treatments-xml/data/4F/0C/86/4F0C8664E74B57D0B692C90D9AE9FBC9.xml

<document ID-CLB-Dataset="100335" ID-DOI="http://dx.doi.org/10.3897/zookeys.1113.82172" ID-GBIF-Dataset="3fc6193c-cd79-4419-a24e-7c5545e0bd30" ID-PMC="PMC9848802" ID-Pensoft-Pub="1313-2970-1113-1" ID-Pensoft-UUID="EB46BF265F2E51B3A83799886C5F084A" ID-PubMed="36762231" ModsDocID="1313-2970-1113-1" checkinTime="1658252710949" checkinUser="pensoft" docAuthor="Bribiesca-Contreras, Guadalupe, Dahlgren, Thomas G., Amon, Diva J., Cairns, Stephen, Drennan, Regan, Durden, Jennifer M., Eleaume, Marc P., Hosie, Andrew M., Kremenetskaia, Antonina, McQuaid, Kirsty, O'Hara, Timothy D., Rabone, Muriel, Simon-Lledo, Erik, Smith, Craig R., Watling, Les, Wiklund, Helena &amp; Glover, Adrian G." docDate="2022" docId="4F0C8664E74B57D0B692C90D9AE9FBC9" docLanguage="en" docName="ZooKeys 1113: 1-110" docOrigin="ZooKeys 1113" docPubDate="2022-07-18" docSource="http://dx.doi.org/10.3897/zookeys.1113.82172" docTitle="Calyptrophora distolos Cairns 2018" docType="treatment" docVersion="5" id="EB46BF265F2E51B3A83799886C5F084A" lastPageNumber="1" masterDocId="EB46BF265F2E51B3A83799886C5F084A" masterDocTitle="Benthic megafauna of the western Clarion-Clipperton Zone, Pacific Ocean" masterLastPageNumber="110" masterPageNumber="1" pageNumber="1" updateTime="1732669336981" updateUser="ExternalLinkService">
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<mods:title id="AA0DCD782A9110E22027A29905565C76">Benthic megafauna of the western Clarion-Clipperton Zone, Pacific Ocean</mods:title>
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<mods:namePart id="28C1444B7215D1DB499776DD43E1786E">Bribiesca-Contreras, Guadalupe</mods:namePart>
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<mods:namePart id="EB5FC7F52A3E5F828EDD755F73095D21">Dahlgren, Thomas G.</mods:namePart>
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<mods:namePart id="D8E859790371AAF768EA5DCF93E7AF8A">Amon, Diva J.</mods:namePart>
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<mods:affiliation id="16D6F12822CCBA7A32BF26A2A53A4B44">Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D. C., USA</mods:affiliation>
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<mods:affiliation id="D8AB415A3EEA0044A9DB762218C0483A">Department of Marine Sciences, University of Gothenburg, Gothenburg, Sweden</mods:affiliation>
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<paragraph id="3896FE3300F3D41CEE52835CEF9EFCF3" pageId="0" pageNumber="1">
<taxonomicName id="184955ACFC48764AD6122F6749F4328C" ID-CoL="6999F" LSID="4F0C8664-E74B-57D0-B692-C90D9AE9FBC9" authority="Cairns, 2018" authorityName="Cairns" authorityYear="2018" class="Anthozoa" family="Primnoidae" genus="Calyptrophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptrophora distolos" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="distolos">Calyptrophora distolos Cairns, 2018</taxonomicName>
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<paragraph id="F0FF0845F9E2EB72B57EAD92917EECE9" pageId="0" pageNumber="1">
<figureCitation id="2D610123B6A898B351E35715F2DADBE0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Calyptrophora distolos Cairns, 2018. Specimen CCZ _ 132: A in situ image B whole colony C detail of polyps before preservation. Scale bars: 2 cm (A, B); 5 mm (C). Image attribution: Durden and Smith (A); Wiklund, Durden, Drennan, and McQuaid (B, C)." figureDoi="10.3897/zookeys.1113.82172.figure18" httpUri="https://binary.pensoft.net/fig/716344" pageId="0" pageNumber="1">Fig. 18</figureCitation>
</paragraph>
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<paragraph id="177DED2E968BE65E54716314E46B880B" pageId="0" pageNumber="1">Material.</paragraph>
<paragraph id="EA216E14F5C44177143740E5988553E0" pageId="0" pageNumber="1">
<materialsCitation id="C912370D4BCA961695D06C5A9CAC47EE" accessionNumber="ON400712, ON406604" collectingDate="2018-06-04" collectorName="Smith, Durden, GenBank, Voucher" elevation="4125" latitude="7.2874" longLatPrecision="7" longitude="-149.8578" specimenCount="1">
Clarion-Clipperton Zone • 
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; APEI 4; 
<geoCoordinate id="2F2C0C12EC52A09D03F30FAD58E7DB67" degrees="7.2874" direction="north" orientation="latitude" precision="5" value="7.2874">7.2874°N</geoCoordinate>
, 
<geoCoordinate id="594C157E83090E7E3D480557767D0E9C" degrees="149.8578" direction="west" orientation="longitude" precision="5" value="-149.8578">149.8578°W</geoCoordinate>
; 
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<elevation id="C059ED54FE9C3CDFB03017204A105EAB" metricMagnitude="3" metricUnit="m" metricValue="4.125" unit="m" value="4125.0">4125 m</elevation>
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deep; 
<collectingDate id="3245C580D56E55DAA6A31A8B14577EBB" value="2018-06-04">04 Jun. 2018</collectingDate>
; 
<collectorName id="CCBE33EA6EDEFB963C3B63E7E0DDF420">Smith</collectorName>
&amp; 
<collectorName id="EEEAD674F4F001BAC56408A13F6CAD2C">Durden</collectorName>
leg.; 
<collectorName id="A7696059A3EF68448CC5F9B26B2BCF2D">GenBank</collectorName>
: 
<accessionNumber id="C9693E54BF09E2C435C02232259ACA0F" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/ON400712">ON400712</accessionNumber>
(COI), 
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(16S); USNM 1550968; 
<collectorName id="82D02518410B3F8312D368DE73B53452">Voucher</collectorName>
: CCZ_131
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.
</paragraph>
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<paragraph id="2E65F5D3987D7DBA124F49B86023775F" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="FEEE41A161C6753D8868C3BB83DC8277" pageId="0" pageNumber="1">
Branching uniplanar, colony ~ 20.8 cm tall, with polyps perpendicular to the stem in in situ images (Fig. 
<figureCitation id="013CDF5A4D692472784617ACD80DED23" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Calyptrophora distolos Cairns, 2018. Specimen CCZ _ 132: A in situ image B whole colony C detail of polyps before preservation. Scale bars: 2 cm (A, B); 5 mm (C). Image attribution: Durden and Smith (A); Wiklund, Durden, Drennan, and McQuaid (B, C)." figureDoi="10.3897/zookeys.1113.82172.figure18" httpUri="https://binary.pensoft.net/fig/716344" pageId="0" pageNumber="1">18A</figureCitation>
). Downward-oriented polyps, arranged parallel to the branch, mostly paired, but a few whorls with three to four polyps are present; polyps are ~ 2.7 mm tall and with an operculum longer than either of the body wall scales (Fig. 
<figureCitation id="CD41A179644E0519C5A2D148987A82D0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Calyptrophora distolos Cairns, 2018. Specimen CCZ _ 132: A in situ image B whole colony C detail of polyps before preservation. Scale bars: 2 cm (A, B); 5 mm (C). Image attribution: Durden and Smith (A); Wiklund, Durden, Drennan, and McQuaid (B, C)." figureDoi="10.3897/zookeys.1113.82172.figure18" httpUri="https://binary.pensoft.net/fig/716344" pageId="0" pageNumber="1">18B, C</figureCitation>
).
</paragraph>
<caption id="5D832467BCBC78ADB414C9695AA66B54" doi="10.3897/zookeys.1113.82172.figure18" httpUri="https://binary.pensoft.net/fig/716344" pageId="0" pageNumber="1" start="Figure 18" startId="F18">
<paragraph id="0D99BC4C8D8F04C66928A89533522A0E" pageId="0" pageNumber="1">
<emphasis id="241DE30FAC3590017EFC8044CD8B4897" bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
<taxonomicName id="BC919E90088A61F203721DFE15F52C50" authorityName="Cairns" authorityYear="2018" class="Anthozoa" family="Primnoidae" genus="Calyptrophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptrophora distolos" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="distolos">
<emphasis id="1320434B957F33D9D3B868365D3B02BA" italics="true" pageId="0" pageNumber="1">Calyptrophora distolos</emphasis>
</taxonomicName>
Cairns, 2018. Specimen CCZ_132: 
<emphasis id="6486B9E5E3C98F398A42EA3706480DA6" bold="true" pageId="0" pageNumber="1">A</emphasis>
in situ image 
<emphasis id="D1D63326585FE121CCC2D1B031C30B04" bold="true" pageId="0" pageNumber="1">B</emphasis>
whole colony 
<emphasis id="FC30D676B32C8AFFC7E5EB1D92E56B19" bold="true" pageId="0" pageNumber="1">C</emphasis>
detail of polyps before preservation. Scale bars: 2 cm (
<emphasis id="967F91CAEFFF12CD07DE3F7BAA9CB5A8" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
); 5 mm (
<emphasis id="C191E300710E973762A71A35D911D01A" bold="true" pageId="0" pageNumber="1">C</emphasis>
). Image attribution: Durden and Smith (
<emphasis id="DCB6695C29725C27FD2626204E1B424B" bold="true" pageId="0" pageNumber="1">A</emphasis>
); Wiklund, Durden, Drennan, and McQuaid (
<emphasis id="98E49C6C262E42AFD6A3EF7D7CA57061" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1E4ED1B68757A016703DD99F71257774" pageId="0" pageNumber="1" type="remarks">
<paragraph id="59BAB6E7E222DE0E7F1816FE86A57541" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="3404AA7393E2CC50D9807AD738EB2531" pageId="0" pageNumber="1">
Morphological characters are concordant with the description of 
<taxonomicName id="447C5905BC48BCA276ED158FA28978F3" authorityName="Cairns" authorityYear="2018" class="Anthozoa" family="Primnoidae" genus="Calyptrophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptrophora distolos" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="distolos">
<emphasis id="8B8AE8DAC5449227F8450B8F3DC4E775" italics="true" pageId="0" pageNumber="1">Calyptrophora distolos</emphasis>
</taxonomicName>
(
<bibRefCitation id="205B3E5AB05122CB9060A402AECACE42" DOI="https://doi.org/10.11646/zootaxa.4532.1.1" author="Cairns, SD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 43" refId="B13" refString="Cairns, SD, 2018. Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific. Zootaxa 4532 (1): 1 - 43, DOI: https://doi.org/10.11646/zootaxa.4532.1.1" title="Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific." url="https://doi.org/10.11646/zootaxa.4532.1.1" volume="4532" year="2018">Cairns 2018</bibRefCitation>
). In addition to the paired polyps mentioned in the species description, this specimen also presents a few whorls with three or four polyps (Fig. 
<figureCitation id="9AADDB5828C9D27037A7D2F2C7B50D8B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Calyptrophora distolos Cairns, 2018. Specimen CCZ _ 132: A in situ image B whole colony C detail of polyps before preservation. Scale bars: 2 cm (A, B); 5 mm (C). Image attribution: Durden and Smith (A); Wiklund, Durden, Drennan, and McQuaid (B, C)." figureDoi="10.3897/zookeys.1113.82172.figure18" httpUri="https://binary.pensoft.net/fig/716344" pageId="0" pageNumber="1">18C</figureCitation>
). Polyps are downward-oriented, therefore belonging to the 
<emphasis id="61FA29BEF6FCCEF2364B40F64D712CE2" italics="true" pageId="0" pageNumber="1">wyvillei</emphasis>
complex (
<bibRefCitation id="A41B224200C7568E5EE0995DF2F1A955" DOI="https://doi.org/10.11646/zootaxa.4532.1.1" author="Cairns, SD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 43" refId="B13" refString="Cairns, SD, 2018. Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific. Zootaxa 4532 (1): 1 - 43, DOI: https://doi.org/10.11646/zootaxa.4532.1.1" title="Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific." url="https://doi.org/10.11646/zootaxa.4532.1.1" volume="4532" year="2018">Cairns 2018</bibRefCitation>
). The species is most similar to 
<taxonomicName id="40B625A23B246A18BBB706A6294B0BFD" lsidName="C. persephone" pageId="0" pageNumber="1" rank="species" species="persephone">
<emphasis id="D87CDED27CA0A3BDD8F38CF614E7A833" italics="true" pageId="0" pageNumber="1">C. persephone</emphasis>
</taxonomicName>
Cairns, 2015, which has been described for the UK-1 and BGR areas in the CCZ (
<bibRefCitation id="A4FA83EB2975E4A792FA978A9519430F" DOI="https://doi.org/10.1007/s12526-015-0340-x" author="Cairns, SD" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" pagination="141 - 150" refId="B12" refString="Cairns, SD, 2015. New abyssal Primnoidae (Anthozoa: Octocorallia) from the Clarion-Clipperton Fracture Zone, equatorial northeastern Pacific. Marine Biodiversity 46 (1): 141 - 150, DOI: https://doi.org/10.1007/s12526-015-0340-x" title="New abyssal Primnoidae (Anthozoa: Octocorallia) from the Clarion-Clipperton Fracture Zone, equatorial northeastern Pacific." url="https://doi.org/10.1007/s12526-015-0340-x" volume="46" year="2015">Cairns 2015</bibRefCitation>
). However, 
<taxonomicName id="6F5D22FDA25BC34F499509CE559FB681" lsidName="C. persephone" pageId="0" pageNumber="1" rank="species" species="persephone">
<emphasis id="C4DAB2DA71C31BD163D9A899BE158D8C" italics="true" pageId="0" pageNumber="1">C. persephone</emphasis>
</taxonomicName>
is characterised as having polyps oriented upwards, therefore belonging to the 
<emphasis id="A90381AF43FA296E4711F2959666E619" italics="true" pageId="0" pageNumber="1">japonica</emphasis>
complex, and that are consistently arranged in whorls of three or four, with each basal scale bearing two prominent distal spines. 
<taxonomicName id="68E578DFD8EAC97B4DCBFDF487E90378" authorityName="Cairns" authorityYear="2018" class="Anthozoa" family="Primnoidae" genus="Calyptrophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptrophora distolos" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="distolos">
<emphasis id="DE32B33488B6163E350A734AC1EF7854" italics="true" pageId="0" pageNumber="1">Calyptrophora distolos</emphasis>
</taxonomicName>
was described from the Enigma Seamount, south of Guam, at 3737 m depth, and has also been recorded for American Samoa at 2994 m depth (
<bibRefCitation id="103B40CBDDE1F8D96D7E7DCCC6723F05" DOI="https://doi.org/10.11646/zootaxa.4532.1.1" author="Cairns, SD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 43" refId="B13" refString="Cairns, SD, 2018. Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific. Zootaxa 4532 (1): 1 - 43, DOI: https://doi.org/10.11646/zootaxa.4532.1.1" title="Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific." url="https://doi.org/10.11646/zootaxa.4532.1.1" volume="4532" year="2018">Cairns 2018</bibRefCitation>
). There are no genetic sequences available for other specimens of 
<taxonomicName id="E02391281C4B5967A554784ADFC4E107" lsidName="C. distolos" pageId="0" pageNumber="1" rank="species" species="distolos">
<emphasis id="7A119E00BBD347447F10E5B24C9DDC58" italics="true" pageId="0" pageNumber="1">C. distolos</emphasis>
</taxonomicName>
, but the sequences generated here cluster with other species of the genus (Fig. 
<figureCitation id="B8B3BC18C3E0DCBA3D27CCCEAFC76A9D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rooted Bayesian phylogeny of Octocorallia. Concatenated (16 S, COI, mtMutS, NADH 2) median consensus BEAST tree with posterior probability (PP) and bootstrap (BS) values indicated. Only values of PP&gt; 0.70 and BS&gt; 50 are shown, with values of PP&gt; 0.95 and BS&gt; 90 indicated with a circle. Nodes not recovered on the RAxML tree are indicated with a hyphen. Sequences generated in this study are highlighted in violet." figureDoi="10.3897/zookeys.1113.82172.figure15" httpUri="https://binary.pensoft.net/fig/716341" pageId="0" pageNumber="1">15</figureCitation>
). However, the genus was not recovered as monophyletic.
</paragraph>
</subSubSection>
<subSubSection id="4EC553EED1C4E195F8230B9ECD3EBBA2" pageId="0" pageNumber="1" type="ecology">
<paragraph id="E956C5FDA3B91A557B88E20252B946B3" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="0E68FD24EE1021D8C4A3061298C37A87" pageId="0" pageNumber="1">The specimen was found attached to a polymetallic crust on the slope of a seamount on APEI 4, at 4124 m depth.</paragraph>
</subSubSection>
<subSubSection id="21D82F1F817C2D1D183746C70CBB0358" pageId="0" pageNumber="1" type="comparisons">
<paragraph id="83B5FD7796413E43CD686A2ED7B8787C" pageId="0" pageNumber="1">Comparison with image-based catalogue.</paragraph>
<paragraph id="B8BC47B5AB73706FFEBA823BE2E93B38" pageId="0" pageNumber="1">
A similar primnoid morphotype (i.e., 
<taxonomicName id="FF98A4C51A680DB1C165360BAE943C30" authorityName="Cairns" authorityYear="2018" class="Anthozoa" family="Primnoidae" genus="Calyptrophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptrophora distolos" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="distolos">
<emphasis id="FF339DB087F7379F2D01A01A99E7CFCC" italics="true" pageId="0" pageNumber="1">Calyptrophora distolos</emphasis>
</taxonomicName>
sp. inc., ALC_016) was catalogued from seabed imagery (also collected on a seamount) in the eastern CCZ (e.g., 
<bibRefCitation id="4E743C1673893D5D72EF9AD67C3EC096" DOI="https://doi.org/10.5194/bg-17-2657-2020" author="Cuvelier, D" journalOrPublisher="Annual Review of Marine Science" pageId="0" pageNumber="1" refId="B19" refString="Cuvelier, D, Ribeiro, PA, Ramalho, SP, Kersken, D, Martinez Arbizu, P, Colaco, A, 2020. Are seamounts refuge areas for fauna from polymetallic nodule fields? Biogeosciences 17(9): 2657-2680. https://doi.org/10.5194/bg-17-2657-2020" title="Are seamounts refuge areas for fauna from polymetallic nodule fields? Biogeosciences 17 (9): 2657 - 2680." url="https://doi.org/10.5194/bg-17-2657-2020" year="2020">Cuvelier et al. 2020</bibRefCitation>
), but not in abyssal areas of the Kiribati EEZ.
</paragraph>
<paragraph id="29D3C7A31A08B1445D2CB8481B250D0C" pageId="0" pageNumber="1">
<emphasis id="D519A9BA5B4C3F9BA50E11772EE6AB91" bold="true" pageId="0" pageNumber="1">
Order 
<taxonomicName id="251FEFEAEEDC7F17B8628C4EAA52F011" class="Anthozoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="order">Pennatulacea</taxonomicName>
Verrill, 1865
</emphasis>
</paragraph>
<paragraph id="2BE96B6E2EA8BCB20ABAA3C51ADA6AAD" pageId="0" pageNumber="1">
A total of 79 records of 
<taxonomicName id="1CAC8C890E3647CFF4EA3FF6C94C3D60" class="Anthozoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="order">Pennatulacea</taxonomicName>
occurring at&gt; 3000 m depth in the CCZ have been recorded in OBIS, but none represent preserved specimens (
<bibRefCitation id="EEED1A07BC4CAF8B70E035AAFCA1686E" author="O'Loughlin, PM" journalOrPublisher="Part II, Topical Studies in Oceanography" pageId="0" pageNumber="1" refId="B75" refString="OBIS (2022) Ocean Biodiversity Information System. Intergovernmental Oceanographic Commission of UNESCO." year="2011">OBIS 2022</bibRefCitation>
). We recovered a single specimen, for which sequences of both 16S and COI genes were obtained, and which were included in the phylogenetic analysis of the 
<taxonomicName id="CF73BA2A53CAC462204815A9FDABDA56" kingdom="Animalia" lsidName="" pageId="0" pageNumber="1" rank="subClass" subClass="Octocorallia">Octocorallia</taxonomicName>
(Fig. 
<figureCitation id="76053086584F4E1729CE326FA2928826" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rooted Bayesian phylogeny of Octocorallia. Concatenated (16 S, COI, mtMutS, NADH 2) median consensus BEAST tree with posterior probability (PP) and bootstrap (BS) values indicated. Only values of PP&gt; 0.70 and BS&gt; 50 are shown, with values of PP&gt; 0.95 and BS&gt; 90 indicated with a circle. Nodes not recovered on the RAxML tree are indicated with a hyphen. Sequences generated in this study are highlighted in violet." figureDoi="10.3897/zookeys.1113.82172.figure15" httpUri="https://binary.pensoft.net/fig/716341" pageId="0" pageNumber="1">15</figureCitation>
).
</paragraph>
<paragraph id="8F9B29179960A1F677AABD42DB5EB4D7" pageId="0" pageNumber="1">
<emphasis id="8C01E259E96EB2CB9C7691589CFE7E6B" bold="true" pageId="0" pageNumber="1">
Suborder 
<taxonomicName id="2A1910387A012703839D2F44EE5B4383" class="Anthozoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="subOrder" subOrder="Sessiliflorae">Sessiliflorae</taxonomicName>
<normalizedToken id="FC7182811C5C295AD779541986F26621" originalValue="Kükenthal">Kuekenthal</normalizedToken>
, 1915
</emphasis>
</paragraph>
<paragraph id="A79FF9EF54F899D29EC583B5E6FF6541" pageId="0" pageNumber="1">
<emphasis id="5BCEC631385C97F54CA030BE9A981C12" bold="true" pageId="0" pageNumber="1">
Family 
<taxonomicName id="C2245F1029B6B7D7D23C175F5FEFE5F1" authorityName="Kolliker" authorityYear="1872" class="Anthozoa" family="Protoptilidae" kingdom="Animalia" lsidName="" order="Pennatulacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="family">Protoptilidae</taxonomicName>
<normalizedToken id="3C87A3D5212BDEA5C99732BD18436E48" originalValue="Kölliker">Koelliker</normalizedToken>
, 1872
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>