectionCode="IBK" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK 00464802</specimenCode>
; 
<typeStatus id="4095D131FDDCE078287B125A1AE87CDC" type="isotype">isotypes</typeStatus>
: 
<specimenCode id="FCFAF6734768F1D4236C6ECDE9845321">IBK 00464803</specimenCode>
, 
<specimenCode id="C840732DF67F6E7D91BAA0FA7FC85E2C" collectionCode="IBK" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK 00464804</specimenCode>
, 
<collectionCode id="7B4D30B4BB54215696EAA3A06E89AFD4" country="China" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" type="Herbarium">IBSC</collectionCode>
, 
<collectionCode id="0E6297DDA329140D4556818FB8F74CB3" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" type="Herbarium">PE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGJAC" type="description">
<paragraph id="025643461D96CDC4F1817343EC4C8A8C">
<heading id="E22343A58BD25C7DD215DE04EFB095FA" reason="title">Description.</heading>
</paragraph>
<paragraph id="8548CF199069C74DD42E13EE3DC1F562">
Trees 
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tall, DBH 
<quantity id="3457CFCBF0D31CF66CB9EA458BF62091" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" unit="cm" value="45.0" valueMax="80.0" valueMin="10.0">10–80 cm</quantity>
. Bark dark grey; branchlets glabrous; winter buds ovoid, glabrous or slightly hairy at tip. Petiole 
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long, glabrous; leaf blades oblong or ovate-oblong, (3.5 –) 4.5–10 (– 14.4) cm long, 2–5.5 (– 6.2) cm wide, thickly papery, glabrous on both sides, with brown tuft domatia in vein axils of abaxial surface, lateral veins 7–8 pairs, raised on abaxial surface, reticulate veins distinct abaxially, base oblique, truncate or cordate, margins entire, apex acuminate. Cymes 5–16 - flowered, 
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long, peduncles glabrous. Bracts narrowly oblong, 
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long, 
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wide, adnate to peduncle for 1 / 3–2 / 5 of its length, glabrous or adaxially slightly hairy along mid-vein, apex obtuse, sessile. Pedicels 
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long, glabrous. Sepals 5, ovate, 
<quantity id="30BA42285F583EE0E854FD1AF6AA851F" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
long, abaxially tomentose, adaxially tomentose or with long tomentose at base. Petals oblong, 
<quantity id="3CE1CDAE7A714FE24B214090210AA3CE" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, glabrous. Stamens ca. 
<quantity id="9DD5BE570BD20F7153B6F967F52C6412" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
long, glabrous; staminodes 5, slightly smaller than petals, glabrous. Ovary densely tomentose; style 
<quantity id="77F086D1B04E5E11D3A27117E3ECB74F" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, glabrous. Fruit ellipsoid, 5 - angled, 
<quantity id="DA587538C12CC63ACF0AD02CC70C594F" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
long, 
<quantity id="184173FE31B0DBAC14E415619E59170B" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
wide, densely appressed tomentose, apex acute; exocarp woody, hard, indehiscent. Seed ellipsoid, ca. 
<quantity id="3B08150118A72AE7071009B2C22816B7" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
long.
</paragraph>
<caption id="A9AC429E2FE3D44B342F924E04B4519F" ID-DOI="10.3897/phytokeys.251.141836.figure2" ID-arpha="4A4F79D1-B100-55AF-A3F7-0F4A9AAF6435" httpUri="https://binary.pensoft.net/fig/1238087" startId="F2">
<paragraph id="583CE96E0829A3D47CC83FFD2CA85821">
<label id="3CE65486A01341B7A828EDAE305CB62C">Figure 2.</label>
</paragraph>
<paragraph id="FBC5CC4377EE8483ACCE288CE9294808">
<taxonomicName id="999E9E408A51BF153034CA52BCEE765D" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis" status="sp. nov.">
<emphasis id="A80573A2ED9584D3B48817B725F2D9BF" italics="true">Tilia saxatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F14F581F6EB9D880FF7BF095EE2582C5" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="70168A35DD15DDB9CFA8507F153A800D" bold="true">A</emphasis>
leaf in abaxial view (Red circle shows tuft domatia in vein axils) 
<emphasis id="8E3282F89235F40251320695FDF5FBBA" bold="true">B</emphasis>
bract in abaxial view 
<emphasis id="A9A5E68CE9FB928A08060D4E9F7405DB" bold="true">C</emphasis>
bract and cyme 
<emphasis id="F321320605B60E33E2ECF1BCEDB8D580" bold="true">D</emphasis>
bract and infructescence 
<emphasis id="AAF55DB354C1440A7EFC12F6F838E367" bold="true">E</emphasis>
flower in frontal view 
<emphasis id="98F6BC3C4018285E63CAD15BE7537995" bold="true">F</emphasis>
flower in lateral view 
<emphasis id="98887EBFA5EF7230F65E1E4171028DB1" bold="true">G</emphasis>
flower in dorsal view 
<emphasis id="6EEE86AE357F6EB39A5E871ED9F005CC" bold="true">H</emphasis>
dissection of flower 
<emphasis id="5F7F9255ADD4A8CCFEFD785A5E1F1AE4" bold="true">I</emphasis>
fruits 
<emphasis id="D54B177183B41012CB62C4C458BACE56" bold="true">J</emphasis>
seeds.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXKAC" type="etymology">
<paragraph id="037B6482287E5C9FDBE82719E3385A7D">
<heading id="0467ADD75F6BFF53BDFB6E7B13C7A915" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A5BB8350031C4F2F6695C76B7512DCE1">
The specific epithet ‘ 
<emphasis id="1423AB7E03920DDF705AADC2B372278E" italics="true">saxatilis</emphasis>
’ refers to the limestone habitats of this new species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHLAC" type="phenology">
<paragraph id="3AEC0766131011D36479CC0FE7BD69B9">
<heading id="3FEDF5A712EC2AFB0D63CBB1A899DCC5" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="7245D48EA94651F27EEF515A975843BD">Flowering July and fruiting from September to October.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMLAC" type="distribution">
<paragraph id="31B784C10E67501B734285B6D0996AD3">
<heading id="68C48E5CB3A8ABEAB16720842474888A" reason="title">Distribution and habitat.</heading>
</paragraph>
<paragraph id="1795B9F64FD2DED739E564653FED75F3">
<taxonomicName id="0FD00BFE0F85AD7C5E3D617E8C2B7350" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="BB31DD471B578E3E957D45216DCD0B66" italics="true">Tilia saxatilis</emphasis>
</taxonomicName>
has only been collected from five localities restricted to central 
<collectingRegion id="ABA61B2A48D46EB4060F4E12E75DD05F" country="China" name="Guangxi">Guangxi</collectingRegion>
of 
<collectingCountry id="C044DF4FB017EC6E15E0E1D260B19219" name="China">China</collectingCountry>
. It grows sporadically in forests on limestone slopes, rare on peaks, at an elevation of 
<quantity id="06AF342F3ED940F63AD9FB4C10D47D23" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="9.5" metricValueMin="7.0" unit="m" value="825.0" valueMax="950.0" valueMin="700.0">
700– 
<specimenCount id="D67B44FCD9E8CF137DDFB7EB7C48C483" count="950" type="generic">950 m.</specimenCount>
</quantity>
The type localities are typical limestone karst landform and belong to the southern subtropical monsoon climate areas, the average annual temperature being 18–21 ° C; the annual sunshine duration is 1220–1590 hours and the annual accumulated temperature is about 6300 ° C; the annual average relative humidity is 74–80 %; the average annual rainfall is 
<quantity id="C7C68DEFE15D4A6479B902B35538353C" metricMagnitude="0" metricUnit="m" metricValue="1.465" metricValueMax="1.68" metricValueMin="1.25" unit="mm" value="1465.0" valueMax="1680.0" valueMin="1250.0">1250–1680 mm</quantity>
and the evaporation is 
<quantity id="EACF05D5D2DADA7B7E3CF4830AB2B63D" metricMagnitude="0" metricUnit="m" metricValue="1.43" metricValueMax="1.65" metricValueMin="1.21" unit="mm" value="1430.0" valueMax="1650.0" valueMin="1210.0">1210–1650 mm</quantity>
.
</paragraph>
<caption id="4467A8170B3A3F5034CAFB013D69FEDF" ID-DOI="10.3897/phytokeys.251.141836.figure3" ID-arpha="2C0B3640-A890-5E60-9E7B-7B0CDE6A069D" httpUri="https://binary.pensoft.net/fig/1238088" startId="F3">
<paragraph id="D5F71B3CA0E3A64314CD48367F770D2F">
<label id="D87754F45397DEDD53F844E614749614">Figure 3.</label>
</paragraph>
<paragraph id="E7D57656DDE2CC26A423146376A4DB61">
The holotype sheet of 
<taxonomicName id="F0947481C6E99BCDC888B0648BCEE2DC" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="130A1A74C330F1D274C00154B5049337" italics="true">Tilia saxatilis</emphasis>
</taxonomicName>
(IBK).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EVMAC" type="conservation status">
<paragraph id="2E5E5E6487B250C688C4607A0981FFC1">
<heading id="7F934B4E8C3E1D0D0E2B06E8406CE0CD" reason="title">Conservation status.</heading>
</paragraph>
<paragraph id="D80E2CE37126CA226CD4C98BC1EF25D6">
The new species has been found in two localities in Du’an County and three localities in Dahua County, 
<collectingRegion id="418540F7B7FC021AA50E27C23C104990" country="China" name="Guangxi">Guangxi</collectingRegion>
, 
<collectingCountry id="D59D820C696CEDC976E14CFEDBE4001A" name="China">China</collectingCountry>
. These five localities are in the assessment area of Southwest Karst National Park, which is currently being prepared. The extent of occurrence is about 
<quantity id="60F6DF95E1F359DF53160068FF45AA63" metricMagnitude="5" metricUnit="m" metricValue="9.6" unit="km" value="960.0">960 km</quantity>
<superScript id="7C006FD2600ECD4444AE2C4353FC25A7">2</superScript>
(&lt;
<quantity id="6F4A054597F9C5016F1B24E6DADAA7F9" metricMagnitude="6" metricUnit="m" metricValue="5.0" unit="km" value="5000.0">5000 km</quantity>
<superScript id="0E69E7AC3EBEA5312B2604F44DC6F64A">2</superScript>
) and its occupancy area is predicted to continuously decline in the future due to grazing and firewood collection by local people. Therefore, according to the IUCN Red List Categories and Criteria (
<bibRefCitation id="DA1C94B6700A47C9827AC41787A40F64" refId="B2" refString="IUCN Standards and Petitions Committee (2022) Guidelines for using the IUCN Red List Categories and Criteria. Version 16. Prepared by the Standards and Petitions Committee of the IUCN Species Survival Commission. https://www.iucnredlist.org/resources/redlistguidelines [accessed on 13 November 2024]" url="https://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN Standards and Petitions Committee 2022</bibRefCitation>
), 
<taxonomicName id="537DA8B38C2CAAA81604A243D5CC5CD4" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="311392722F4F2E0A6959578ADAF2E039" italics="true">Tilia saxatilis</emphasis>
</taxonomicName>
should be considered in the Endangered (
<abbrev id="ABBRID0EONAC" xlink_title="Endangered">EN</abbrev>
) [B 1 ab (iii)] category at present.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESNAC" type="material">
<paragraph id="79529EC783199379F7191847BB07F82F">
<heading id="54CE594CB0B3BA6C4E0E2A0315EEB009" reason="title">
Additional specimens examined (
<typeStatus id="DFB7D6694BCAD66A05E9004E8E532A8A" type="paratype">paratypes</typeStatus>
).
</heading>
</paragraph>
<paragraph id="E94B386779B868A433016C29B72E958A">
<materialsCitation id="4BE9C11F6C40632D7474B04C2FE3A394" collectedFrom="limestone slope" collectingDate="2023-07-19" collectionCode="IBK, KUN, PE" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Dahua County" elevation="880" latitude="24.121" location="Nonghe Village" longLatPrecision="1" longitude="107.72767" municipality="Qibainong Town" specimenCount="1" stateProvince="Guangxi">
<collectingCountry id="3FBEBB403E9B2AA19BF56DDB07FB602D" name="China">China</collectingCountry>
• 
<collectingRegion id="FA6601441878564C735CF9310D965120" country="China" name="Guangxi">Guangxi</collectingRegion>
: 
<collectingCounty id="6C9B54325E83B1AC4B7A22AFF16814E9">Dahua County</collectingCounty>
, 
<collectingMunicipality id="DFEFC0E1B7C7B9B67422B2561A7CDC32">Qibainong Town</collectingMunicipality>
, 
<location id="E59AFB6EAE62BB229134848A3C5C15DC" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:E59AFB6EAE62BB229134848A3C5C15DC" country="China" county="Dahua County" latitude="24.121" longLatPrecision="1" longitude="107.72767" municipality="Qibainong Town" name="Nonghe Village" stateProvince="Guangxi">Nonghe Village</location>
, 
<location id="4ECE03C0596056A1B7B222B80FFC194C" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:4ECE03C0596056A1B7B222B80FFC194C" country="China" county="Dahua County" latitude="24.121" longLatPrecision="1" longitude="107.72767" municipality="Qibainong Town" name="Nongge" stateProvince="Guangxi">Nongge</location>
, around the point 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2NAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.727671,24.121001]}">
<geoCoordinate id="356C7CA127762ED5A8674E0397E6C27D" degrees="24.121001" direction="north" orientation="latitude" precision="1" value="24.121">24.121001 ° N</geoCoordinate>
, 
<geoCoordinate id="10E2C270EDFE4E32960BC05062002901" degrees="107.727671" direction="east" orientation="longitude" precision="1" value="107.72767">107.727671 ° E</geoCoordinate>
</named-content>
, 
<collectedFrom id="059ACF5FB206F62E467F021FF8B75B7E">limestone slope</collectedFrom>
; alt. 
<quantity id="E67608B94287721B467D4A44B4DD7869" metricMagnitude="2" metricUnit="m" metricValue="8.8" unit="m" value="880.0">
<elevation id="62A1CD49647CC14E0D12B4CBE91CD182" metricMagnitude="2" metricUnit="m" metricValue="8.8" unit="m" value="880.0">880 m</elevation>
</quantity>
; 
<date id="B9D2ABA0B38C795B48FF895F3BB9289E" value="2023-07-19">
<collectingDate id="118D601422089C69A49485B5CE9BDE79" value="2023-07-19">19 July 2023</collectingDate>
</date>
; 
<emphasis id="DF52B67B0FDEC00D37B8E4CFFBA84BC7" italics="true">
<collectorName id="835C0B697747FF1DC592A0F30A4A2E88">W. B. Xu</collectorName>
, 
<collectorName id="C78330458DB4B24E77D01C17372BE378">Z. C. Lu</collectorName>
, 
<collectorName id="28B9C9FE35FBEB538D8146DB2528746A">M. L Mo</collectorName>
, 
<collectorName id="DACEF6AEB2ED45CCFA3499EABF90306C">S. L. Chang</collectorName>
&amp; 
<collectorName id="850A2176F18A7849662DAE5F8A469DBD">J. Q. Huang</collectorName>
16285
</emphasis>
(
<collectionCode id="977AD8C03ECE50004764D21A122408B6" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
, 
<collectionCode id="F055654538AEDF7B6434A0A138EC23FC" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15842" lsid="urn:lsid:biocol.org:col:15842" name="Kunming Institute of Botany, Chinese Academy of Sciences" type="Herbarium">KUN</collectionCode>
, 
<collectionCode id="8E7E7C267C8F26D299C723B3C2AD1F2B" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" type="Herbarium">PE</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="0409E7C0ED1F151DC24531D4931FF7FB" collectedFrom="limestone slope" collectingDate="2023-07-19" collectionCode="IBK, CSH, IBSC" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Dahua County" elevation="880" latitude="24.121" location="Nonghe Village" longLatPrecision="1" longitude="107.72767" municipality="Qibainong Town" specimenCount="1" stateProvince="Guangxi">
ibid.; 
<date id="0F34BDF475017EA43632F6FCD114F513" value="2023-07-19">
<collectingDate id="625C6DBFCB281E2DC335C69B9C20F631" value="2023-07-19">19 July 2023</collectingDate>
</date>
; 
<emphasis id="F7AF2FBB709DE8CFF474435548152B41" italics="true">
<collectorName id="ACC3A3870A5459B84CFA1EA38137D96A">W. B. Xu</collectorName>
, 
<collectorName id="E28DE4635678761377121F9116367C4E">Z. C. Lu</collectorName>
, 
<collectorName id="F51414E5E48778FABF45C6341E5ACF73">M. L Mo</collectorName>
, 
<collectorName id="910CB784A74544AC950869CF43C4D868">S. L. Chang</collectorName>
&amp; 
<collectorName id="F304AB72C23C649A92323858B47757BE">J. Q. Huang</collectorName>
16312
</emphasis>
(
<collectionCode id="F46ADD65EBA5B681271D7FAC4EDA8F90" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
, 
<collectionCode id="5C7AAE18F27F4BD6ECC3F385EA112E5D" country="China" name="Chenshan Botanical Garden" type="Herbarium">CSH</collectionCode>
, 
<collectionCode id="5EAF5C4C69F37E82FF138053B3C41BB6" country="China" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" type="Herbarium">IBSC</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="E7B57C298C0619810F0A418DC244D18B" collectedFrom="limestone slope" collectingDate="2023-09-28" collectionCode="IBK" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="769" latitude="24.079994" location="Shangzhen Village" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
<collectingCounty id="9D91B003FEA0E77E674F7694BA5EC054">Du’an County</collectingCounty>
, 
<collectingMunicipality id="A6384BF178BB0B46FAB3681930856D96">Bao’an Town</collectingMunicipality>
, 
<location id="7F23EF0882D66AE33881063179DB250E" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:7F23EF0882D66AE33881063179DB250E" country="China" county="Du'an County" latitude="24.079994" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" name="Shangzhen Village" stateProvince="Guangxi">Shangzhen Village</location>
, 
<location id="F7C9D0A3A0623807D635F2436B538A5D" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:F7C9D0A3A0623807D635F2436B538A5D" country="China" county="Du'an County" latitude="24.079994" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" name="Nongwen" stateProvince="Guangxi">Nongwen</location>
, around the point 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHOAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.828760,24.079994]}">
<geoCoordinate id="263D0F0471666218D9A6548AF7FEDCE7" degrees="24.07999417" direction="north" orientation="latitude" precision="1" value="24.079994">24.07999417 ° N</geoCoordinate>
, 
<geoCoordinate id="8E28D90C32C137AF450E21F5E96B1763" degrees="107.82876" direction="east" orientation="longitude" precision="1" value="107.82876">107.82876 ° E</geoCoordinate>
</named-content>
, 
<collectedFrom id="F7EEACD95DDC62DA65731F6D34CFA3DF">limestone slope</collectedFrom>
; alt. 
<quantity id="568434A3664F0C12FF37BD6E327BB870" metricMagnitude="2" metricUnit="m" metricValue="7.69" unit="m" value="769.0">
<elevation id="195D386A13B486915560C5338AD1E774" metricMagnitude="2" metricUnit="m" metricValue="7.69" unit="m" value="769.0">769 m</elevation>
</quantity>
; 
<date id="C493DE5E2A9AF1AC2379E2D127DE42A6" value="2023-09-28">
<collectingDate id="1EE85CDABAE6DD65BE81DACE74463E68" value="2023-09-28">28 September 2023</collectingDate>
</date>
; 
<emphasis id="C9BAC66C23A31D52ECF655AE70A2EEFB" italics="true">
<collectorName id="A821A92A3F0A0340D1AF8FDCD717CC1C">W. B. Xu</collectorName>
, 
<collectorName id="F30936D12ACFC881D60F0D92212EED11">Z. C. Lu</collectorName>
, 
<collectorName id="897208598022A5898EA92DA7522E8780">M. L Mo</collectorName>
, 
<collectorName id="F9C9CBF08ABCC7DF8C30CC4EA5AF3DF9">S. L. Chang</collectorName>
&amp; 
<collectorName id="CB9177CBE241B5992812DEA93348D78D">J. Q. Huang</collectorName>
17300
</emphasis>
(
<collectionCode id="8AAAA2E2C2689289112B624DE15DEF0A" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="8D61DA9918FF3DC2FFEE71155C023794" collectedFrom="limestone slope" collectingDate="2023-09-28" collectionCode="IBK, GXMG, GXMI" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="769" latitude="24.079994" location="Shangzhen Village" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
ibid.; 
<date id="0F4B223E327BBFA604AC903CD2ABD20D" value="2023-09-28">
<collectingDate id="9799344C28F9FD2EBD577B47A52E6513" value="2023-09-28">28 September 2023</collectingDate>
</date>
; 
<emphasis id="CCA81519578898F2BEB7BAC09D0A0A18" italics="true">
<collectorName id="F2693F226013D40B8EC7D5F0A994C465">W. B. Xu</collectorName>
, 
<collectorName id="30E7647246FC40A61C6442F136AFB357">Z. C. Lu</collectorName>
, 
<collectorName id="08EA209A30AE360A44ABB58FB2BABA5D">M. L Mo</collectorName>
, 
<collectorName id="F63177B40D2DD72638330655607D107B">S. L. Chang</collectorName>
&amp; 
<collectorName id="F6A570691C26D46095AA823FC1CE8534">J. Q. Huang</collectorName>
17301
</emphasis>
(
<collectionCode id="32A4B195717F4B6922F63C0B58BAB8E2" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
, 
<collectionCode id="ED24BD785FB4D7B73D88FED7B80E8356" country="China" lsid="urn:lsid:biocol.org:col:13154" name="Guangxi Medicinal Botanic Garden" type="Herbarium">GXMG</collectionCode>
, 
<collectionCode id="7562AD16D1257A0A0EE601D3F19F40D7" country="China" lsid="urn:lsid:biocol.org:col:13153" name="Guangxi Institute of Traditional Medical and Pharmaceutical Sciences" type="Herbarium">GXMI</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="249FFAEE563AF2B769DB8FD18206284D" collectedFrom="limestone slope" collectingDate="2023-09-28" collectionCode="IBK, GXMI" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="769" latitude="24.079994" location="Shangzhen Village" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
ibid.; 
<date id="B6390D4DCCBECB827B915F2537AC576D" value="2023-09-28">
<collectingDate id="3FBE6002523EF74E3CAE776060B19F0A" value="2023-09-28">28 September 2023</collectingDate>
</date>
; 
<emphasis id="009052E13A261DC7A816907966FC2772" italics="true">
<collectorName id="C2F30238695CBF2D09F49407ED88E38D">W. B. Xu</collectorName>
, 
<collectorName id="7B666C58CB69E8465AC13519D3A56197">Z. C. Lu</collectorName>
, 
<collectorName id="8C25ED0F1357A83BDDBB0BD5C2920559">M. L Mo</collectorName>
, 
<collectorName id="DE80848AB58B80BAC5510E483D3B2F68">S. L. Chang</collectorName>
&amp; 
<collectorName id="F98FE06E1994314A4FDF129229589113">J. Q. Huang</collectorName>
17302
</emphasis>
(
<collectionCode id="65EC16F1607A5E4735316D98879FB925" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
, 
<collectionCode id="E9112CF2D063E7AE3CE8AAAAA2B03BE9" country="China" lsid="urn:lsid:biocol.org:col:13153" name="Guangxi Institute of Traditional Medical and Pharmaceutical Sciences" type="Herbarium">GXMI</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="AB32594ED18DD2174595F5F834546502" collectedFrom="limestone slope" collectingDate="2023-09-28" collectionCode="IBK, GXMG, CSH" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="769" latitude="24.079994" location="Shangzhen Village" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
ibid.; 
<date id="C6BEBE0F3A72AB99CC2F9DA03E2266D5" value="2023-09-28">
<collectingDate id="AF92C8BD853FBEA367C684491C0EC2D7" value="2023-09-28">28 September 2023</collectingDate>
</date>
; 
<emphasis id="6D88618D1624A0A0F5CFADDFE56DD721" italics="true">
<collectorName id="46BA8E3002B35FBAB2142D1464EFBA75">W. B. Xu</collectorName>
, 
<collectorName id="2920F49B66C3FC645C986F396080738B">Z. C. Lu</collectorName>
, 
<collectorName id="11217382C0A32DD81517BF0091FB7493">M. L Mo</collectorName>
, 
<collectorName id="2ACE9FF75A887C430CA2E331696C01D8">S. L. Chang</collectorName>
&amp; 
<collectorName id="29E43CF2DECC1551DE1BA8AA3CD6F8E9">J. Q. Huang</collectorName>
17303
</emphasis>
(
<collectionCode id="BE42FA63734FDBF4278E40B196E88070" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
, 
<collectionCode id="3F9E19D5BC1D12C84E8074A037CA7BC9" country="China" lsid="urn:lsid:biocol.org:col:13154" name="Guangxi Medicinal Botanic Garden" type="Herbarium">GXMG</collectionCode>
, 
<collectionCode id="E0F23F6E7514B9E99BC2CD38DE0EC611" country="China" name="Chenshan Botanical Garden" type="Herbarium">CSH</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="55B9239B38F35BB30D5C7B2B127112E0" collectedFrom="limestone slope" collectingDate="2023-09-28" collectionCode="IBK, PE, KUN" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="769" latitude="24.079994" location="Shangzhen Village" longLatPrecision="1" longitude="107.82876" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
ibid.; 
<date id="5C687B8960238E145138B2C54142654E" value="2023-09-28">
<collectingDate id="BAEFB4596A20F1DF2EB45A9760385472" value="2023-09-28">28 September 2023</collectingDate>
</date>
; 
<emphasis id="69ED6F7A3CB3D062D69C335CEC5C131B" italics="true">
<collectorName id="619030B74EE6DCE91FFD6499EEF659AF">W. B. Xu</collectorName>
, 
<collectorName id="E1B3C3FF9A5E8A81542D6564EE6D694E">Z. C. Lu</collectorName>
, 
<collectorName id="915D0413E5B7D91B0CC0FBF576F76754">M. L Mo</collectorName>
, 
<collectorName id="E6EDB23ADB25C8E5A5C6E45428E24FB2">S. L. Chang</collectorName>
&amp; 
<collectorName id="D06262A121CBE3E753DA2D906A4B9321">J. Q. Huang</collectorName>
17322
</emphasis>
(
<collectionCode id="A324FE248DFEF60509DFBB2BF4DDD96F" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
, 
<collectionCode id="484D2F0B77938C612CAAAE3110BFE513" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" type="Herbarium">PE</collectionCode>
, 
<collectionCode id="E410151B8A2F8A97FE3A90559DD6CE0D" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15842" lsid="urn:lsid:biocol.org:col:15842" name="Kunming Institute of Botany, Chinese Academy of Sciences" type="Herbarium">KUN</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="DFDFD139998F493412A8604CAC1D5BE4" collectedFrom="limestone slope" collectingDate="2023-09-30" collectionCode="IBK" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="860" latitude="24.10944" location="Yuanli Village" longLatPrecision="1" longitude="107.81167" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
<collectingCounty id="2E1B2DE09ACD54418C11E35659ED4F34">Du’an County</collectingCounty>
, 
<collectingMunicipality id="84E39FE8902DEFD8A92C419772A099F3">Bao’an Town</collectingMunicipality>
, 
<location id="40FA5006267705A17A62CF429B91BBB6" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:40FA5006267705A17A62CF429B91BBB6" country="China" county="Du'an County" latitude="24.10944" longLatPrecision="1" longitude="107.81167" municipality="Bao'an Town" name="Yuanli Village" stateProvince="Guangxi">Yuanli Village</location>
, 
<location id="2005AFAEC42F6632023881D6600FBB93" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:2005AFAEC42F6632023881D6600FBB93" country="China" county="Du'an County" latitude="24.10944" longLatPrecision="1" longitude="107.81167" municipality="Bao'an Town" name="Nongkou" stateProvince="Guangxi">Nongkou</location>
, around the point 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZOAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.811669,24.109440]}">
<geoCoordinate id="F759325FC86C3FB8CD45022C96EEEBC2" degrees="24.10944" direction="north" orientation="latitude" precision="1" value="24.10944">24.10944 ° N</geoCoordinate>
, 
<geoCoordinate id="3992D06C509925DDCBADD9727B7C644B" degrees="107.811669" direction="east" orientation="longitude" precision="1" value="107.81167">107.811669 ° E</geoCoordinate>
</named-content>
, 
<collectedFrom id="53D24EC053DB733A1AB07D2AA607AC57">limestone slope</collectedFrom>
; alt. 
<quantity id="2B732E7C5D812C715EA5F98F5F49E803" metricMagnitude="2" metricUnit="m" metricValue="8.6" unit="m" value="860.0">
<elevation id="8418A0EB2EE7E6CCEC9165E6FB1DA45B" metricMagnitude="2" metricUnit="m" metricValue="8.6" unit="m" value="860.0">860 m</elevation>
</quantity>
; 
<date id="C22423875908C835438557C01D6453DC" value="2023-09-30">
<collectingDate id="102E510156BEC219CEADD51EE09A5F82" value="2023-09-30">30 September 2023</collectingDate>
</date>
; 
<emphasis id="EB7E93E4D19D72A66B4D36E9BAFC014A" italics="true">
<collectorName id="6A7AD5DA59F681BCC7FA4E59AE82BBAD">W. B. Xu</collectorName>
, 
<collectorName id="C37A1A309BC7ACA5643FFFB7632C717D">Z. C. Lu</collectorName>
, 
<collectorName id="02829AE0CC4AB638F14003C546707869">M. L Mo</collectorName>
, 
<collectorName id="B651483391F9DA7D4B3ACE89894CE194">S. L. Chang</collectorName>
&amp; 
<collectorName id="6D9DFC32042DFF07E7C9343B53A58204">J. Q. Huang</collectorName>
17367
</emphasis>
(
<collectionCode id="93C4839D0DCA7972153DC2AABDF6F849" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="DD9779E4F8D357AFA5FF0F876B1B5DDC" collectedFrom="limestone slope" collectingDate="2023-09-30" collectionCode="IBK" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Du'an County" elevation="860" latitude="24.10944" location="Yuanli Village" longLatPrecision="1" longitude="107.81167" municipality="Bao'an Town" specimenCount="1" stateProvince="Guangxi">
ibid.; 
<date id="F9F49DEE3A529B11CC41A1AFA8541478" value="2023-09-30">
<collectingDate id="EC73F56EE30E0FE2C9144CA0977BDFA4" value="2023-09-30">30 September 2023</collectingDate>
</date>
; 
<emphasis id="8DAE19CB25A9E311AE7AB59F962B9128" italics="true">
<collectorName id="946304E58ABE42BB95F3046570C11A0E">W. B. Xu</collectorName>
, 
<collectorName id="973BC0F291B2232F0E1BDFD79A55E8DA">Z. C. Lu</collectorName>
, 
<collectorName id="D85E9EF1B710AF55560F1EB2CAFD80D8">M. L Mo</collectorName>
, 
<collectorName id="D178C37F6F12E827776EFEE70BED40DB">S. L. Chang</collectorName>
&amp; 
<collectorName id="29F7E60E502D6326ABD1936F17BBB9FF">J. Q. Huang</collectorName>
17368
</emphasis>
(
<collectionCode id="F5E4F5F36330761B955AD41919F09166" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="352BD370D63E5D116FFCAACFCAE106B1" collectedFrom="limestone slope" collectingDate="2023-10-01" collectionCode="IBK" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Dahua County" elevation="936" latitude="24.22271" location="Nongcong Village" longLatPrecision="1" longitude="107.75722" municipality="Bansheng Town" specimenCount="1" stateProvince="Guangxi">
<collectingCounty id="0E9AEC06AD586AB2E20AB2A0D6FBC083">Dahua County</collectingCounty>
, 
<collectingMunicipality id="690E8D850BF9F4FB0F77D87E040DB824">Bansheng Town</collectingMunicipality>
, 
<location id="F7E290B2BDD2C9B6206B92ECF90A1B40" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:F7E290B2BDD2C9B6206B92ECF90A1B40" country="China" county="Dahua County" latitude="24.22271" longLatPrecision="1" longitude="107.75722" municipality="Bansheng Town" name="Nongcong Village" stateProvince="Guangxi">Nongcong Village</location>
, 
<location id="D170CE03CD5CD5C747002B9EAC57DD77" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:D170CE03CD5CD5C747002B9EAC57DD77" country="China" county="Dahua County" latitude="24.22271" longLatPrecision="1" longitude="107.75722" municipality="Bansheng Town" name="Nongji" stateProvince="Guangxi">Nongji</location>
, around the point 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFPAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.757219,24.222708]}">
<geoCoordinate id="05CBE22F671A2B5867014D2BA698A9A9" degrees="24" direction="north" minutes="13" orientation="latitude" precision="1" seconds="21.75" value="24.22271">24 ° 13 ′ 21.75 ″ N</geoCoordinate>
, 
<geoCoordinate id="E4D4576CCAC6A6E9422C3B2E9EFB2658" degrees="107" direction="east" minutes="45" orientation="longitude" precision="1" seconds="25.99" value="107.75722">107 ° 45 ′ 25.99 ″ E</geoCoordinate>
</named-content>
, 
<collectedFrom id="D29E26AB82E7662C4EEF45A7F41386F4">limestone slope</collectedFrom>
; alt. 
<quantity id="9B1713B31DBBE9788C2829549AB2D4CE" metricMagnitude="2" metricUnit="m" metricValue="9.36" unit="m" value="936.0">
<elevation id="2E34444D06104D672CB57CB0A12F7A87" metricMagnitude="2" metricUnit="m" metricValue="9.36" unit="m" value="936.0">936 m</elevation>
</quantity>
; 
<date id="0DE073AFD863DE78D89D0D8E7635F5FC" value="2023-10-01">
<collectingDate id="C3ACE696A54B4CA2123043F69470B5AF" value="2023-10-01">1 October 2023</collectingDate>
</date>
; 
<emphasis id="E5ED21D9462DB0EA812B1A6094DE1F63" italics="true">
<collectorName id="980D95CC96E003A6CE2E9F3EBA469D55">W. B. Xu</collectorName>
, 
<collectorName id="A917F0E1308AC77E978A0A8F0EACD816">Z. C. Lu</collectorName>
, 
<collectorName id="EDE1760E111B4237E5467E83529AFFA1">M. L Mo</collectorName>
, 
<collectorName id="95CF5E1B87971D02420A2253B15BDAC1">S. L. Chang</collectorName>
&amp; 
<collectorName id="8399F995E36697F86FC7464121FE65D7">J. Q. Huang</collectorName>
17403
</emphasis>
(
<collectionCode id="5D8706729DD077EAA6B82EF4AAA1BD2D" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
)
</materialsCitation>
• 
<materialsCitation id="E5367830F2403C4D3D17C2BC04452EA1" collectedFrom="limestone peak" collectingDate="2023-10-04" collectionCode="IBK" collectorName="W. B. Xu &amp; Z. C. Lu &amp; M. L Mo &amp; S. L. Chang &amp; J. Q. Huang" country="China" county="Dahua County" elevation="920" latitude="24.088762" location="Nongjing Village" longLatPrecision="1" longitude="107.773285" municipality="Qibainong Town" specimenCount="1" stateProvince="Guangxi">
<collectingCounty id="6E59684F3E252D705D21E0AE3BA4A4CB">Dahua County</collectingCounty>
, 
<collectingMunicipality id="9757E20E612B5C3724ABE9E36372B3CF">Qibainong Town</collectingMunicipality>
, 
<location id="89D4E8C84CA88D83F6EBD17E5BA083AE" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:89D4E8C84CA88D83F6EBD17E5BA083AE" country="China" county="Dahua County" latitude="24.088762" longLatPrecision="1" longitude="107.773285" municipality="Qibainong Town" name="Nongjing Village" stateProvince="Guangxi">Nongjing Village</location>
, 
<location id="E0F2763EE3C5E9F2B3D25C8F4D261F31" LSID="urn:lsid:plazi:treatment:F6CAD151C84054CF8014305006360C57:E0F2763EE3C5E9F2B3D25C8F4D261F31" country="China" county="Dahua County" latitude="24.088762" longLatPrecision="1" longitude="107.773285" municipality="Qibainong Town" name="Baxiang" stateProvince="Guangxi">Baxiang</location>
, around the point 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPPAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.773282,24.088763]}">
<geoCoordinate id="A9778928803289099C397526F78974E5" degrees="24.088763" direction="north" orientation="latitude" precision="1" value="24.088762">24.088763 ° N</geoCoordinate>
, 
<geoCoordinate id="15425E19A1EA9A2CFEB3A1AEC220185D" degrees="107.773282" direction="east" orientation="longitude" precision="1" value="107.773285">107.773282 ° E</geoCoordinate>
</named-content>
, 
<collectedFrom id="0D6B4DBE101B9EAC6B446C5825AC085B">limestone peak</collectedFrom>
; alt. 
<quantity id="02E9B427F0DCA5B67A3FEBED4681D5EF" metricMagnitude="2" metricUnit="m" metricValue="9.2" unit="m" value="920.0">
<elevation id="B8DCD10249BEEC6BF38DF4F90EE35825" metricMagnitude="2" metricUnit="m" metricValue="9.2" unit="m" value="920.0">920 m</elevation>
</quantity>
; 
<date id="FF0EE64C2605D1AC1FB0AB799F479DFF" value="2023-10-04">
<collectingDate id="BF48C486CBAEE6A276B9A16F18CDF78A" value="2023-10-04">4 October 2023</collectingDate>
</date>
; 
<emphasis id="E3B47767F4C7E0C0F12E4A65CE9C3ACB" italics="true">
<collectorName id="C328A9EAE240B85378FD24FB8BC0F7F0">W. B. Xu</collectorName>
, 
<collectorName id="F4AD86F634A71CCA69C2B09A5F48DDB5">Z. C. Lu</collectorName>
, 
<collectorName id="78C0C5F03D8E8C38F463DFA6D7A32EC9">M. L Mo</collectorName>
, 
<collectorName id="353C6379E10B258A7B41F98490D854BF">S. L. Chang</collectorName>
&amp; 
<collectorName id="229D48D40BDE5BDA620EB393E2F3C3B1">J. Q. Huang</collectorName>
17503
</emphasis>
(
<collectionCode id="4035179DD932A26F508BD1BA24C57B1A" country="China" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWPAC" type="notes">
<paragraph id="87029E1FC2C825AE4228FB58F284A51D">
<heading id="2CE4F201BA810012457279F8BD500C00" reason="title">Notes.</heading>
</paragraph>
<paragraph id="34D0634EA3A06BD94CB14E9029C183CE">
<taxonomicName id="0EF9687984FB6219CA8B38143C03E489" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="DD0C75A460E2841EF53161F46BC8D60E" italics="true">Tilia saxatilis</emphasis>
</taxonomicName>
has glabrous branchlets, leaf blades base oblique, truncate or cordate, abaxially hairy only in vein axils, bracts narrowly oblong, adnate to inflorescence peduncle, sessile, staminodes 5 and fruits indehiscent. Based on these morphological characters, 
<taxonomicName id="2DCA7CC5252D6AFB636F7E2CB236B9A5" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="C03C1B432460B5EF09466B16B50235DB" italics="true">T. saxatilis</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4102A3AAC8E8AD7DEFCE23A68C774E7E" authority="(Tang et al. 2007)" baseAuthorityName="Tang" baseAuthorityYear="2007" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="tuan">
<emphasis id="988BA0BEF91049C34A8E004FE7825BA2" italics="true">T. tuan</emphasis>
(
<bibRefCitation id="CE28708FACB007E6E16FEF4A48262512" author="Tang" editor="Wu ZY &amp; Raven PH &amp; Hong DY" etAl="et al." firstAuthor="Tang" journalOrPublisher="Vol. 12. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis" pagination="240-248" refId="B7" refString="Tang Y, Gilbert MG, Dorr LJ (2007) Tilia. In: Wu ZY, Raven PH, Hong DY (Eds) Flora of China. Vol. 12. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis, 240–248." title="Tilia." volumeTitle="Flora of China." year="2007">Tang et al. 2007</bibRefCitation>
)
</taxonomicName>
, but differs from the latter in having leaf blades that are oblong or ovate-oblong, margin entire, fruit ellipsoid, 5 - angled, apex acute. The morphological differences between 
<taxonomicName id="C8012CCB7B7F39F8BB1D088BD997A489" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="F0E0DF0CF1AEEC3920CE909CFEB6612F" italics="true">T. saxatilis</emphasis>
</taxonomicName>
and 
<taxonomicName id="D69B4E177574AD17C60FC69172785027" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="tuan">
<emphasis id="D6691DF20A0144FC1132664F14CE3547" italics="true">T. tuan</emphasis>
</taxonomicName>
are shown in Table 
<tableCitation id="D2A612DB746D234E94E2EC946585331A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters distinguishing T. saxatilis from T. tuan." httpUri="http://table.plazi.org/id/9D9DD60B5592F7E887E897217197B9A1" tableUuid="9D9DD60B5592F7E887E897217197B9A1">1</tableCitation>
.
</paragraph>
<caption id="9D9DD60B5592F7E887E897217197B9A1" ID-Table-UUID="9D9DD60B5592F7E887E897217197B9A1" httpUri="http://table.plazi.org/id/9D9DD60B5592F7E887E897217197B9A1" startId="T1" targetIsTable="true">
<paragraph id="2D67E8259FE6F26C94045A3230881EA9">
<label id="3B2C199C6E3C8DFAB1EFC8169E4ED22B">Table 1.</label>
</paragraph>
<paragraph id="D7E323EEF6DA450875F2939F3753E1D0">
Morphological characters distinguishing 
<taxonomicName id="AD8EED05E47685E7CC4DA08C32CAC948" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="E30888E04C094BF2671BA6C8097524D3" italics="true">T. saxatilis</emphasis>
</taxonomicName>
from 
<taxonomicName id="620DBE16B7DE9C8686148C8BE40A54A2" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="tuan">
<emphasis id="F5E379F79FA44EEB8E84022EC2ED63EF" italics="true">T. tuan</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="B4F8CE83218621518C07F4990F840532">
<table id="TID0EXJAE" rules="all">
<tbody id="955705A4196B77D4CB8D55586666BCEA">
<tr id="0AAB960A361361A268D24F39F4B1443C">
<th id="6502183B68C72C60B9135670EDB0D42F" colspan="1" rowspan="1">Characters</th>
<emphasis id="CA53B3033235DFE16982B6CAFAAA63B4" italics="true">
<th id="C030763E3080715C0D045349F43F9DA7" colspan="1" rowspan="1">
<taxonomicName id="D3A5A8B7F40F6DCA4D056C1B4FDD1004" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">T. saxatilis</taxonomicName>
</th>
<taxonomicName id="D340E1A20CFAF85CF0BC8FA6BC51FFE6" authority="Branchlets" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="tuan">
<th id="4F7B9BF21B2282A2D5B3505F55FF1830" colspan="1" rowspan="1">T. tuan</th>
<tr id="934708F51EA41030C9BC244B4D465C00">
<td id="7DDB1A439979DD8E466D09FB0C4AFF8F" colspan="1" rowspan="1">Branchlets</td>
<td id="F2A11B04356F0787821D3FC36D8D092F" colspan="1" rowspan="1">Glabrous</td>
<td id="912B4DA142226D3ADC59DD95546B09F5" colspan="1" rowspan="1">Glabrous or tomentose</td>
</tr>
</taxonomicName>
</emphasis>
</tr>
<tr id="858DB96952546EA598AD97BD6E8F82E3">
<td id="8B1686CEE91D5292EABBC8F05C363EA6" colspan="1" rowspan="1">Leaf blades</td>
<td id="152B371D7E539FBDC248878428DE1F0B" colspan="1" rowspan="1">Oblong or ovate-oblong, margin entire</td>
<td id="8875EDC1F14C0A36DCF444A3863805F0" colspan="1" rowspan="1">Narrowly ovate or ovate-oblong to ovate-orbicular, margin entire or with a few minute teeth near apex or prominently dentate</td>
</tr>
<tr id="E1737D117D6F90B524D7E50F1323E5A0">
<td id="835001B86552BFE1DEAE3B2D48E0988C" colspan="1" rowspan="1">Cymes</td>
<td id="043E9C5FAB533C0A4888227A34B76C39" colspan="1" rowspan="1">5–16 - flowered, 3.5–6 cm long</td>
<td id="EDF78DE114A4BDF4E33C06ACBD9C29AA" colspan="1" rowspan="1">3–22 - flowered, 5–14 cm long</td>
</tr>
<tr id="A033E597A5DF265441D028EFE0A6CE77">
<td id="2D262A061E20C6907D9C62D2174571F3" colspan="1" rowspan="1">Bracts</td>
<td id="8F38A4C92D5E75DA7CB3507AAD806932" colspan="1" rowspan="1">3.8–9 cm long, 1–2 cm wide, sessile</td>
<td id="5F6A2671886BD8E4CDD5B98C26561382" colspan="1" rowspan="1">6–16 cm long, 1–3 cm wide, sessile or with stalks 5–8 mm long</td>
</tr>
<tr id="26C826066D4CF870FAB2DD09B856EC67">
<td id="8E01C25D19917020B609088E98AE7C75" colspan="1" rowspan="1">Pedicel</td>
<td id="DB2A75CEC28D498B82D11F22E7D9346D" colspan="1" rowspan="1">2–4 mm long</td>
<td id="E43912FE16569ACB36C6F66ECDEC5609" colspan="1" rowspan="1">4–9 mm long</td>
</tr>
<tr id="8C3438D9AA6D8837556061C563C7CB20">
<td id="D9581CF30355025EC6A2A83C5330AF0A" colspan="1" rowspan="1">Fruit</td>
<td id="CBD5D9E3B77F0A270851E34E6E539388" colspan="1" rowspan="1">Ellipsoid, 5 - angled, apex acute</td>
<td id="2871FCCE02C2A2FAA6CB1AED606FE231" colspan="1" rowspan="1">Globose or obovoid-globose, not ridged, apex rounded</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="40C68D9C2FFEF5A771384CFC9599B1FA">
In addition, several schemes for subdividing the genus 
<taxonomicName id="8A88082359724ACBEA242CDAB9608A6A" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="B1D954F4FACDF64F1C9759BFE7A9FB38" italics="true">Tilia</emphasis>
</taxonomicName>
have been described (
<bibRefCitation id="DACD95EF1A9F3A7529F17D17055CE008" author="Chang" editor="Chang HT &amp; Miau RH" firstAuthor="Chang" journalOrPublisher="Vol. 49 (1). Science Press, Beijing" pagination="51-76" refId="B1" refString="Chang HT (1989) Tilia. In: Chang HT, Miau RH (Eds) Flora Republicae Popularis Sinicae. Vol. 49 (1). Science Press, Beijing, 51–76." title="Tilia." volumeTitle="Flora Republicae Popularis Sinicae." year="1989">Chang 1989</bibRefCitation>
; 
<bibRefCitation id="89F4401C0BF7C8E919D414B4B6D30381" author="Zhuge and Tang" firstAuthor="Zhuge" issue="4" journalOrPublisher="Journal of Southwest Forestry College" pagination="1-14" refId="B9" refString="Zhuge R, Tang Y (1995) Morphological evolution and biogeography of Tilia. Journal of Southwest Forestry College 15 (4): 1–14." title="Morphological evolution and biogeography of Tilia." volume="15" year="1995">Zhuge and Tang 1995</bibRefCitation>
; 
<bibRefCitation id="79C8412C04E3BF0529CBB3FA0D01418D" DOI="10.1017/CBO9781139033275" author="Pigott" firstAuthor="Pigott" refId="B5" refString="Pigott CD (2012) Lime-trees and basswoods: A biological monograph of the genus Tilia. Cambridge University Press, Cambridge, 45–247. https://doi.org/10.1017/CBO9781139033275" year="2012">Pigott 2012</bibRefCitation>
) and, according to the different morphological characteristics, 
<taxonomicName id="ABCFCC842BB5A6F7CB5E139C24F7E8FD" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="8BF70F8732641A26F8EC0956412E3EAF" italics="true">T. saxatilis</emphasis>
</taxonomicName>
would be placed in different position. 
<bibRefCitation id="E9F0A9AB9C36D4A211DAD969DBD55C17" author="Chang HT" editor="Chang HT &amp; Miau RH" firstAuthor="Chang" journalOrPublisher="Vol. 49 (1). Science Press, Beijing" pagination="51-76" refId="B1" refString="Chang HT (1989) Tilia. In: Chang HT, Miau RH (Eds) Flora Republicae Popularis Sinicae. Vol. 49 (1). Science Press, Beijing, 51–76." title="Tilia." volumeTitle="Flora Republicae Popularis Sinicae." year="1989">Chang (1989)</bibRefCitation>
gave two sections for 
<taxonomicName id="884FB0A933F5B7599788BABD3851F523" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="B3E0DB887DBCAB577F3ADD7DD0A1B66A" italics="true">Tilia</emphasis>
</taxonomicName>
in the &quot; Flora Republicae Popularis Sinicae &quot;; 
<taxonomicName id="E2DDFF07365CA3C5CB725E3120D7DC37" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="53BA267E0C920B2B0D328D6D631D8B90" italics="true">T. saxatilis</emphasis>
</taxonomicName>
would be placed in the sect. 
<taxonomicName id="140B9BA197C337B60835D60BE8FBCAD3" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="EFF02686A41FC5249E363B7B9455A9DA" italics="true">Tilia</emphasis>
</taxonomicName>
L. based on the fruit indehiscent when dry. 
<bibRefCitation id="B5EAF093543EB0287627FBD47B1E897F" author="Zhuge R &amp; Tang Y" firstAuthor="Zhuge" issue="4" journalOrPublisher="Journal of Southwest Forestry College" pagination="1-14" refId="B9" refString="Zhuge R, Tang Y (1995) Morphological evolution and biogeography of Tilia. Journal of Southwest Forestry College 15 (4): 1–14." title="Morphological evolution and biogeography of Tilia." volume="15" year="1995">Zhuge and Tang (1995)</bibRefCitation>
divided 
<taxonomicName id="B0F13838FCF86FF3724BFABBAE01D2C2" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="85A9E65FCD1787671D193263EDB13DFE" italics="true">Tilia</emphasis>
</taxonomicName>
into three sections; 
<taxonomicName id="77E1BCAF5B550E8811757F12E6BDAC7D" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="B4B76A30F5ABE813FE7773186F1F16D0" italics="true">T. saxatilis</emphasis>
</taxonomicName>
would be placed in the sect. 
<taxonomicName id="541894B5392A15D69F39F0FD3692AA85" authority="Reichb." authorityName="Reichb." class="Magnoliopsida" family="Malvaceae" genus="Lindnera" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="FFBBA49F7F7E9AE28FCC360D883FB75D" italics="true">Lindnera</emphasis>
Reichb.
</taxonomicName>
based on the woody fruit exocarp. In 
<bibRefCitation id="C94923D32DC68AFDB34E8632409B0CD6" DOI="10.1017/CBO9781139033275" author="Pigott CD" firstAuthor="Pigott's" refId="B5" refString="Pigott CD (2012) Lime-trees and basswoods: A biological monograph of the genus Tilia. Cambridge University Press, Cambridge, 45–247. https://doi.org/10.1017/CBO9781139033275" year="2012">Pigott’s (2012)</bibRefCitation>
monograph, the genus 
<taxonomicName id="A07C46B7CE375C84FB3CB7E223CF6A1D" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="6CCC240545DF1DC4B6EEB75D2B59A34C" italics="true">Tilia</emphasis>
</taxonomicName>
was divided into four sections; 
<taxonomicName id="DFF8DADBA3DB5121DAC41503A89B2AB3" authorityName="Z. C. Lu &amp; W. B. Xu" authorityYear="2025" class="Magnoliopsida" family="Malvaceae" genus="Tilia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="saxatilis">
<emphasis id="B0F69573F038636ECE61AE092B551C73" italics="true">T. saxatilis</emphasis>
</taxonomicName>
would be placed in the section 
<taxonomicName id="2C851189CB4A43A8F390D39B799F83CF" authority="V. Engler" authorityName="V. Engler" rank="section" section="Astrophilyra">
<emphasis id="F2D784786A463DA942898DAA952740FF" italics="true">Astrophilyra</emphasis>
V. Engler
</taxonomicName>
based on the leaf blades upper surface green, glabrous, lower surface pale green, hairy only in vein axils, fruits not splitting, flowers with staminodes.
</paragraph>
</subSubSection>
</treatment>
</document>