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149 lines
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<document id="0FC0DF3148D1228F8C6949AA0398E513" ID-CLB-Dataset="24943" ID-DOI="10.11646/zootaxa.4695.2.5" ID-GBIF-Dataset="7f34a626-f4fc-4484-83cf-ea8a2fdc963c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3531280" ID-ZooBank="CC111070-3A28-4D52-A26F-B808CC200975" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573110041946" checkinUser="plazi" docAuthor="Sobral, Rafael, De Morais, José W. & Grossi, Paschoal C." docDate="2019" docId="03F74F327A47162167D8FDDEFA12FC3B" docLanguage="en" docName="zootaxa.4695.2.5.pdf" docOrigin="Zootaxa 4695 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colacus Ohaus 1910" docType="key" docVersion="8" lastPageNumber="167" masterDocId="FFCE374A7A4F1629674FFFDFFF8EFF9F" masterDocTitle="A new species of Colacus Ohaus, 1910 (Coleoptera: Scarabaeidae: Dynastinae) from the Mata Seca biotope of Brazil, and notes on Colacus morio Ohaus, 1910" masterLastPageNumber="167" masterPageNumber="159" pageNumber="167" updateTime="1698763786448" updateUser="ExternalLinkService">
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<mods:title id="4204215141BDBD6ED73A4B5967D773FF">A new species of Colacus Ohaus, 1910 (Coleoptera: Scarabaeidae: Dynastinae) from the Mata Seca biotope of Brazil, and notes on Colacus morio Ohaus, 1910</mods:title>
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<mods:namePart id="F2513BF70A962C94FF579131EBA93DFD">Sobral, Rafael</mods:namePart>
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<mods:namePart id="3CFAD86481549A5E26FED28F775FF5FB">De Morais, José W.</mods:namePart>
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<mods:namePart id="FC2546A10448DEF5E2336483D7BA7E17">Grossi, Paschoal C.</mods:namePart>
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<mods:title id="FF9F2150DE7EE56AD23B08C246F13CDC">Zootaxa</mods:title>
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<treatment id="03F74F327A47162167D8FDDEFA12FC3B" ID-DOI="http://doi.org/10.5281/zenodo.5922990" ID-GBIF-Taxon="160164833" ID-Zenodo-Dep="5922990" LSID="urn:lsid:plazi:treatment:03F74F327A47162167D8FDDEFA12FC3B" httpUri="http://treatment.plazi.org/id/03F74F327A47162167D8FDDEFA12FC3B" lastPageNumber="167" pageId="8" pageNumber="167">
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<subSubSection id="C344ADAF7A47162167D8FDDEFDBFFDA0" pageId="8" pageNumber="167" type="nomenclature">
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<paragraph id="8BE1FE247A47162167D8FDDEFDFDFD84" blockId="8.[151,627,513,575]" box="[151,627,513,540]" pageId="8" pageNumber="167">
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<heading id="D0A949487A47162167D8FDDEFDFDFD84" bold="true" box="[151,627,513,540]" fontSize="11" level="1" pageId="8" pageNumber="167" reason="1">
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<emphasis id="B92A22367A47162167D8FDDEFDFDFD84" bold="true" box="[151,627,513,540]" pageId="8" pageNumber="167">
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Key to species of
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<taxonomicName id="4C5E85A77A4716216621FDDEFDFDFD84" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" box="[366,627,513,540]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="genus">
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<emphasis id="B92A22367A4716216621FDDEFE5EFD83" bold="true" box="[366,464,513,540]" italics="true" pageId="8" pageNumber="167">Colacus</emphasis>
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Ohaus, 1910
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE1FE247A47162167D8FDFAFDBFFDA0" blockId="8.[151,627,513,575]" box="[151,561,548,575]" pageId="8" pageNumber="167">
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(modified from
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<bibRefCitation id="EFCF83D57A4716216605FDFAFDA4FDA0" author="Neita-Moreno, J. C." box="[330,554,548,575]" pageId="8" pageNumber="167" pagination="187 - 199" refId="ref3531" refString="Neita-Moreno, J. C. (2015) Revision del genero Colacus Ohaus (Coleoptera: Scarabaeidae: Dynastinae: Agaocephalini). Dugesiana, 22, 187 - 199." type="journal article" year="2015">Neita-Moreno 2015</bibRefCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C344ADAF7A47162167D8FDB5FA12FC3B" pageId="8" pageNumber="167" type="key">
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<keyStep id="30AAE6817A47162167D8FDB5FA12FD26" pageId="8" pageNumber="167">
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<paragraph id="8BE1FE247A47162167D8FDB5FA12FD03" blockId="8.[151,1437,617,933]" pageId="8" pageNumber="167">
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<keyLead id="30AF5D117A47162167D8FDB5FA12FD03" pageId="8" pageNumber="167">1. Head, pronotum, and elytra black. Pronotum with a pair of tubercles and a small, anterior fovea. Elytra rugopunctate, punctures strongly impressed.................................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="8BE1FE247A47162167D8FD7AFA12FD26" blockId="8.[151,1437,617,933]" box="[151,1436,676,698]" pageId="8" pageNumber="167">
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<keyLead id="30AF5D117A47162167D8FD7AFA12FD26" box="[151,1436,676,698]" pageId="8" pageNumber="167">- Head and pronotum black, elytra yellow or red. Pronotum lacking tubercles. Elytra punctate, punctures shallow.......... 3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30AAE6817A47162167D8FD1DFA13FCB0" pageId="8" pageNumber="167">
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<paragraph id="8BE1FE247A47162167D8FD1DFA12FD6B" blockId="8.[151,1437,617,933]" pageId="8" pageNumber="167">
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<keyLead id="30AF5D117A47162167D8FD1DFA12FD6B" pageId="8" pageNumber="167">
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2. Cephalic tubercle entire, sharp, conical (
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<figureCitation id="1365E2A17A471621653EFD1EFD4FFD48" box="[625,705,705,727]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: A–C, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="8" pageNumber="167">Fig. 7D</figureCitation>
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). Femora ventrally black (
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<figureCitation id="1365E2A17A4716216489FD1EFB9BFD48" box="[966,1045,705,727]" captionStart="FIGURE 6" captionStartId="6.[151,250,1987,2012]" captionTargetBox="[157,1424,812,1955]" captionTargetId="figure@6.[151,1435,803,1966]" captionTargetPageId="6" captionText="FIGURE 6. Female of Colacus morio. A, Habitus in dorsal view; B, habitus in ventral view. Scale bars: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531292" httpUri="https://zenodo.org/record/3531292/files/figure.png" pageId="8" pageNumber="167">Fig. 6B</figureCitation>
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). External margin of mandibles slightly concave (
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<figureCitation id="1365E2A17A4716216671FD00FE09FD6B" box="[318,391,735,757]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: A–C, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="8" pageNumber="167">Fig. 7B</figureCitation>
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)..............................................................
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<taxonomicName id="4C5E85A77A47162163C8FD00FA12FD6B" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" box="[1159,1436,735,757]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="morio">
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<emphasis id="B92A22367A47162163C8FD00FA12FD6B" bold="true" box="[1159,1436,735,757]" pageId="8" pageNumber="167">
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<emphasis id="B92A22367A47162163C8FD00FA96FD6B" bold="true" box="[1159,1304,735,757]" italics="true" pageId="8" pageNumber="167">Colacus morio</emphasis>
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Ohaus, 1910
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BE1FE247A47162167D8FD22FA13FCB0" blockId="8.[151,1437,617,933]" pageId="8" pageNumber="167">
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<keyLead id="30AF5D117A47162167D8FD22FA13FCB0" pageId="8" pageNumber="167">
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- Cephalic tubercle apically emarginate, compressed, transverse (
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<figureCitation id="1365E2A17A471621640DFD23FC1EFC8D" box="[834,912,764,786]" captionStart="FIGURE 2" captionStartId="2.[151,250,1473,1498]" captionTargetBox="[188,1399,309,1454]" captionTargetId="figure@2.[188,1399,301,1454]" captionTargetPageId="2" captionText="FIGURE 2. Head and pronotum of Colacus rubrofemoratus. A, Head in frontal view, white arrow indicating the apex of cephalic tubercle; B, head and anterior portion of pronotum in dorsal view, white arrow indicating the pronotal fovea; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrow indicating pronotal tubercles. Scale bars: A–C, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531284" httpUri="https://zenodo.org/record/3531284/files/figure.png" pageId="8" pageNumber="167">Fig. 2A</figureCitation>
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). Femora ventrally dark red (
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<figureCitation id="1365E2A17A47162163EAFD23FB7EFC8D" box="[1189,1264,764,786]" captionStart="FIGURE 1" captionStartId="1.[151,250,1968,1993]" captionTargetBox="[151,1435,184,1949]" captionTargetId="figure@1.[151,1435,174,1949]" captionTargetPageId="1" captionText="FIGURE 1. Male of Colacus rubrofemoratus. A, Habitus in dorsal view, black arrow indicating the fourth protibial tooth; B, habitus in ventral view; C, living specimen on a log. Scale bars: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531282" httpUri="https://zenodo.org/record/3531282/files/figure.png" pageId="8" pageNumber="167">Fig. 1B</figureCitation>
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). External margin of mandibles convex (
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<figureCitation id="1365E2A17A47162166F8FCC6FE71FCAF" box="[439,511,793,816]" captionStart="FIGURE 3" captionStartId="3.[151,250,1911,1936]" captionTargetBox="[184,1403,841,1891]" captionTargetId="figure@3.[184,1403,841,1891]" captionTargetPageId="3" captionText="FIGURE 3. Mouthparts of Colacus rubrofemoratus. A, Left mandible in dorsal view; B, left mandible in ventral view, white arrows indicating shallow punctures on mandible; C, left maxilla in ventral view, black arrow indicating maxillary palpifer; D, labrum in dorsal view; E, head in ventral view, black arrow indicating outer edge of right mandible, white arrow indicating punctures on mentum. Scale bars: A–D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3531286" httpUri="https://zenodo.org/record/3531286/files/figure.png" pageId="8" pageNumber="167">Fig. 3E</figureCitation>
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).......................
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<emphasis id="B92A22367A471621646EFCC6FA13FCB0" bold="true" box="[801,1437,793,815]" pageId="8" pageNumber="167">
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<taxonomicName id="4C5E85A77A471621646EFCC6FB3AFCB0" authority="Sobral, Morais" authorityName="Sobral, Morais" box="[801,1204,793,815]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="rubrofemoratus">
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<emphasis id="B92A22367A471621646EFCC6FB9DFCB0" bold="true" box="[801,1043,793,815]" italics="true" pageId="8" pageNumber="167">Colacus rubrofemoratus</emphasis>
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Sobral, Morais
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</taxonomicName>
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, & Grossi, new species
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</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30AAE6817A47162167D8FCE8FA12FC3B" pageId="8" pageNumber="167">
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<paragraph id="8BE1FE247A47162167D8FCE8FA12FCF6" blockId="8.[151,1437,617,933]" pageId="8" pageNumber="167">
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<keyLead id="30AF5D117A47162167D8FCE8FA12FCF6" pageId="8" pageNumber="167">
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3. Clypeal apex sharp, bristles absent. Labium, in lateral view, slightly convex. Lateral edges of paraglossae as long as labial palpomere I (see Fig. 10a in
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<bibRefCitation id="EFCF83D57A47162166BAFC8BFD6BFCF5" author="Neita-Moreno, J. C." box="[501,741,852,874]" pageId="8" pageNumber="167" pagination="187 - 199" refId="ref3531" refString="Neita-Moreno, J. C. (2015) Revision del genero Colacus Ohaus (Coleoptera: Scarabaeidae: Dynastinae: Agaocephalini). Dugesiana, 22, 187 - 199." type="journal article" year="2015">Neita-Moreno 2015: 193</bibRefCitation>
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)................................
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<taxonomicName id="4C5E85A77A4716216333FC8BFA12FCF6" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" box="[1148,1436,852,874]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis id="B92A22367A4716216333FC8BFA12FCF6" bold="true" box="[1148,1436,852,874]" pageId="8" pageNumber="167">
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<emphasis id="B92A22367A4716216333FC8BFA99FCF5" bold="true" box="[1148,1303,852,874]" italics="true" pageId="8" pageNumber="167">Colacus bicolor</emphasis>
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Ohaus, 1910
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BE1FE247A47162167D8FCADFA12FC3B" blockId="8.[151,1437,617,933]" pageId="8" pageNumber="167">
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<keyLead id="30AF5D117A47162167D8FCADFA12FC3B" pageId="8" pageNumber="167">
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- Clypeal apex truncate, slender bristles present. Labium, in lateral view, strongly convex. Lateral edges of paraglossae shorter than labial palpomere I (see Fig. 29a in
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<bibRefCitation id="EFCF83D57A471621652FFC50FCC1FC3A" author="Neita-Moreno, J. C." box="[608,847,911,933]" pageId="8" pageNumber="167" pagination="187 - 199" refId="ref3531" refString="Neita-Moreno, J. C. (2015) Revision del genero Colacus Ohaus (Coleoptera: Scarabaeidae: Dynastinae: Agaocephalini). Dugesiana, 22, 187 - 199." type="journal article" year="2015">Neita-Moreno 2015: 197</bibRefCitation>
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)................
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<taxonomicName id="4C5E85A77A4716216362FC50FA12FC3B" authority="Neita-Moreno, 2015" authorityName="Neita-Moreno" authorityYear="2015" box="[1069,1436,911,933]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="moroni">
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<emphasis id="B92A22367A4716216362FC50FA12FC3B" bold="true" box="[1069,1436,911,933]" pageId="8" pageNumber="167">
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<emphasis id="B92A22367A4716216362FC50FB45FC3B" bold="true" box="[1069,1227,911,933]" italics="true" pageId="8" pageNumber="167">Colacus moroni</emphasis>
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<bibRefCitation id="EFCF83D57A471621639EFC50FA12FC3B" author="Neita-Moreno, J. C." box="[1233,1436,911,933]" pageId="8" pageNumber="167" pagination="187 - 199" refId="ref3531" refString="Neita-Moreno, J. C. (2015) Revision del genero Colacus Ohaus (Coleoptera: Scarabaeidae: Dynastinae: Agaocephalini). Dugesiana, 22, 187 - 199." type="journal article" year="2015">Neita-Moreno, 2015</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document>
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