treatments-xml/data/03/A9/87/03A987A8493CFFCC6FB9FC41660EF987.xml

<document id="252238AD60A3F6C3CCF72295E440C1B4" ID-DOI="10.11646/zootaxa.5362.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10149411" ID-ZooBank="9D68D967-AE6A-4ED0-A063-E82E93F2DC3D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700227262364" checkinUser="plazi" docAuthor="Hu, Shao-Ji, Cotton, Adam M., Lamas, Gerardo, Duan, Kuang &amp; Zhang, Xin" docDate="2023" docId="03A987A8493CFFCC6FB9FC41660EF987" docLanguage="en" docName="zootaxa.5362.1.1.pdf.imf" docOrigin="Zootaxa 5362 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5362.1.1/52217" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Graphium agetes subsp. agetes" docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="FF90FFD0492EFFDE6F2EFFEF630BFFF2" masterDocTitle="Checklist of Yunnan Papilionidae (Lepidoptera: Papilionoidea) with nomenclatural notes and descriptions of new subspecies" masterLastPageNumber="69" masterPageNumber="1" pageNumber="19" updateTime="1732743792442" updateUser="felipe">
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<mods:title id="ADF8FBACD1A1C7D11AB78DCA75FB4589">Checklist of Yunnan Papilionidae (Lepidoptera: Papilionoidea) with nomenclatural notes and descriptions of new subspecies</mods:title>
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<mods:namePart id="0ED8A32250D21F98406A2172E68E7680">Hu, Shao-Ji</mods:namePart>
<mods:affiliation id="3E205C1FBF76087F9418C01E5AA2D2E6">Institute of International Rivers and Eco-security, Yunnan University, Kunming, 650500, China &amp; Yunnan Key Laboratory of International Rivers and Transboundary Eco-security, Yunnan University, Kunming, 650500, China</mods:affiliation>
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<mods:namePart id="B5850A3FC279700392415D6708BBB595">Cotton, Adam M.</mods:namePart>
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<mods:affiliation id="5B4A12F548B7FAA65F9E5D839D82E3DB">86 / 2 Moo 5, Tambon Nong Kwai, Hang Dong, Chiang Mai, Thailand https: // orcid. org / 0000 - 0002 - 4852 - 903 X</mods:affiliation>
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<mods:namePart id="B6E1FF3F5A566BD0323619C5EBEA6354">Lamas, Gerardo</mods:namePart>
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<mods:affiliation id="4DCB4341E883E0BB101B09DB1DEAC693">Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima- 14, Peru https: // orcid. org / 0000 - 0002 - 3664 - 6730</mods:affiliation>
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<mods:namePart id="FD70593E348AB808409577212BF64599">Duan, Kuang</mods:namePart>
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<mods:affiliation id="7CA5781903B57464A5DC58FD23F712D2">School of Agriculture, Yunnan University, Kunming, 650500, China https: // orcid. org / 0000 - 0003 - 1123 - 8631</mods:affiliation>
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<mods:namePart id="F72D2EAC59AFAA69F77B603AF34C6F8B">Zhang, Xin</mods:namePart>
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<mods:affiliation id="1A37C13226DE64C65A0A96747C25768F">Institute of International Rivers and Eco-security, Yunnan University, Kunming, 650500, China &amp; Kunming Youning Biotech Co., Ltd, Kunming, 650031, China https: // orcid. org / 0000 - 0002 - 4638 - 3997 Corresponding author: shaojihu @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1615 - 9601 &amp; Institute of International Rivers and Eco-security, Yunnan University, Kunming, 650500, China</mods:affiliation>
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<subSubSection id="C31A6535493CFFCC6FB9FC4161D2FC3A" box="[151,729,942,969]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BBF36BE493CFFCC6FB9FC4161D2FC3A" blockId="18.[151,1437,942,1653]" box="[151,729,942,969]" pageId="18" pageNumber="19">
<emphasis id="B974EAAC493CFFCC6FB9FC4161D2FC3A" bold="true" box="[151,729,942,969]" pageId="18" pageNumber="19">
22a. 
<taxonomicName id="4C004D3D493CFFCC6FFCFC4161D2FC3A" ID-CoL="6L3P4" authority="(Westwood, 1843)" baseAuthorityName="Westwood" baseAuthorityYear="1843" box="[210,729,942,969]" class="Insecta" family="Papilionidae" genus="Graphium" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="agetes">
<emphasis id="B974EAAC493CFFCC6FFCFC4162FFFC3B" bold="true" box="[210,500,942,969]" italics="true" pageId="18" pageNumber="19">Graphium agetes agetes</emphasis>
(Westwood, 1843)
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</emphasis>
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<paragraph id="8BBF36BE493CFFCC6FB9FC3C6167FBE3" blockId="18.[151,1437,942,1653]" pageId="18" pageNumber="19">
<taxonomicName id="4C004D3D493CFFCC6FB9FC3C61A2FC1F" authority="Fruhstorfer, 1901" authorityName="Fruhstorfer" authorityYear="1901" box="[151,681,979,1005]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="tenuilineatas">
<emphasis id="B974EAAC493CFFCC6FB9FC3C62DDFC1F" box="[151,470,979,1005]" italics="true" pageId="18" pageNumber="19">Papilio agetes tenuilineatas</emphasis>
<bibRefCitation id="EF914B4F493CFFCC6ECEFC3C61A2FC1F" author="Fruhstorfer, H." box="[480,681,979,1005]" pageId="18" pageNumber="19" pagination="89 - 90" refId="ref49670" refString="Fruhstorfer, H. (1901) Zwei neue Papilio aus Indochina. Societas Entomologica, 16 (12), 89 - 90." year="1901">Fruhstorfer, 1901</bibRefCitation>
</taxonomicName>
; Soc. ent., 16 (12): 89; TL: ‘Xom-Gom, S ü d-Annam’ [near Phan Rang, 
<collectingRegion id="49C4F85C493CFFCC6FCEFC1862D8FBE3" box="[224,467,1015,1041]" country="Vietnam" name="Ninh Thuan" pageId="18" pageNumber="19">Ninh Thuan Province</collectingRegion>
, S. 
<collectingCountry id="F317762E493CFFCC6ED2FC18616AFBE3" box="[508,609,1015,1041]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
].
</paragraph>
<paragraph id="8BBF36BE493CFFCC6FE1FBF46766FBC7" blockId="18.[151,1437,942,1653]" box="[207,1133,1051,1077]" pageId="18" pageNumber="19">
<emphasis id="B974EAAC493CFFCC6FE1FBF463EEFBC6" box="[207,229,1051,1076]" italics="true" pageId="18" pageNumber="19">P.</emphasis>
[
<emphasis id="B974EAAC493CFFCC6FDAFBF4623EFBC7" box="[244,309,1051,1077]" italics="true" pageId="18" pageNumber="19">apilio</emphasis>
] 
<emphasis id="B974EAAC493CFFCC6E68FBF4612EFBC7" box="[326,549,1051,1077]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C004D3D493CFFCC6E68FBF46285FBC6" authorityName="Westwood" authorityYear="1843" box="[326,398,1051,1076]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="agetes">agetes</taxonomicName>
tenuilineatus
</emphasis>
<bibRefCitation id="EF914B4F493CFFCC6D02FBF461F9FBC7" author="Fruhstorfer, H." box="[556,754,1051,1077]" pageId="18" pageNumber="19" pagination="167 - 234" refId="ref49919" refString="Fruhstorfer, H. (1903) Verzeichnis der in Tonkin, Annam und Siam gesammelten Papilioniden und Besprechung verwandter formen. Berliner Entomologische Zeitschrift, 43 (3 - 4), 167 - 234. https: // doi. org / 10.1002 / mmnd. 47919030406" year="1903">Fruhstorfer, 1903</bibRefCitation>
; Berl. ent. Z., 47 (3/4): 199. [ISS]
</paragraph>
</subSubSection>
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<paragraph id="8BBF36BE493CFFCC6FE9FBD163D3FB8F" blockId="18.[151,1437,942,1653]" pageId="18" pageNumber="19">
<emphasis id="B974EAAC493CFFCC6FE9FBD16268FBAA" bold="true" box="[199,355,1086,1112]" pageId="18" pageNumber="19">Distribution:</emphasis>
found in lowlands and river valleys in W., S., and S.E. Yunnan; also found in Indochina and N.E. 
<collectingCountry id="F317762E493CFFCC6FB9FB8C63DFFB8F" box="[151,212,1123,1149]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
.
</paragraph>
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<subSubSection id="C31A6535493CFFCC6FE9FB68660EF987" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BBF36BE493CFFCC6FE9FB68660EF987" blockId="18.[151,1437,942,1653]" pageId="18" pageNumber="19">
<emphasis id="B974EAAC493CFFCC6FE9FB686203FB53" bold="true" box="[199,264,1159,1185]" pageId="18" pageNumber="19">Note:</emphasis>
<bibRefCitation id="EF914B4F493CFFCC6E3FFB6862EAFB53" author="Chou, I." box="[273,481,1159,1185]" pageId="18" pageNumber="19" refId="ref46886" refString="Chou, I. (Ed.) (1994) Monographia Rhopalocerorum Sinensium (Monograph of Chinese Butterflies). Vol. 1. Henan Science and Technology Publishing House, Zhengzhou, 880 pp." year="1994">Chou &amp; Li (1994)</bibRefCitation>
described 
<taxonomicName id="4C004D3D493CFFCC6D70FB686072FB53" box="[606,889,1159,1185]" class="Insecta" family="Papilionidae" genus="Pathysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="chinensis">
<emphasis id="B974EAAC493CFFCC6D70FB686072FB53" box="[606,889,1159,1185]" italics="true" pageId="18" pageNumber="19">Pathysa agetes chinensis</emphasis>
</taxonomicName>
from 
<collectingRegion id="49C4F85C493CFFCC6CEFFB68674FFB53" box="[961,1092,1159,1185]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
and Daweishan, S.E. 
<collectingRegion id="49C4F85C493CFFCC6A6EFB686692FB53" box="[1344,1433,1159,1185]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
. Examination of the specimen from Daweishan deposited in the Kunming Institute of Zoology, Chinese Academy of Sciences, shows that it belongs to 
<taxonomicName id="4C004D3D493CFFCC6D1AFB2061A5FB1A" authorityName="Chou &amp; Li" authorityYear="1994" box="[564,686,1231,1257]" class="Insecta" family="Papilionidae" genus="Pathysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="agetes">
ssp. 
<emphasis id="B974EAAC493CFFCC6D48FB2061A5FB1A" box="[614,686,1231,1256]" italics="true" pageId="18" pageNumber="19">agetes</emphasis>
</taxonomicName>
, while the specimen from 
<collectingRegion id="49C4F85C493CFFCC6CF9FB206751FB1B" box="[983,1114,1231,1257]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
illustrated in 
<bibRefCitation id="EF914B4F493CFFCC6BDAFB206688FB1B" author="Chou, I." box="[1268,1411,1231,1257]" pageId="18" pageNumber="19" refId="ref46886" refString="Chou, I. (Ed.) (1994) Monographia Rhopalocerorum Sinensium (Monograph of Chinese Butterflies). Vol. 1. Henan Science and Technology Publishing House, Zhengzhou, 880 pp." year="1994">Chou (1994)</bibRefCitation>
is applicable to 
<taxonomicName id="4C004D3D493CFFCC6E02FB1C61D9FAFF" authority="(Mell, 1923)" baseAuthorityName="Mell" baseAuthorityYear="1923" box="[300,722,1267,1293]" class="Insecta" family="Papilionidae" genus="Graphium" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="hoenei">
<emphasis id="B974EAAC493CFFCC6E02FB1C6136FAFF" box="[300,573,1267,1293]" italics="true" pageId="18" pageNumber="19">Graphium agetes hoenei</emphasis>
(
<bibRefCitation id="EF914B4F493CFFCC6D65FB1C61C1FAFF" author="Mell, R." box="[587,714,1267,1293]" pageId="18" pageNumber="19" pagination="153 - 160" refId="ref54080" refString="Mell, R. (1923) Noch unbeschriebene Lepidopteren aus S ¸ dchina II. Deutsche Entomologische Zeitschrift, (2), 153 - 160." year="1923">Mell, 1923</bibRefCitation>
)
</taxonomicName>
. The original description of the new subspecies clearly indicated the characters of the specimen from 
<collectingRegion id="49C4F85C493CFFCC6D1CFAF861B3FAC3" box="[562,696,1303,1329]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
, which is illustrated as ‘ 
<taxonomicName id="4C004D3D493CFFCC6CE7FAF86661FAC3" authority="Chou et Li." authorityName="Chou et Li." authorityYear="2023" box="[969,1386,1303,1329]" class="Insecta" family="Papilionidae" genus="Pathysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" status="subsp. nov." subSpecies="chinensis">
<emphasis id="B974EAAC493CFFCC6CE7FAF867E9FAC3" box="[969,1250,1303,1329]" italics="true" pageId="18" pageNumber="19">Pathysa agetes chinensis</emphasis>
Chou et Li.
</taxonomicName>
<taxonomicNameLabel id="A24757D7493CFFCC6A5FFAF863C1FAA7" pageId="18" pageNumber="19" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
’ in 
<figureCitation id="133B2A3B493CFFCC6FDBFAD46233FAA7" box="[245,312,1339,1365]" captionStart="FIGURE 8" captionStartId="29.[151,250,1782,1806]" captionTargetBox="[343,1243,1039,1758]" captionTargetId="figure-62@29.[343,1243,1039,1758]" captionTargetPageId="29" captionText="FIGURE 8. Male genitalia of Byasa genestieri genestieri (Oberthür, 1918) stat. nov. from Kunming, Yunnan, China.All: whole genitalia with right valve removed, R.: ring, V.: right valve, TSU: tegumen, socii, and uncus, Ae.: aedeagus, Ju.: juxta. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10149425" httpUri="https://zenodo.org/record/10149425/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
on page 752 and with a red dot next to the photo of the same specimen, bottom left on page 173. However, the authors designated the Daweishan specimen (illustrated with a yellow dot on page 173) as 
<typeStatus id="54BB881C493CFFCC6A30FAB0668BFA8B" box="[1310,1408,1375,1401]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
in the original description rather than the 
<collectingRegion id="49C4F85C493CFFCC6D10FA6C61CAFA6F" box="[574,705,1411,1437]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
specimen. Since the 
<specimenCount id="9D06FD37493CFFCC6C8BFA6C6742FA6F" box="[933,1097,1411,1437]" count="2" pageId="18" pageNumber="19" type="generic">two specimens</specimenCount>
belong to different subspecies, we hereby invoke the First Reviser Principle to give priority to the characters stated in the original description over the specimen designated as the 
<typeStatus id="54BB881C493CFFCC6D2FFA24616CFA17" box="[513,615,1483,1509]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
. The illustration of the 
<collectingRegion id="49C4F85C493CFFCC6C5BFA2460F3FA17" box="[885,1016,1483,1509]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
specimen in 
<figureCitation id="133B2A3B493CFFCC6BBCFA2467DEFA17" box="[1170,1237,1483,1509]" captionStart="FIGURE 8" captionStartId="29.[151,250,1782,1806]" captionTargetBox="[343,1243,1039,1758]" captionTargetId="figure-62@29.[343,1243,1039,1758]" captionTargetPageId="29" captionText="FIGURE 8. Male genitalia of Byasa genestieri genestieri (Oberthür, 1918) stat. nov. from Kunming, Yunnan, China.All: whole genitalia with right valve removed, R.: ring, V.: right valve, TSU: tegumen, socii, and uncus, Ae.: aedeagus, Ju.: juxta. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10149425" httpUri="https://zenodo.org/record/10149425/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
on page 752 and photograph of the same specimen on page 173 are thus treated as illustrations of the correct 
<typeStatus id="54BB881C493CFFCC6B8BFA00660CF9FB" box="[1189,1287,1519,1545]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
of the taxon. The depository of this specimen is unknown. As a result, we hereby synonymise 
<taxonomicName id="4C004D3D493CFFCC6B3BF9FC63FDF9A3" authority="Chou and Li, 1994" authorityName="Chou and Li" authorityYear="1994" class="Insecta" family="Papilionidae" genus="Pathysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="chinensis">
<emphasis id="B974EAAC493CFFCC6B3BF9FC6620F9DF" box="[1045,1323,1555,1581]" italics="true" pageId="18" pageNumber="19">Pathysa agetes chinensis</emphasis>
<bibRefCitation id="EF914B4F493CFFCC6A1FF9FC63FDF9A3" author="Chou, I." pageId="18" pageNumber="19" refId="ref46886" refString="Chou, I. (Ed.) (1994) Monographia Rhopalocerorum Sinensium (Monograph of Chinese Butterflies). Vol. 1. Henan Science and Technology Publishing House, Zhengzhou, 880 pp." year="1994">Chou and Li, 1994</bibRefCitation>
</taxonomicName>
with 
<taxonomicName id="4C004D3D493CFFCC6E1AF9D861D1F9A3" authority="(Mell, 1923)" baseAuthorityName="Mell" baseAuthorityYear="1923" box="[308,730,1591,1617]" class="Insecta" family="Papilionidae" genus="Graphium" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" status="syn. nov." subSpecies="hoenei">
<emphasis id="B974EAAC493CFFCC6E1AF9D8614CF9A3" box="[308,583,1591,1617]" italics="true" pageId="18" pageNumber="19">Graphium agetes hoenei</emphasis>
(
<bibRefCitation id="EF914B4F493CFFCC6D7BF9D861D9F9A3" author="Mell, R." box="[597,722,1591,1617]" pageId="18" pageNumber="19" pagination="153 - 160" refId="ref54080" refString="Mell, R. (1923) Noch unbeschriebene Lepidopteren aus S ¸ dchina II. Deutsche Entomologische Zeitschrift, (2), 153 - 160." year="1923">Mell, 1923</bibRefCitation>
)
</taxonomicName>
<emphasis id="B974EAAC493CFFCC6DCFF9D86043F9A3" bold="true" box="[737,840,1591,1617]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A24757D7493CFFCC6DCFF9D86043F9A3" box="[737,840,1591,1617]" pageId="18" pageNumber="19">syn. nov.</taxonomicNameLabel>
</emphasis>
, rather than with the nominate subspecies. 
<taxonomicName id="4C004D3D493CFFCC6A07F9D8623AF987" baseAuthorityName="Mell" baseAuthorityYear="1923" class="Insecta" family="Papilionidae" genus="Graphium" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="agetes" subSpecies="hoenei">
<emphasis id="B974EAAC493CFFCC6A07F9D8623AF987" italics="true" pageId="18" pageNumber="19">Graphium agetes hoenei</emphasis>
</taxonomicName>
does not occur anywhere in 
<collectingRegion id="49C4F85C493CFFCC6D5FF9B461C1F987" box="[625,714,1627,1653]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
, but is found in 
<collectingRegion id="49C4F85C493CFFCC6C50F9B4670AF987" box="[894,1025,1627,1653]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
and northern 
<collectingRegion id="49C4F85C493CFFCC6BB3F9B4660AF987" box="[1181,1281,1627,1653]" country="China" name="Guangxi" pageId="18" pageNumber="19">Guangxi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>