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243 lines
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.3.920" ID-GBIF-Dataset="2b0a82dd-c694-440c-974f-f600476d450e" ID-PMC="PMC3174433" ID-Pensoft-Pub="1314-2003-3-1" ID-Pensoft-UUID="9E71FFF5FFF2CB01FFE7FFA3AF0C5917" ID-PubMed="22171174" ID-Zenodo-Dep="576069" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1314-2003-3-1" ModsDocOrigin="PhytoKeys 3" ModsDocTitle="Hieracium maccoshiana, a new Scottish hawkweed related to H. dovrense (Section Alpestria, Asteraceae)" checkinTime="1451252131426" checkinUser="pensoft" docAuthor="Rich, Timothy C. G." docDate="2011" docId="FEB99493EA5A8D65FFA35805FA30D44C" docLanguage="en" docName="PhytoKeys 3: 1-8" docOrigin="PhytoKeys 3" docSource="http://dx.doi.org/10.3897/phytokeys.3.920" docTitle="Hieracium maccoshiana T. C. G. Rich 2011, sp. nov." docType="treatment" docVersion="5" id="9E71FFF5FFF2CB01FFE7FFA3AF0C5917" lastPageNumber="4" masterDocId="9E71FFF5FFF2CB01FFE7FFA3AF0C5917" masterDocTitle="Hieracium maccoshiana, a new Scottish hawkweed related to H. dovrense (Section Alpestria, Asteraceae)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="2" updateTime="1668140988382" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Hieracium maccoshiana, a new Scottish hawkweed related to H. dovrense (Section Alpestria, Asteraceae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rich, Timothy C. G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>8</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.3.920</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.3.920</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-3-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">9E71FFF5FFF2CB01FFE7FFA3AF0C5917</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576069</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152024896" LSID="urn:lsid:plazi:treatment:FEB99493EA5A8D65FFA35805FA30D44C" httpUri="http://treatment.plazi.org/id/FEB99493EA5A8D65FFA35805FA30D44C" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection pageId="1" pageNumber="2" type="nomenclature">
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<paragraph pageId="1" pageNumber="2">
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<taxonomicName LSID="45394E62-BF79-5D27-8325-271613C0AA10" authority="T. C. G. Rich" authorityName="T. C. G. Rich" authorityYear="2011" class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hieracium maccoshiana" order="Campanulales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="maccoshiana" status="sp. nov.">Hieracium maccoshiana T.C.G.Rich</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="2">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Hieracium maccoshiana (NMW)." httpUri="https://binary.pensoft.net/fig/9871" pageId="1" pageNumber="2">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Middle stem leaves. A-C possible Hieracium maccoshiana. A-B Ben Loyal (NMW 28.131.5185). C Ben Loyal (BM). D-N Hieracium maccoshiana. D-F Rhiconich gorge (NMW V. 2005.1.159). G-H Foinaven (CGE). I-L Rhiconich (CGE). M-N Craig na H'Uidhe (CGE). O-U Hieracium dovrense, Drivstua, Dovrefj, Norway (S). O-Q S 09 - 16213. R S 09 - 16217. S S 09 - 16224. T-U S 09 - 16222. Scale bar 5 cm." httpUri="https://binary.pensoft.net/fig/9872" pageId="1" pageNumber="2">
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2
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<normalizedToken originalValue="D–N">D-N</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="latin">
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<paragraph pageId="1" pageNumber="2">Latin</paragraph>
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<paragraph pageId="1" pageNumber="2">
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<emphasis italics="true" pageId="1" pageNumber="2">Rosula basalia sub anthesi fere absenti. Foliis caulinis 4-8 ellipticis dentibus magnis. Pedunculis pilis multis stellatis pilis eglandulosis simplicibus multis pilisque glandulosis paucis. Bracteis involucralibus 2-2.5 mm latitudinis pilis stellatis paucis pilis eglandulosis simplicibus multis pilisque glandulosis multis. Ligulis apice glabris. Stylis obscuris.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="holotype">
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<paragraph pageId="1" pageNumber="2">Holotype:</paragraph>
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<paragraph pageId="1" pageNumber="2">
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<emphasis bold="true" pageId="1" pageNumber="2">Scotland</emphasis>
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. Sutherland: by small stream, Allt na Cuile, Rhiconich, 58.4470, -4.9300, 150 m alt., 26 July 2010, M. Jannink (
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<emphasis bold="true" pageId="1" pageNumber="2">NMW</emphasis>
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, accession number V.2010.1.213).
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/9871" pageId="1" pageNumber="2" start="Figure 1" startId="F1">
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<paragraph pageId="1" pageNumber="2">
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<emphasis bold="true" pageId="1" pageNumber="2">Figure 1.</emphasis>
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Holotype of
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<taxonomicName authorityName="T. C. G. Rich" authorityYear="2011" class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="maccoshiana">
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<emphasis italics="true" pageId="1" pageNumber="2">Hieracium maccoshiana</emphasis>
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</taxonomicName>
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(NMW).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
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<paragraph pageId="1" pageNumber="2">Description.</paragraph>
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<paragraph lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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Stem to 50 cm, pale green, slightly
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<normalizedToken originalValue="purple–tinted">purple-tinted</normalizedToken>
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below, slender to fairly robust; stellate hairs sparse and mainly above; simple eglandular hairs long, pale, sparse and glabrescent to dense (especially below); glandular hairs very small, occasional; sparsely and minutely puberulous. Basal leaves absent or withering before or up to flowering time or rarely persistent, few. Lamina elliptical or subrotund; apex
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<normalizedToken originalValue="obtuse–mucronulate">obtuse-mucronulate</normalizedToken>
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; base attenuate; margins remotely denticulate to coarsely dentate; with a few, pale, medium simple eglandular hairs above and below or nearly glabrous above. Petioles winged and broadened at base, with numerous medium to long, pale simple eglandular hairs. Stem leaves and inflorescence bracts (3
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<normalizedToken originalValue="–)4–">-)4-</normalizedToken>
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8, 2-10
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<normalizedToken originalValue="×">x</normalizedToken>
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0.8-3.5 cm, gradually decreasing in size upwards, pale green on upper surface, paler beneath. Lamina of the lower leaves elliptical; apex
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<normalizedToken originalValue="obtuse–mucronulate">obtuse-mucronulate</normalizedToken>
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to acute; base attenuate; margins denticulate to sharply and irregularly dentate with ascending teeth; petiole winged, semiamplexicaul. Lamina of the median leaves elliptical; apex
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<normalizedToken originalValue="obtuse–mucronulate">obtuse-mucronulate</normalizedToken>
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to acute; base rounded or abruptly contracted, sessile, semiamplexicaul; margins denticulate to sharply and irregularly dentate with large or small, ascending, mammiform teeth. Lamina of the upper leaves lanceolate; apex acute to acuminate; base cuneate, sessile, semiamplexicaul, margins denticulate to shallowly dentate. All stem leaves with stellate hairs few to sparse on both surfaces; simple eglandular hairs few to numerous below and nearly glabrous above, pale, medium. Inflorescence usually with 2
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<normalizedToken originalValue="–9(–">-9(-</normalizedToken>
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12) capitula, rather compactly cymose. Peduncles 1-5 cm (acladium 0.3-2 cm), suberect, slender; stellate hairs sparse to dense; simple eglandular hairs few to numerous, short to medium,
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<normalizedToken originalValue="dark–based">dark-based</normalizedToken>
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; glandular hairs few, very short, black. Capitula 20
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<normalizedToken originalValue="–30(–">-30(-</normalizedToken>
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45) mm in diameter, subtruncate at base. Involucral bracts 9-11
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<normalizedToken originalValue="×">x</normalizedToken>
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(1.7
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<normalizedToken originalValue="–)2.0–">-)2.0-</normalizedToken>
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2.5 mm (the outermost from
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<emphasis italics="true" pageId="1" pageNumber="2">c.</emphasis>
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5 mm long), all
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<normalizedToken originalValue="linear–lanceolate">linear-lanceolate</normalizedToken>
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,
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<normalizedToken originalValue="blackish–green">blackish-green</normalizedToken>
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; apex obtuse; stellate hairs
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<pageBreakToken pageId="2" pageNumber="3" start="start">sparse</pageBreakToken>
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, often with a tuft at the apex; simple eglandular hairs many, short to medium,
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<normalizedToken originalValue="dark–based">dark-based</normalizedToken>
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; glandular hairs many, very short, black. Ligules yellow,
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<normalizedToken originalValue="glabrous–tipped">glabrous-tipped</normalizedToken>
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. Styles discoloured. Achenes 4.0-4.5 mm,
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<normalizedToken originalValue="blackish–brown">blackish-brown</normalizedToken>
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. Margins of receptacle pits
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<normalizedToken originalValue="long–dentate">long-dentate</normalizedToken>
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. Chromosome number 2n = 36 (cf.
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<bibRefCitation author="Sell, PD" journalOrPublisher="Cambridge University Press, Cambridge" pageId="4" pageNumber="5" pagination="218 - 421" refId="B7" refString="Sell, PD, Murrell, G, 2006. Flora of Great Britain and Ireland, volume 4. Cambridge University Press, Cambridge: 218 - 421" title="Flora of Great Britain and Ireland, volume 4." year="2006">Sell and Murrell 2006</bibRefCitation>
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, as
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="dovrense">
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<emphasis italics="true" pageId="2" pageNumber="3">Hieracium dovrense</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/9872" pageId="2" pageNumber="3" start="Figure 2" startId="F2">
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<paragraph pageId="2" pageNumber="3">
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<emphasis bold="true" pageId="2" pageNumber="3">Figure 2.</emphasis>
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Middle stem leaves.
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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</emphasis>
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possible
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<taxonomicName authorityName="T. C. G. Rich" authorityYear="2011" class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="maccoshiana">
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<emphasis italics="true" pageId="2" pageNumber="3">Hieracium maccoshiana</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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</emphasis>
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Ben Loyal (NMW 28.131.5185).
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<emphasis bold="true" pageId="2" pageNumber="3">C</emphasis>
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Ben Loyal (BM).
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="D–N">D-N</normalizedToken>
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</emphasis>
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<taxonomicName authorityName="T. C. G. Rich" authorityYear="2011" class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="maccoshiana">
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<emphasis italics="true" pageId="2" pageNumber="3">Hieracium maccoshiana</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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</emphasis>
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Rhiconich gorge (NMW V.2005.1.159).
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="G–H">G-H</normalizedToken>
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</emphasis>
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Foinaven (CGE).
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="I–L">I-L</normalizedToken>
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</emphasis>
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Rhiconich (CGE).
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="M–N">M-N</normalizedToken>
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</emphasis>
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Craig na
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<normalizedToken originalValue="H’Uidhe">H'Uidhe</normalizedToken>
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(CGE).
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="O–U">O-U</normalizedToken>
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</emphasis>
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="dovrense">
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<emphasis italics="true" pageId="2" pageNumber="3">Hieracium dovrense</emphasis>
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</taxonomicName>
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, Drivstua, Dovrefj, Norway (S).
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="O–Q">O-Q</normalizedToken>
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</emphasis>
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S09-16213.
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<emphasis bold="true" pageId="2" pageNumber="3">R</emphasis>
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S09-16217.
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<emphasis bold="true" pageId="2" pageNumber="3">S</emphasis>
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S09-16224.
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<emphasis bold="true" pageId="2" pageNumber="3">
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<normalizedToken originalValue="T–U">T-U</normalizedToken>
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</emphasis>
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S09-16222. Scale bar 5 cm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="distribution">
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<paragraph pageId="2" pageNumber="3">Distribution.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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Endemic to Sutherland, Scotland, where it is known from four sites in Sutherland: in a gorge and on a burn side near Rhiconich; on a burn side in Coire
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<normalizedToken originalValue="Dùail">Duail</normalizedToken>
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, Foinaven; and on rocks at Creag na h-Uidhe (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of Hieracium maccoshiana in Scotland." httpUri="https://binary.pensoft.net/fig/9873" pageId="2" pageNumber="3">Fig. 3</figureCitation>
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). The status of plants from Ben Loyal remains to be clarified when it has been refound in the field
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<pageBreakToken pageId="3" pageNumber="4" start="start">.</pageBreakToken>
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/9873" pageId="3" pageNumber="4" start="Figure 3" startId="F3">
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<paragraph pageId="3" pageNumber="4">
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<emphasis bold="true" pageId="3" pageNumber="4">Figure 3.</emphasis>
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Distribution of
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<taxonomicName authorityName="T. C. G. Rich" authorityYear="2011" class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="4" phylum="Angiospermae" rank="species" species="maccoshiana">
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<emphasis italics="true" pageId="3" pageNumber="4">Hieracium maccoshiana</emphasis>
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</taxonomicName>
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in Scotland.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="ecology">
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<paragraph pageId="3" pageNumber="4">Ecology.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Mountain cliff ledges and rocks, in rocky ravines and on riversides. It mostly occurs out of the reach of large herbivores such as deer or sheep as, like most leafy hawkweeds, is likely to be selectively eaten. The altitudes range from 130-414 m. It flowers from late July to August, and is probably apomictic. Some plants may produce sparse pollen and others none (
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<bibRefCitation author="Thomas, S" journalOrPublisher="Edinburgh Journal of Botany" pageId="4" pageNumber="5" refId="B11" refString="Thomas, S, Bailey, J, Rich, TCG, in press. Pollen and cytological studies in Hieracium section Alpestria (Asteraceae). In: Nordic Journal of Botany." title="Pollen and cytological studies in Hieracium section Alpestria (Asteraceae)" volumeTitle="Nordic Journal of Botany." year="in press">Thomas et al. 2011</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<emphasis bold="true" pageId="3" pageNumber="4">Etymology</emphasis>
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. The epithet honours David J. McCosh for his work in mapping British and Irish
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="4" phylum="Angiospermae" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Hieracium</emphasis>
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</taxonomicName>
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taxa over the last 30 years, and for mentoring me. The spelling
|
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<normalizedToken originalValue="‘maccoshiana’">
|
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'
|
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<emphasis italics="true" pageId="3" pageNumber="4">maccoshiana</emphasis>
|
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</normalizedToken>
|
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' follows recommendation 60C.5.a of the International Code of Botanical Nomenclature (Vienna Code McNeil et al. 2006).
|
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</paragraph>
|
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<paragraph pageId="3" pageNumber="4">
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|
<emphasis bold="true" pageId="3" pageNumber="4">Conservation status</emphasis>
|
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. Surveys in 2010 revealed about 60 plants in four sites (see supplementary file 1:
|
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Hieracium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="4" phylum="Angiospermae" rank="species" species="dovrense">
|
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<emphasis italics="true" pageId="3" pageNumber="4">Hieracium dovrense</emphasis>
|
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</taxonomicName>
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survey 2010). It is thus best considered at
|
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<bibRefCitation author="IUCN," journalOrPublisher="International Union for Conservation of Nature, Gland" pageId="4" pageNumber="5" refId="B2" refString="IUCN, , 2001. IUCN Red List Categories. Version 3.1. International Union for Conservation of Nature, Gland" title="IUCN Red List Categories. Version 3.1." year="2001">IUCN (2001)</bibRefCitation>
|
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threat status
|
|||
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|
<normalizedToken originalValue="‘Endangered’">'Endangered'</normalizedToken>
|
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due to the small population size.
|
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</paragraph>
|
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</subSubSection>
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</treatment>
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</document>
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