<documentID-CLB-Dataset="3007"ID-DOI="http://dx.doi.org/10.3897/subtbiol.41.75613"ID-GBIF-Dataset="d409aeb2-e394-4143-a84d-77ce363a8b73"ID-Pensoft-Pub="1314-2615-41-69"ID-Pensoft-UUID="5992EA8732135F93917415967CC3DDE0"ID-ZooBank="917780D7509B43D7B6499D1D4FF0EC54"ModsDocID="1314-2615-41-69"checkinTime="1639196403999"checkinUser="pensoft"docAuthor="Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B. & Rađa 5, Tonci"docDate="2021"docId="075A8FD8A3B45743840D9C7DBB0D8B38"docLanguage="en"docName="SubterBiol 41: 69-85"docOrigin="Subterranean Biology 41"docPubDate="2021-12-10"docSource="http://dx.doi.org/10.3897/subtbiol.41.75613"docTitle="Leonhardia solaki Curcic, Rađa, Vesovic & Vrbica 2021, sp. nov."docType="treatment"docVersion="5"id="5992EA8732135F93917415967CC3DDE0"lastPageNumber="69"masterDocId="5992EA8732135F93917415967CC3DDE0"masterDocTitle="A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus"masterLastPageNumber="85"masterPageNumber="69"pageNumber="69"updateTime="1732866781713"updateUser="ExternalLinkService">
<mods:titleid="CD1D8D6258A36CAF2E05B8D9897D751D">A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus</mods:title>
<mods:affiliationid="2F54DE346D06D041B0A72E9DABB317DA">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:affiliationid="318619179AAC07898715EA28716CE304">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:affiliationid="C961889A39BAB8A8C0CF593532BBE02A">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:affiliationid="6116330A36E5C9612FAE209FE2403DCE">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:affiliationid="C4CC11D2FB1DD0F1A2F50B7CDBAA8149">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:affiliationid="B8BF1982414F8E5FA3ED1FD6983A9D0C">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:affiliationid="B5C1DC92180D45233E6DA3DA48C9716F">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<figureCitationid="220E07DAEA74C78C1C40A827D539C31E"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">http://zoobank.org/32557822-9245-4CE3-9E5B-3E8ABF5DEF12 Figures 2</figureCitation>
<figureCitationid="80C3722C0F98859B780694554B81A0E0"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">, 3</figureCitation>
<figureCitationid="95C9C36A8BB4A703EB4EEF39D9D895C6"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
(seven specimens). Same data as for holotype [two males and two females, IZFB, SSM]; same data as for holotype except for date [one male and two females, IZFB, June, 12, 2018]. All paratypes are labeled with white, printed locality labels and with red printed labels "Paratypus
<figureCitationid="1DCA72C730CC7CD414758B2F15628300"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Map of Bosnia and Herzegovina showing locality records for all taxa of Leonhardia. The grey pattern indicates karst terrain L. reitteri reitteri (blue circles), L. reitteri mersa (blue cross), L. reitteri zariquieyi (blue rhombuses), L. delminiumica (green squares), L. jajcensis (lime deltoid), L. hilfi hilfi (orange star), L. hilfi robusta (orange sun), L. sebesicensis (pink pentagon), L. droveniki (red hexagon), L. solaki sp. nov. (yellow triangle). Scale bar: 50 km."figureDoi="10.3897/subtbiol.41.75613.figure1"httpUri="https://binary.pensoft.net/fig/621562"pageId="0"pageNumber="69">1</figureCitation>
<figureCitationid="FF7A8ADCC7D45063C673E8D13A1258A9"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<bibRefCitationid="8823519D67521EFB1714F353ECA38CC9"author="Jeannel, R"journalOrPublisher="Archives de Zoologie experimentale et generale"pageId="0"pageNumber="69"pagination="1 - 436"refId="B14"refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436"title="Monographie des Bathysciinae. Biospeologica L."volume="63"year="1924">Jeannel 1924</bibRefCitation>
<bibRefCitationid="A687B88267CAC31609D5F00FBD3E4554"author="Nonveiller, G"journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie"pageId="0"pageNumber="69"pagination="165 - 168"refId="B19"refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168"title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]."volume="24"year="2002">Nonveiller et al. 2002</bibRefCitation>
<bibRefCitationid="2BC33D77568263D7B104B3FBEB9E8E44"DOI="https://doi.org/10.2298/ABS1402919C"author="Curcic, S"journalOrPublisher="Archives of Biological Sciences, Belgrade"pageId="0"pageNumber="69"pagination="919 - 933"refId="B6"refString="Curcic, S, Rađa, T, Mulaomerovic, J, Vrbica, M, Antic, D, Curcic, B, Rađa, B, Vesovic, N, 2014. Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina. Archives of Biological Sciences, Belgrade 66: 919 - 933, DOI: https://doi.org/10.2298/ABS1402919C"title="Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina."url="https://doi.org/10.2298/ABS1402919C"volume="66"year="2014">
<bibRefCitationid="0F3F70D744AE7766628C7A00A0216AE6"DOI="https://doi.org/10.11646/zootaxa.4483.3.6"author="Curcic, S"journalOrPublisher="Zootaxa"pageId="0"pageNumber="69"pagination="523 - 548"refId="B7"refString="Curcic, S, Pavicevic, D, Vesovic, N, Mulaomerovic, J, Rađa, T, Antic, D, Bosco, F, Markovic, Đ, Petkovic, M, 2018. Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula. Zootaxa 4483: 523 - 548, DOI: https://doi.org/10.11646/zootaxa.4483.3.6"title="Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula."url="https://doi.org/10.11646/zootaxa.4483.3.6"volume="4483"year="2018">2018</bibRefCitation>
in regard to AL in males (antennae exceeding the middle of elytra vs. antennae reaching the middle of elytra); A10L/A10W (less than 2.375 vs. more than 2.50); pronotum form (bell-shaped, lateral margins well-rounded anteriorly vs. subquadrate, lateral margins obtuse anteriorly); PL/PW (pronotum wider than long vs. pronotum as long as wide); shape of the mesosternal carina (higher, almost right-angled, posterior margin concave vs. lower, obtuse-angled, posterior margin somewhat elevated); shape of elytra (inversely ovate vs. widely oval); shape of the median lobe in dorsal view (apically flattened vs. apically narrowed); length of the median lobe (barely longer than parameres vs. markedly longer than parameres); shape and size of the basal bulb in dorsal view (larger, rounded vs. smaller, not rounded); and shape of the aedeagus in lateral view (median lobe more curved, basal bulb larger vs. median lobe less curved, basal bulb smaller) (Figs
<figureCitationid="EA61A0470B4E8717BC545A07C0EE94CA"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
<figureCitationid="3CCD8B8DAE1928A8CA35ECD62D4F95B0"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<bibRefCitationid="B586C48CBE4231A96937774277254963"DOI="https://doi.org/10.5962/bhl.part.7311"author="Reitter, E"journalOrPublisher="Berichte des Naturwissenschaftlich-medizinischen Vereins in Innsbruck"pageId="0"pageNumber="69"refId="B28"refString="Reitter, E, 1901. Ein neuer blinder Grotten-Silphide aus der Herzegowina. Wiener entomologische Zeitung 20: 128. https://doi.org/10.5962/bhl.part.7311"title="Ein neuer blinder Grotten-Silphide aus der Herzegowina. Wiener entomologische Zeitung 20: 128."url="https://doi.org/10.5962/bhl.part.7311"year="1901">Reitter 1901</bibRefCitation>
<bibRefCitationid="1F1074ED9044CBC9BEDBCD1323884F6C"author="Jeannel, R"journalOrPublisher="Archives de Zoologie experimentale et generale"pageId="0"pageNumber="69"pagination="1 - 436"refId="B14"refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436"title="Monographie des Bathysciinae. Biospeologica L."volume="63"year="1924">Jeannel 1924</bibRefCitation>
<bibRefCitationid="BB772E77FBE6FD1D0678303230F0D24E"DOI="https://doi.org/10.3406/bsef.1999.16602"author="Perreau, M"journalOrPublisher="Bulletin de la Societe entomologique de France"pageId="0"pageNumber="69"pagination="399 - 406"refId="B20"refString="Perreau, M, 1999. Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae). Bulletin de la Societe entomologique de France 104: 399 - 406, DOI: https://doi.org/10.3406/bsef.1999.16602"title="Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae)."url="https://doi.org/10.3406/bsef.1999.16602"volume="104"year="1999">Perreau 1999</bibRefCitation>
<bibRefCitationid="E9CC4DD64D63AC61C9FAC347FA1ABF92"author="Nonveiller, G"journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie"pageId="0"pageNumber="69"pagination="165 - 168"refId="B19"refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168"title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]."volume="24"year="2002">Nonveiller et al. 2002</bibRefCitation>
, from which it differs with respect to AL (antennae exceeding the middle of elytra in males and reaching the middle of elytra in females vs. antennae barely reaching the middle of body); A2L/A3L+A4L (antennomere II shorter than the following two antennomeres combined vs. antennomere II as long as the following two antennomeres combined); A8L/A8W (M in males 2.06, in females 1.58 vs. 1.50 in both genders); A10L/A10W (R in females 1.54-1.78 vs. more than 2.00 in both genders); shape of the hind pronotal angles (obtuse vs. right or weakly acute); shape of the mesosternal carina (almost right-angled, anterior margin convex, posterior margin regularly concave vs. obtuse-angled, anterior margin obtuse, posterior margin deeply incised); shape of elytra (inversely ovate vs. oval); shape of the median lobe in dorsal (wider in apical half, apically flattened vs. thinner in apical half, apically narrowed) and lateral (more elongate vs. less elongate) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); and position of parameral setae in dorsal view (inner pre-apical seta somewhat separated from two remaining setae vs. all setae equidistant) (Figs
<figureCitationid="6073A6342473FD130034AF74C70063E0"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
<figureCitationid="801C4A3AEAEDF7625C05E29DE9B4E711"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<bibRefCitationid="514F2479469C575A13250A16CD2AF9DF"author="Breit, J"journalOrPublisher="Cladistics"pageId="0"pageNumber="69"refId="B2"refString="Breit, J, 1902. Ein neuer Hoehlenkaefer aus Bosnien. Societas entomologica 17: 89."title="Ein neuer Hoehlenkaefer aus Bosnien. Societas entomologica 17: 89."year="1902">Breit 1902</bibRefCitation>
<bibRefCitationid="D813A72E1057D4B2D05F339CE9251575"author="Jeannel, R"journalOrPublisher="Archives de Zoologie experimentale et generale"pageId="0"pageNumber="69"pagination="1 - 436"refId="B14"refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436"title="Monographie des Bathysciinae. Biospeologica L."volume="63"year="1924">Jeannel 1924</bibRefCitation>
<bibRefCitationid="3F5B38B4C049D234B30E84D067907A33"author="Mueller, G"journalOrPublisher="Atti del Museo Civico di Storia Naturale di Trieste"pageId="0"pageNumber="69"pagination="107 - 117"refId="B16"refString="Mueller, G, 1937. Nuovi silfidi cavernicoli della Balcania e osservazioni su specie gia descritte. Atti del Museo Civico di Storia Naturale di Trieste 13: 107 - 117"title="Nuovi silfidi cavernicoli della Balcania e osservazioni su specie gia descritte."volume="13"year="1937">
<bibRefCitationid="3A0C2CD5B3E758A65A2C952B67E7AAAD"DOI="https://doi.org/10.3406/bsef.1999.16602"author="Perreau, M"journalOrPublisher="Bulletin de la Societe entomologique de France"pageId="0"pageNumber="69"pagination="399 - 406"refId="B20"refString="Perreau, M, 1999. Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae). Bulletin de la Societe entomologique de France 104: 399 - 406, DOI: https://doi.org/10.3406/bsef.1999.16602"title="Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae)."url="https://doi.org/10.3406/bsef.1999.16602"volume="104"year="1999">Perreau 1999</bibRefCitation>
<bibRefCitationid="DB81A5BD7CCD0C0E30BCC251FA4CA94D"author="Nonveiller, G"journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie"pageId="0"pageNumber="69"pagination="165 - 168"refId="B19"refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168"title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]."volume="24"year="2002">Nonveiller et al. 2002</bibRefCitation>
in regard to TL (R 3.40-3.65 mm in males, 3.55-3.93 mm in females vs. R 3.10-3.20 mm in males, 3.40 mm in a female); A8L/A8W (M 2.06 in males, 1.58 in females vs. antennomere VIII barely longer than broad); shape of the mesosternal carina (almost right-angled, posterior margin less concave vs. obtuse-angled, posterior margin more concave); shape of the median lobe in dorsal (apically flattened, with a rounded apex vs. apically narrowed, with a pointed apex) and lateral (more curved, wider vs. less curved, narrower) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); size of the basal bulb in dorsal view (larger vs. smaller); and position of parameral setae in dorsal view (inner pre-apical seta closer to the two remaining setae vs. inner pre-apical seta farther away from the two remaining setae) (Figs
<figureCitationid="2D9DFF8E55353D1ADC559FFBED91DFF8"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
<figureCitationid="7E2FD288E12CC857F902F0FC5302A2F1"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<bibRefCitationid="A8002ED3517A4D05BF43E8BECC0633DB"author="Nonveiller, G"journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie"pageId="0"pageNumber="69"pagination="165 - 168"refId="B19"refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168"title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]."volume="24"year="2002">Nonveiller et al. 2002</bibRefCitation>
, from which it differs with respect to AL in females (antennae reaching the middle of elytra vs. antennae ending before the middle of elytra); shape of the hind pronotal angles (obtuse vs. almost right); shape of the mesosternal carina (higher, anterior margin more convex, posterior margin more concave vs. lower, anterior margin less convex, posterior margin less concave); form of the elytra (more elongate vs. less elongate); shape of the median lobe in dorsal (apically flattened vs. apically narrowed) and lateral (more thickened in its basal half, less convex ventrally vs. less thickened in its basal half, more convex ventrally) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); size of the basal bulb in dorsal view (larger vs. smaller); and position of parameral setae in dorsal view (inner pre-apical seta farther away from the two remaining setae vs. inner pre-apical seta closer to the two remaining setae) (Figs
<figureCitationid="435310AB61BA864106E6CDB2782B764E"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
<figureCitationid="1B24E6655A79C082A6054ACAFC8DE5A6"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<bibRefCitationid="7EE90AA01CD074FF0F760A2B012DDE96"DOI="https://doi.org/10.2298/ABS1402919C"author="Curcic, S"journalOrPublisher="Archives of Biological Sciences, Belgrade"pageId="0"pageNumber="69"pagination="919 - 933"refId="B6"refString="Curcic, S, Rađa, T, Mulaomerovic, J, Vrbica, M, Antic, D, Curcic, B, Rađa, B, Vesovic, N, 2014. Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina. Archives of Biological Sciences, Belgrade 66: 919 - 933, DOI: https://doi.org/10.2298/ABS1402919C"title="Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina."url="https://doi.org/10.2298/ABS1402919C"volume="66"year="2014">
in regard to AL in females (antennae reaching the middle of elytra vs. antennae ending before the middle of elytra); shape of the hind pronotal angles (obtuse-angled in both genders vs. almost right-angled in males); shape of the pronotum and elytra (less elongate vs. more elongate); shape of the mesosternal carina (apically rounded vs. apically toothed); shape of the median lobe in dorsal (apically flattened vs. apically rounded) and lateral (more curved, narrower in apical half vs. less curved, wider in apical half) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); and position of parameral setae in dorsal view (inner pre-apical seta somewhat separated from two remaining setae vs. all setae equidistant) (Figs
<figureCitationid="51ED045F6F5716BE388DFE07BD8FF169"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
<figureCitationid="EAEBBE4BD1D0804B3A168D9A7D514148"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<bibRefCitationid="CDE29F76D94DA90CB2B71A199E6558E8"DOI="https://doi.org/10.11646/zootaxa.4483.3.6"author="Curcic, S"journalOrPublisher="Zootaxa"pageId="0"pageNumber="69"pagination="523 - 548"refId="B7"refString="Curcic, S, Pavicevic, D, Vesovic, N, Mulaomerovic, J, Rađa, T, Antic, D, Bosco, F, Markovic, Đ, Petkovic, M, 2018. Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula. Zootaxa 4483: 523 - 548, DOI: https://doi.org/10.11646/zootaxa.4483.3.6"title="Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula."url="https://doi.org/10.11646/zootaxa.4483.3.6"volume="4483"year="2018">
<paragraphid="48AFE70289B4943446C9249AF32FAFDC"pageId="0"pageNumber="69">Medium-sized leptodirine. TL M 3.62 mm (3.55 mm in males, 3.69 mm in females), R 3.40-3.93 mm (3.40-3.65 mm in males, 3.55-3.93 mm in females).</paragraph>
<figureCitationid="3189F081F2227C375FD14BDA9B0ADAE2"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="0357287151333A28EA6C38ACCB8CBAD4"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3A, B</figureCitation>
<figureCitationid="6363A30EE29E09A774A6FCDF863AF19B"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3D, G, L</figureCitation>
). Body covered with a number of densely distributed deep punctures and yellow pubescence of medium length (erect on head, recumbent on both pronotum and elytra).
<figureCitationid="7E2CED2873BA0657049BFDDA3B9A4AD8"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="D57D86E11927E1BAE84045E525BA7117"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3C</figureCitation>
). Labrum slightly emarginate, with several long setae. Penultimate maxillary palpomere widened apically. Ultimate maxillary palpomere short, thin, gradually narrowing apically. Vertex with a longitudinal impression. Antennae inserted in middle third of head, slender, narrow proximally, slightly dilated distally, longer in males, AL M 2.15 mm, R 1.93-2.33 mm (2.165-2.33 mm in males, 1.93-2.165 mm in females), exceeding middle of elytra in males and reaching middle of elytra in females (Figs
<figureCitationid="BCB2831D9995176B92B44C1B31607287"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="106452000B011F0D7EB60FF85BEC7EFA"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3E</figureCitation>
). Antennomere I short and wide. A1L/A2L M 0.54, R 0.52-0.58. Antennomere II narrow, elongate, shorter than III and IV combined (A2L/A3+A4L M 0.86, R 0.79-0.90). Antennomeres III-VI of similar shape and size, narrow, among which V is the longest. Antennomeres VII and IX-XI widened (VII, IX, and X dilated distally), large, among which VII is the shortest in males, VIII the shortest in females, and XI the longest in both sexes. Antennomere VIII small, elongate in males (A8L/A8W M 2.06, R 1.82-2.40), wide in females (A8L/A8W M 1.58, R 1.33-1.90). A9L/A9W in males M 2.35 (R 2.25-2.50), in females M 1.68 (R 1.60-1.78). A10L/A10W in males M 2.27 (R 2.21-2.375), in females M 1.65 (R 1.54-1.78). Ultimate antennomere slender, narrowing apically, A11L/A11W in males M 3.10 (R 2.54-3.50), in females M 2.82 (R 2.67-3.00).
<bibRefCitationid="E1D7E078318359AF279EE9930C3E42A1"author="Nonveiller, G"journalOrPublisher="Revue francaise d'Entomologie, Nouvelle serie"pageId="0"pageNumber="69"pagination="165 - 168"refId="B19"refString="Nonveiller, G, Pavicevic, D, Rada, T, Vujcic-Karlo, S, 2002. A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]. Revue francaise d'Entomologie, Nouvelle serie 24: 165 - 168"title="A new species of the genus Leonhardia Reitter, 1901 from Herzegovina [Coleoptera, Leiodidae, Cholevinae, Leptodirini]."volume="24"year="2002">Nonveiller et al. (2002)</bibRefCitation>
<bibRefCitationid="7C35459E194A553C69E84AA997DAC1FE"DOI="https://doi.org/10.2298/ABS1402919C"author="Curcic, S"journalOrPublisher="Archives of Biological Sciences, Belgrade"pageId="0"pageNumber="69"pagination="919 - 933"refId="B6"refString="Curcic, S, Rađa, T, Mulaomerovic, J, Vrbica, M, Antic, D, Curcic, B, Rađa, B, Vesovic, N, 2014. Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina. Archives of Biological Sciences, Belgrade 66: 919 - 933, DOI: https://doi.org/10.2298/ABS1402919C"title="Three new cave-dwelling leiodid beetles (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina."url="https://doi.org/10.2298/ABS1402919C"volume="66"year="2014">
<bibRefCitationid="A720E153EE29B06236E061F2D1F8BF08"DOI="https://doi.org/10.11646/zootaxa.4483.3.6"author="Curcic, S"journalOrPublisher="Zootaxa"pageId="0"pageNumber="69"pagination="523 - 548"refId="B7"refString="Curcic, S, Pavicevic, D, Vesovic, N, Mulaomerovic, J, Rađa, T, Antic, D, Bosco, F, Markovic, Đ, Petkovic, M, 2018. Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula. Zootaxa 4483: 523 - 548, DOI: https://doi.org/10.11646/zootaxa.4483.3.6"title="Seven new taxa of Leptodirini (Coleoptera: Leiodidae: Cholevinae) from the Balkan Peninsula."url="https://doi.org/10.11646/zootaxa.4483.3.6"volume="4483"year="2018">2018</bibRefCitation>
<figureCitationid="9925F5774BB8C25D580E479D95E8C7BA"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="F28AFCACB8E0C4FE675D9C68F3B6C4BD"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3F</figureCitation>
<figureCitationid="C91B4CDBE5304F7C9F2206B5762B8547"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3B</figureCitation>
<figureCitationid="B42948E075F767908411D72B18D8E06A"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2B</figureCitation>
<figureCitationid="06FA5B320E39E52FF2E54CC00CE32E14"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3H</figureCitation>
<figureCitationid="210C39D221555865E3373333842A46C1"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3I</figureCitation>
<figureCitationid="602005BD090795A8D46FD3CD22642C72"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="ECB4191672DBE140B2BEC4275EDDB982"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3J</figureCitation>
<figureCitationid="97F4233391BA4C048FA894C4DAC53E22"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="99BF4AA4C819AC1170977D2CBF4ABF46"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3K</figureCitation>
<figureCitationid="3D6D0F9497FF406E67273692B230FB6B"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3B</figureCitation>
<figureCitationid="24A3961BCE7D2A61D6BCBAFD8842BE2B"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
). Femora widened basally, constricted sub-apically. Tibiae thin, very gently curved, gradually widening distally, each with several spines. Each fore tibia with an outer distal brush-like structure. Male fore tarsi five-segmented, weakly dilated. Tarsal claws elongate, narrow, curved, pointed apically.
<figureCitationid="0F33DD69B72E472AE2C17B31ED9375D2"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2C, D</figureCitation>
<figureCitationid="6E9AA172F8C4A0CB2D91F23E089383D6"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2C</figureCitation>
<figureCitationid="F1A4884C8E3162CE91DDFD360ADDC233"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2D</figureCitation>
<figureCitationid="A8537BE6683EDC02F6FADFA918A39940"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2C</figureCitation>
). Median lobe in lateral view curved, straight in basal half, sub-apically concave dorsally, almost straight ventrally in basal part, with an acute apex curved downwards (Fig.
<figureCitationid="AA1D6D18CB49208BA8DDC1803AD14296"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2D</figureCitation>
<figureCitationid="EDB401498291D283148A42BE3ED95845"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2C, E</figureCitation>
<figureCitationid="E818721B4C774A795ED60B5D5199D1F6"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2D</figureCitation>
). Paramere bearing three setae: two strong, long, in apical and pre-apical position, respectively, and one inner, thin, short, in pre-apical position (Fig.
<figureCitationid="AF8FAE8EE502EFBC981DC23D7DC41484"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2E</figureCitation>
<figureCitationid="AD379E2CAC3BE39241CD7B5F1287FFD4"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2E</figureCitation>
). Endophallus elongate, tubular. Copulatory piece having a Y-shaped phanera and two lateral basal sclerifications, in front of which a weakly chitinized structure resembling teeth is present. In addition, two lateral bands are present in apical half of median lobe (Fig.
<figureCitationid="BE2E201A46349DCFD4EBBAED1B54D5F1"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2C</figureCitation>
<figureCitationid="38AD7D91D855FDA18E4ED883AE6C0A84"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2F</figureCitation>
<figureCitationid="389D9901DD8F3A579B2134D92221D3EF"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2F</figureCitation>
<figureCitationid="16B19A2AD32B4FDF621FF554E15BC5C5"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2G</figureCitation>
<figureCitationid="26AC84FDFF4C7921A2A1B670F039ABB6"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2H</figureCitation>
<paragraphid="C44EF5B599CAB9F2FEBFEE1D9D0B0210"pageId="0"pageNumber="69">A certain level of intraspecific variability is observed in the new species. Several morphological differences between males and females point to the occurrence of sexual dimorphism. To be specific: (i) males are on average slightly shorter than females; (ii) the antennae in males are longer than in females; (iii) antennomeres VIII-XI in males are more slender than in females; (iv) the head in males is larger than in females; (v) the pronotum in males is more elongate than in females; (vi) the elytra in males are more elongate than in females.</paragraph>
species. We assume that the new species probably inhabits other subterranean sites in the surrounding areas of western Bosnia and Herzegovina, although one of us (TR) visited the
kod Skucana Pit. They were recorded at this subterranean site at a depth of 30 m under conditions of permanent darkness and high humidity, along with the presence of trickling water. All individuals of
, 1943, which is recorded in the same habitat where individuals of the new species were found, so these two taxa can be treated as sympatric. The same pit is also the type locality of the recently described moth fly
<taxonomicNameid="0B94E2A332D8477FBD816A6AD9BA3396"authorityName="Wagner & Rada"authorityYear="2020"class="Insecta"family="Psychodidae"genus="Psychoda"kingdom="Animalia"lsidName="Psychoda glamocensis"order="Diptera"pageId="0"pageNumber="69"phylum="Arthropoda"rank="species"species="glamocensis">
<bibRefCitationid="D8C80F4238F4C2A1F16489CC450E1471"DOI="https://doi.org/10.11646/zootaxa.4845.2.8"author="Wagner, R"journalOrPublisher="Zootaxa"pageId="0"pageNumber="69"pagination="275 - 282"refId="B32"refString="Wagner, R, Rada, T, 2020. Moth flies (Diptera, Psychodidae) living in the dark of caves in the Dinaric Karst. Zootaxa 4845: 275 - 282, DOI: https://doi.org/10.11646/zootaxa.4845.2.8"title="Moth flies (Diptera, Psychodidae) living in the dark of caves in the Dinaric Karst."url="https://doi.org/10.11646/zootaxa.4845.2.8"volume="4845"year="2020">Wagner and Rada 2020</bibRefCitation>
<subSubSectionid="3C182C3BDCDBFC2FE5BE54D63F51116A"pageId="0"pageNumber="69"type="key for identification of species and subspecies of leonhardia (figs 2–4)">
<figureCitationid="F0FF9467C4EEF4AEF612386C058BE70C"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2</figureCitation>
<figureCitationid="202412CCD5DEF1E99658F6C90D06B9F3"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4</figureCitation>
<figureCitationid="8FEF6C607C85B1BA5C5B5806A1186E82"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4A</figureCitation>
<figureCitationid="A20097B426CFA143ACBA427263C02B73"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2B</figureCitation>
<figureCitationid="2CC2D961255796577D03CC3434AA45F8"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3H</figureCitation>
<figureCitationid="3D333FA93F8A37FE0438DD223AF40C38"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4B-F</figureCitation>
<figureCitationid="41B553F11E23F8ED7002F3AFA9D6B828"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4C</figureCitation>
<figureCitationid="ACBF1276011F87239FBD942E08D6AFCD"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4M</figureCitation>
<figureCitationid="E67E63D96750E38041AF186446A95A79"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4B, D-F</figureCitation>
<figureCitationid="5DBA04C03B97671BB248752D6F891A74"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4L, N-P</figureCitation>
<tdid="0D3A589E19832094A09304BF6A90545E"colspan="1"pageId="0"pageNumber="69"rowspan="1">Larger body size (3.50-3.70 mm in length). Pronotum less transverse. Lateral elytral margins more arcuate, elytra apically rounded</td>
<tdid="38B9DBE59E61E0A6D4E639EDB2690FC1"colspan="1"pageId="0"pageNumber="69"rowspan="1">Smaller body size (3.20 mm in length). Pronotum more transverse. Lateral elytral margins less arcuate, elytra apically pointed</td>
<tdid="03DF988F8B90BDAB4B373912DC24205D"colspan="1"pageId="0"pageNumber="69"rowspan="1">Pronotum narrowed basally, elytra narrower and less convex, pronotal depressions lacking or barely discernible</td>
<tdid="A0DD5851B4B2D884BD6E749A2D2F5539"colspan="1"pageId="0"pageNumber="69"rowspan="1">Pronotum not narrowed basally, elytra wider and more convex, two depressions present on pronotum</td>
<figureCitationid="B96ACC8098F3C919512F049912233945"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="E7998239BBE26FB877ADCC126BB02513"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3H</figureCitation>
<figureCitationid="066C389FB906EE23BF1374E491EAAF67"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4E</figureCitation>
<figureCitationid="8D3029E0C287091237549656A3CB2B90"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4B, D, F</figureCitation>
<figureCitationid="E7274B61EE43CE1AA6C24280593C06F0"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4E</figureCitation>
<figureCitationid="736C2B401F34C8C49091AC0930C23A8C"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4J</figureCitation>
<figureCitationid="6EDFBB8949E615294F055A4F71599553"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2A</figureCitation>
<figureCitationid="25D29706BCFE5A398D050F1EDBBF84BF"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3F</figureCitation>
<figureCitationid="4905B0B0E3D839710C0D9B5E3A5B05E1"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2B</figureCitation>
<figureCitationid="A7A63A89FD69DA32D8703258E16F833A"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. SEM images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A paratype male, habitus, dorsal aspect B paratype male, habitus, lateral aspect C paratype male, head, dorsal aspect D paratype male, microsculpture of head, dorsal aspect E paratype male, right antenna, dorsal aspect F paratype male, pronotum, dorsal aspect G paratype male, microsculpture of pronotum, dorsal aspect H paratype male, mesosternal carina, lateral aspect I paratype male, mesoventrite, ventral aspect J paratype male, scutellum, dorsal aspect K paratype male, elytra, dorsal aspect L paratype male, microsculpture of elytra, dorsal aspect. Scale bars: 1.0 mm (A, B); 0.5 mm (E, F, H, I, K); 0.2 mm (C, J); 0.1 mm (D, G, L)."figureDoi="10.3897/subtbiol.41.75613.figure3"httpUri="https://binary.pensoft.net/fig/621564"pageId="0"pageNumber="69">3H</figureCitation>
<figureCitationid="5CC25CD764D14222B43F94B3782275AC"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Bright-field images of morphological features of Leonhardia solaki sp. nov. from the Golubnjaca kod Skucana Pit, village of Skucani, close to the town of Glamoc, western Bosnia and Herzegovina A holotype male, habitus, dorsal aspect B holotype male, mesosternal carina, lateral aspect C holotype male, aedeagus, dorsal aspect D holotype male, aedeagus, lateral aspect E holotype male, left paramere apex, dorsal aspect F paratype female, left gonostylus, dorsal aspect G paratype female, spermatheca, lateral aspect H paratype female, abdominal segment VIII, ventral aspect. Scale bars: 1.0 mm (A); 0.2 mm (B-D, H); 0.1 mm (F, G); 0.05 mm (E)."figureDoi="10.3897/subtbiol.41.75613.figure2"httpUri="https://binary.pensoft.net/fig/621563"pageId="0"pageNumber="69">2C</figureCitation>
<figureCitationid="921825B75A63023FEFEDFDDB99F267CF"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4I</figureCitation>
<figureCitationid="6327CF5D4760079F584BB05A0434602B"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4S</figureCitation>
<figureCitationid="13355B02C8E2BAFECE7DE07F2D098B44"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4G, K</figureCitation>
<figureCitationid="79BE9A605DB0D9FE248EDC7E98A39305"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4Q, U</figureCitation>
<figureCitationid="6486E2C498D501B0FB8BEA055545262E"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4F</figureCitation>
<figureCitationid="84D42D1F633FE1668ACC645EEB6B4626"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4K</figureCitation>
<figureCitationid="DB51BC9A28878F8C1C84076E561F1146"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4U</figureCitation>
<figureCitationid="5BE98F95E74990F4D63BC8A2F62BB7EE"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4B</figureCitation>
<figureCitationid="A88F01FD7AFE98AEDA89EC900770D890"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4G</figureCitation>
<figureCitationid="038DDC80FEC13D125E7CA9947A28B393"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Different morphological features of Leonhardia [L. droveniki (A) L. hilfi (B, G, L, Q) L. reitteri (C, H, M, R) L. delminiumica (D, I, N, S) L. jajcensis (E, J, O, T) and L. sebesicensis (F, K, P, U)] [modified after Nonveiller et al. (2002) and Curcic et al. (2014, 2018)] A-F mesosternal carina, lateral aspect G-K aedeagus, dorsal aspect L-P aedeagus, lateral aspect Q-U left paramere apex, dorsal aspect. Scale bars: 0.5 mm (G-P)."figureDoi="10.3897/subtbiol.41.75613.figure4"httpUri="https://binary.pensoft.net/fig/621565"pageId="0"pageNumber="69">4Q</figureCitation>
<tdid="B6B28EEE3317F4FAC806C7534A8C19EB"colspan="1"pageId="0"pageNumber="69"rowspan="1">Smaller body size (3.40-3.50 mm in length). Antennae shorter and narrower. Punctures smaller. Pubescence shorter</td>
<tdid="7B6323F9141D73CF62F8CB6441574EC7"colspan="1"pageId="0"pageNumber="69"rowspan="1">Larger body size (3.80-4.00 mm in length). Antennae longer and wider. Punctures larger. Pubescence longer</td>
