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<document id="32FD7689CFE7CC31B1DD4A5B47AEC22B" ID-CLB-Dataset="26985" ID-DOI="10.5852/ejt.2019.519" ID-GBIF-Dataset="c6f362bc-4d6b-4e3a-b69a-a9e8392e6756" ID-ISSN="2118-9773" ID-Zenodo-Dep="2650112" ID-ZooBank="D9E64DD6-D675-4E25-B9B8-A44C14CE0B22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556106999488" checkinUser="plazi" docAuthor="Oliveira, Livia M., Hoeksema, Bert W. &amp; Rocha, Rosana M." docDate="2019" docId="039C8788FFAF2165FE108EDCFD7E7479" docLanguage="en" docName="ejt-519_oliveira_hoeksema_rocha.pdf.imf" docOrigin="European Journal of Taxonomy 519" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Polysyncraton purou C. Monniot &amp; F. Monniot 1987" docType="treatment" docVersion="19" lastPageNumber="17" masterDocId="FFA5FFF0FFBF2175FFCB8F2FFFB4710F" masterDocTitle="Polysyncraton (Ascidiacea, Didemnidae): a re-examination of some specimens and descriptions of three new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="17" updateTime="1698723430310" updateUser="ExternalLinkService">
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<mods:title id="A13AAB3B5624652802AAD83C7FF10373">Polysyncraton (Ascidiacea, Didemnidae): a re-examination of some specimens and descriptions of three new species</mods:title>
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<mods:namePart id="0DB4DE28BB6F0E4FB6CCF5AC9CF2CB0F">Oliveira, Livia M.</mods:namePart>
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<mods:affiliation id="4BE897346849BDED0E9634C6FBF6B972">Laboratório de Sistemática e Ecologia de Invertebrados Marinhos, Departamento de Zoologia, Universidade Federal do Paraná, CP 19020, 81531 - 980 Curitiba, Brazil.</mods:affiliation>
<mods:nameIdentifier id="49C3082F6532CB53193A9ACD367775B4" type="email">liviaoliveira@ufpr.br</mods:nameIdentifier>
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<mods:namePart id="EFE2E5CAF133E23C646EA70E8550B9D5">Hoeksema, Bert W.</mods:namePart>
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<mods:affiliation id="F36A74F41A728F68909062BEE67A6FDB">Department of Taxonomy and Systematics, Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.</mods:affiliation>
<mods:nameIdentifier id="46F31AEEF3895107B1E3F5206146C477" type="email">bert.hoeksema@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="8FBF70D7D873708FA5CB613974852B58">Rocha, Rosana M.</mods:namePart>
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<mods:affiliation id="37D1DD65B30F153F0951EAEE5C8D35C8">Laboratório de Sistemática e Ecologia de Invertebrados Marinhos, Departamento de Zoologia, Universidade Federal do Paraná, CP 19020, 81531 - 980 Curitiba, Brazil.</mods:affiliation>
<mods:nameIdentifier id="853F64C6C2E651631613DC908A7D7587" type="email">rmrocha@ufpr.br</mods:nameIdentifier>
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<mods:title id="42CFFBCACB171B3F8BE5E8C485EBD52B">European Journal of Taxonomy</mods:title>
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<mods:number id="AB790999E0155A540A2AF3C4D1B4AA0F">519</mods:number>
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<subSubSection id="C32F6515FFAF2165FE108EDCFBEC7301" box="[475,1112,499,526]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B8A369EFFAF2165FE108EDCFBEC7301" blockId="16.[475,1112,499,560]" box="[475,1112,499,526]" pageId="16" pageNumber="17">
<taxonomicName id="4C354D1DFFAF2165FE108EDCFBEC7301" authority="C. Monniot &amp; F. Monniot, 1987" authorityName="C. Monniot &amp; F. Monniot" authorityYear="1987" box="[475,1112,499,526]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="purou">
<emphasis id="B941EA8CFFAF2165FE108EDCFD6C7301" bold="true" box="[475,728,499,526]" italics="true" pageId="16" pageNumber="17">Polysyncraton purou</emphasis>
C. Monniot &amp; F.
<bibRefCitation id="EFA44B6FFFAF2165FC618EDBFBEC7301" author="Monniot" box="[938,1112,500,526]" firstAuthor="Monniot" pageId="16" pageNumber="17" pagination="1 - 155" refId="ref9102" refString="Monniot C. &amp; Monniot F. 1987. Les ascidies de Polynesie francaise. Memoires du Museum national d'Histoire naturelle, Zoologie 136: 1 - 155." type="journal article" year="1987">Monniot, 1987</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C32F6515FFAF2165FD3B8D39FCF7733F" box="[752,835,534,560]" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8A369EFFAF2165FD3B8D39FCF7733F" blockId="16.[475,1112,499,560]" box="[752,835,534,560]" pageId="16" pageNumber="17">
<figureCitation id="130E2A1BFFAF2165FD3B8D39FCF7733F" box="[752,835,534,560]" captionStart="Fig" captionStartId="17.[189,232,1895,1921]" captionTargetBox="[194,1399,404,1840]" captionTargetId="figure@17.[189,1399,388,1845]" captionTargetPageId="17" captionText="Fig. 10. Polysyncraton purou C. Monniot &amp; F. Monniot, 1987, with structures stained with hemalum (MNHN A2-843). a. Zooid with bifurcated atrial languet (arrow). b. Thorax with button-like atrial languet (big arrow) and muscular process (small arrow). c. Abdomen with gonads (large arrow indicates oocyte). d. Immature larva. e. Mature larva. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3239961/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F6515FFAF2165FF768D78FDBE73DB" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8B8A369EFFAF2165FF768D78FB83737E" blockId="16.[189,1079,599,626]" box="[189,1079,599,626]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB2511B0FFAF2165FF768D78FB83737E" box="[189,1079,599,626]" pageId="16" pageNumber="17">
<taxonomicName id="4C354D1DFFAF2165FF768D78FCD5737E" authority="C. Monniot &amp; F. Monniot, 1987: 49" authorityName="C. Monniot &amp; F. Monniot" authorityPageNumber="49" authorityYear="1987" box="[189,865,599,626]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="purou">
<emphasis id="B941EA8CFFAF2165FF768D78FE07737E" box="[189,435,599,625]" italics="true" pageId="16" pageNumber="17">Polysyncraton purou</emphasis>
C. Monniot &amp; F.
<treatmentCitation id="0A94108FFFAF2165FD488D78FCD5737E" author="Monniot C. &amp; Monniot F." box="[643,865,599,625]" page="49" pageId="16" pageNumber="17" year="1987">
<bibRefCitation id="EFA44B6FFFAF2165FD488D78FCD5737E" author="Monniot" box="[643,865,599,625]" firstAuthor="Monniot" pageId="16" pageNumber="17" pagination="1 - 155" refId="ref9102" refString="Monniot C. &amp; Monniot F. 1987. Les ascidies de Polynesie francaise. Memoires du Museum national d'Histoire naturelle, Zoologie 136: 1 - 155." type="journal article" year="1987">Monniot, 1987: 49</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 16d–h, pl.
<date id="FF8B105EFFAF2165FBD38D78FB83737E" box="[1048,1079,599,625]" pageId="16" pageNumber="17">3i.</date>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8A369EFFAF2165FF768DB7FDBE73DB" blockId="16.[189,1398,664,725]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB2511B0FFAF2165FF768DB7FDBE73DB" pageId="16" pageNumber="17">
<taxonomicName id="4C354D1DFFAF2165FF768DB7FE0573BD" authorityName="C. Monniot &amp; F. Monniot" authorityYear="1987" box="[189,433,664,690]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="purou">
<emphasis id="B941EA8CFFAF2165FF768DB7FE0573BD" box="[189,433,664,690]" italics="true" pageId="16" pageNumber="17">Polysyncraton purou</emphasis>
</taxonomicName>
– F.
<treatmentCitation id="0A94108FFFAF2165FE2D8DB7FD0C73BD" author="Monniot F." box="[486,696,664,690]" page="11" pageId="16" pageNumber="17" year="1993">
<bibRefCitation id="EFA44B6FFFAF2165FE2D8DB7FD0C73BD" author="Monniot" box="[486,696,664,690]" firstAuthor="Monniot" pageId="16" pageNumber="17" pagination="3 - 17" refId="ref9261" refString="Monniot F. 1993. Ascidies de Nouvelle-Caledonie XIII. Le genre Polysyncraton. Bulletin du Museum national d'Histoire naturelle, 4 ieme serie 15 ª (1 - 4): 3 - 17." type="journal article" year="1993">Monniot 1993: 11</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0A94108FFFAF2165FD2E8DB7FC2273BD" author="Kott P." box="[741,918,664,690]" page="126" pageId="16" pageNumber="17" year="2001">
<bibRefCitation id="EFA44B6FFFAF2165FD2E8DB7FC2273BD" author="Kott" box="[741,918,664,690]" firstAuthor="Kott" pageId="16" pageNumber="17" pagination="1 - 410" refId="ref8579" refString="Kott P. 2001. The Australian Ascidiacea. Part 4, Aplousobranchia (3), Didemnidae. Memoirs of the Queensland Museum 47 (1): 1 - 410. Available from https: // biodiversitylibrary. org / page / 40460926 [accessed 20 Jul. 2018]." type="journal article" year="2001">Kott 2001: 126</bibRefCitation>
</treatmentCitation>
, figs 60, 162C, pl. 6F; 2002: 33, fig. 12A– D; 2005: 2436, fig. 19D.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32F6515FFAF2165FF768DD4FB96724C" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B8A369EFFAF2165FF768DD4FE1E721A" blockId="16.[189,426,763,789]" box="[189,426,763,789]" pageId="16" pageNumber="17">
<heading id="D0C281F2FFAF2165FF768DD4FE1E721A" bold="true" box="[189,426,763,789]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B941EA8CFFAF2165FF768DD4FE1E721A" bold="true" box="[189,426,763,789]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A369EFFAF2165FF768C06FB96724C" blockId="16.[189,1058,809,835]" box="[189,1058,809,835]" pageId="16" pageNumber="17">
<collectingCountry id="F322760EFFAF2165FF768C06FEE5724C" box="[189,337,809,835]" name="Tanzania" pageId="16" pageNumber="17">TANZANIA</collectingCountry>
• 3 slides; MNHN A2-843, MNHN A2-846, MNHN A2-847.
</paragraph>
</subSubSection>
<subSubSection id="C32F6515FFAF2165FF768C45FAD375EB" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B8A369EFFAF2165FF768C45FE9B728B" blockId="16.[189,303,874,900]" box="[189,303,874,900]" pageId="16" pageNumber="17">
<heading id="D0C281F2FFAF2165FF768C45FE9B728B" bold="true" box="[189,303,874,900]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B941EA8CFFAF2165FF768C45FE9B728B" bold="true" box="[189,303,874,900]" pageId="16" pageNumber="17">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A369EFFAF2165FF768CB8FC9E72F9" blockId="16.[189,1399,919,1014]" pageId="16" pageNumber="17">
This species was described from
<collectingCountry id="F322760EFFAF2165FDAB8CB8FC8572BE" box="[608,817,919,945]" name="French Polynesia" pageId="16" pageNumber="17">French Polynesia</collectingCountry>
by C. Monniot &amp; F.
<bibRefCitation id="EFA44B6FFFAF2165FB878CB8FAA472BD" author="Monniot" box="[1100,1296,919,946]" firstAuthor="Monniot" pageId="16" pageNumber="17" pagination="1 - 155" refId="ref9102" refString="Monniot C. &amp; Monniot F. 1987. Les ascidies de Polynesie francaise. Memoires du Museum national d'Histoire naturelle, Zoologie 136: 1 - 155." type="journal article" year="1987">Monniot (1987)</bibRefCitation>
. All the characteristics of the specimens from
<collectingCountry id="F322760EFFAF2165FDBC8C95FD5472DB" box="[631,736,954,980]" name="Tanzania" pageId="16" pageNumber="17">Tanzania</collectingCountry>
correspond to those in the original description (
<figureCitation id="130E2A1BFFAF2165FADA8C95FADC72DB" box="[1297,1384,954,980]" captionStart="Fig" captionStartId="17.[189,232,1895,1921]" captionTargetBox="[194,1399,404,1840]" captionTargetId="figure@17.[189,1399,388,1845]" captionTargetPageId="17" captionText="Fig. 10. Polysyncraton purou C. Monniot &amp; F. Monniot, 1987, with structures stained with hemalum (MNHN A2-843). a. Zooid with bifurcated atrial languet (arrow). b. Thorax with button-like atrial languet (big arrow) and muscular process (small arrow). c. Abdomen with gonads (large arrow indicates oocyte). d. Immature larva. e. Mature larva. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3239961/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
), thus enlarging the known distribution of this species.
</paragraph>
<paragraph id="8B8A369EFFAF2165FF768B32FAD375EB" blockId="16.[189,1399,1053,1252]" pageId="16" pageNumber="17">
<bibRefCitation id="EFA44B6FFFAF2165FF768B32FDB67538" accessDate="2018-07-20" author="Monniot C. &amp; Monniot F. &amp; Griffiths C. L. &amp; Schleyer M." box="[189,514,1053,1079]" journalOrPublisher="Annals of the South African Museum" pageId="16" pageNumber="17" pagination="1 - 141" part="108" publicationUrl="https://biodiversitylibrary.org/page/40905473" refId="ref9163" refString="Monniot C., Monniot F., Griffiths C. L. &amp; Schleyer M. 2001. South African ascidians. Annals of the South African Museum 108 (1): 1 - 141. Available from https: // biodiversitylibrary. org / page / 40905473 [accessed 20 Jul. 2018]." title="South African ascidians" type="journal article" year="2001">Monniot &amp; Monniot (2001)</bibRefCitation>
described
<taxonomicName id="4C354D1DFFAF2165FD498B31FD487538" authorityName="F.Monniot &amp; C.Monniot" authorityYear="2001" box="[642,764,1054,1079]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vestiens">
<emphasis id="B941EA8CFFAF2165FD498B31FD487538" box="[642,764,1054,1079]" italics="true" pageId="16" pageNumber="17">P. vestiens</emphasis>
</taxonomicName>
from
<collectingCountry id="F322760EFFAF2165FC8F8B32FC317538" box="[836,901,1053,1079]" name="Palau" pageId="16" pageNumber="17">Palau</collectingCountry>
as a different species from
<taxonomicName id="4C354D1DFFAF2165FB018B31FA987538" authorityName="C. Monniot &amp; F. Monniot" authorityYear="1987" box="[1226,1324,1054,1079]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="purou">
<emphasis id="B941EA8CFFAF2165FB018B31FA987538" box="[1226,1324,1054,1079]" italics="true" pageId="16" pageNumber="17">P. purou</emphasis>
</taxonomicName>
based on the smaller spicules, a wider atrial aperture and the absence of the muscular process in
<taxonomicName id="4C354D1DFFAF2165FB3F8B6FFADB7556" authorityName="F.Monniot &amp; C.Monniot" authorityYear="2001" box="[1268,1391,1088,1113]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vestiens">
<emphasis id="B941EA8CFFAF2165FB3F8B6FFADB7556" box="[1268,1391,1088,1113]" italics="true" pageId="16" pageNumber="17">P. vestiens</emphasis>
</taxonomicName>
. However, in the specimens from
<collectingCountry id="F322760EFFAF2165FD808B4DFD007573" box="[587,692,1122,1148]" name="Tanzania" pageId="16" pageNumber="17">Tanzania</collectingCountry>
there is a variation in the size of the spicules and they are smaller (
<quantity id="4CCD9B7BFFAF2165FEE98BAAFE067590" box="[290,434,1156,1183]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" metricValueMax="5.5" metricValueMin="4.3" pageId="16" pageNumber="17" unit="mm" value="49.0" valueMax="55.0" valueMin="43.0">43 to 55 µm</quantity>
) than in specimens from Tahiti. The muscular process in
<taxonomicName id="4C354D1DFFAF2165FB818BAAFB1F7591" authorityName="C. Monniot &amp; F. Monniot" authorityYear="1987" box="[1098,1195,1157,1182]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="purou">
<emphasis id="B941EA8CFFAF2165FB818BAAFB1F7591" box="[1098,1195,1157,1182]" italics="true" pageId="16" pageNumber="17">P. purou</emphasis>
</taxonomicName>
is very small and not visible in some zooids (
<figureCitation id="130E2A1BFFAF2165FDC98B88FDE375CD" box="[514,599,1191,1218]" captionStart="Fig" captionStartId="17.[189,232,1895,1921]" captionTargetBox="[194,1399,404,1840]" captionTargetId="figure@17.[189,1399,388,1845]" captionTargetPageId="17" captionText="Fig. 10. Polysyncraton purou C. Monniot &amp; F. Monniot, 1987, with structures stained with hemalum (MNHN A2-843). a. Zooid with bifurcated atrial languet (arrow). b. Thorax with button-like atrial languet (big arrow) and muscular process (small arrow). c. Abdomen with gonads (large arrow indicates oocyte). d. Immature larva. e. Mature larva. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3239961/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
a–b). Since the size of the atrial aperture is also a variable character that depends on the contraction of the zooid, there is doubt as to whether
<taxonomicName id="4C354D1DFFAF2165FBD38BE5FB2675EC" authorityName="F.Monniot &amp; C.Monniot" authorityYear="2001" box="[1048,1170,1226,1251]" class="Ascidiacea" family="Didemnidae" genus="Polysyncraton" kingdom="Animalia" order="Enterogona" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vestiens">
<emphasis id="B941EA8CFFAF2165FBD38BE5FB2675EC" box="[1048,1170,1226,1251]" italics="true" pageId="16" pageNumber="17">P. vestiens</emphasis>
</taxonomicName>
is a valid species.
</paragraph>
</subSubSection>
<subSubSection id="C32F6515FFAF2165FF768A25FD7E7479" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B8A369EFFAF2165FF768A25FEE2742B" blockId="16.[189,342,1290,1316]" box="[189,342,1290,1316]" pageId="16" pageNumber="17">
<heading id="D0C281F2FFAF2165FF768A25FEE2742B" bold="true" box="[189,342,1290,1316]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B941EA8CFFAF2165FF768A25FEE2742B" bold="true" box="[189,342,1290,1316]" pageId="16" pageNumber="17">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A369EFFAF2165FF768A16FD7E7479" blockId="16.[189,1398,1336,1398]" pageId="16" pageNumber="17">
<typeStatus id="548E883CFFAF2165FF768A16FF43745C" box="[189,247,1337,1363]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
<collectingCountry id="F322760EFFAF2165FEA88A17FD99745D" box="[355,557,1336,1362]" name="French Polynesia" pageId="16" pageNumber="17">French Polynesia</collectingCountry>
(C. Monniot &amp; F.
<bibRefCitation id="EFA44B6FFFAF2165FCCB8A17FC1E745D" author="Monniot" box="[768,938,1336,1362]" firstAuthor="Monniot" pageId="16" pageNumber="17" pagination="1 - 155" refId="ref9102" refString="Monniot C. &amp; Monniot F. 1987. Les ascidies de Polynesie francaise. Memoires du Museum national d'Histoire naturelle, Zoologie 136: 1 - 155." type="journal article" year="1987">Monniot 1987</bibRefCitation>
).
<collectingCountry id="F322760EFFAF2165FC708A17FB93745D" box="[955,1063,1336,1362]" name="Australia" pageId="16" pageNumber="17">Australia</collectingCountry>
(
<bibRefCitation id="EFA44B6FFFAF2165FBFF8A17FB1B745D" author="Kott" box="[1076,1199,1336,1362]" firstAuthor="Kott" pageId="16" pageNumber="17" pagination="1 - 410" refId="ref8579" refString="Kott P. 2001. The Australian Ascidiacea. Part 4, Aplousobranchia (3), Didemnidae. Memoirs of the Queensland Museum 47 (1): 1 - 410. Available from https: // biodiversitylibrary. org / page / 40460926 [accessed 20 Jul. 2018]." type="journal article" year="2001">Kott 2001</bibRefCitation>
),
<collectingCountry id="F322760EFFAF2165FB098A16FAC2745D" box="[1218,1398,1336,1363]" name="New Caledonia" pageId="16" pageNumber="17">New Caledonia</collectingCountry>
(F.
<bibRefCitation id="EFA44B6FFFAF2165FF288A74FE38747A" author="Monniot" box="[227,396,1371,1397]" firstAuthor="Monniot" pageId="16" pageNumber="17" pagination="3 - 17" refId="ref9261" refString="Monniot F. 1993. Ascidies de Nouvelle-Caledonie XIII. Le genre Polysyncraton. Bulletin du Museum national d'Histoire naturelle, 4 ieme serie 15 ª (1 - 4): 3 - 17." type="journal article" year="1993">Monniot 1993</bibRefCitation>
) and
<collectingCountry id="F322760EFFAF2165FE068A74FD82747A" box="[461,566,1371,1397]" name="Tanzania" pageId="16" pageNumber="17">Tanzania</collectingCountry>
(this study).
</paragraph>
</subSubSection>
</treatment>
</document>
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