treatments-xml/data/03/D5/69/03D56948FFA5403D4CA9FF47FECA6F91.xml

<document id="2C49BD98C76D0AECEBA5BE865644D624" ID-DOI="10.11646/phytotaxa.642.1.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13214159" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722601820667" checkinUser="felipe" docAuthor="Lyu, Tong, Tang, Song-Ming, He, Jun, Wang, Shuai, Wu, Xiao-Qu, Chen, De-Chao, Ao, Cheng-Ce &amp; Li, Shu-Hong" docDate="2024" docId="03D56948FFA5403D4CA9FF47FECA6F91" docLanguage="en" docName="phytotaxa.642.1.4.pdf" docOrigin="Phytotaxa 642 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.642.1.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Lyophyllum lixivium S. M. Tang &amp; S. H. Li, sp. nov." docType="treatment" docVersion="2" lastPageNumber="57" masterDocId="FFEC1130FFA0403A4C21FFB3FFD16D1D" masterDocTitle="Lyophyllum lixivium (Lyophyllaceae, Agaricales), a new species from Yunnan, China" masterLastPageNumber="60" masterPageNumber="50" pageNumber="55" updateTime="1722804533030" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0C6B393B63EA84C0F782C49B700BC50F">Lyophyllum lixivium (Lyophyllaceae, Agaricales), a new species from Yunnan, China</mods:title>
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<mods:namePart id="FDC04A6095BE010C844B59484EC55FE8">Lyu, Tong</mods:namePart>
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<mods:namePart id="2212A3F2255C68CE5952AA8B99717EF9">Tang, Song-Ming</mods:namePart>
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<mods:namePart id="6BA8DDA35092FD1FEB7FAF4776A556EF">He, Jun</mods:namePart>
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<mods:affiliation id="1ABA19600D9346512BEE85CACD2E83C4">College of Biotechnology and Bioengineering, West Yunnan University, Lincang 677000, China</mods:affiliation>
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<mods:namePart id="D0BE14EE3620D5CFD57D14B4AC3E5629">Wang, Shuai</mods:namePart>
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<mods:affiliation id="A9DADB24C1E842A9DB7884324B31FD43">School of Agriculture, Yunnan University, Kunming 650500, China &amp; Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences, Kunming 650205, China</mods:affiliation>
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<mods:namePart id="1B8771769307088EDBA91558D49BD53C">Wu, Xiao-Qu</mods:namePart>
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<mods:affiliation id="6E99FD283BB8FF82718E82FFE68FFE9D">School of Agriculture, Yunnan University, Kunming 650500, China &amp; Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences, Kunming 650205, China</mods:affiliation>
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<mods:namePart id="783CD0766A8F812FDF6B3BFC641E9E6E">Ao, Cheng-Ce</mods:namePart>
<mods:affiliation id="0AB2DEB1A86790013544ED56643B6CE5">Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences, Kunming 650205, China &amp; College of Agriculture and Biological Science, Dali University, Dali 671003, China</mods:affiliation>
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<mods:namePart id="B696E5B5FA70EADB3AB0FE4002961AC9">Li, Shu-Hong</mods:namePart>
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<mods:date id="6F09FAA456D9B24AA0E969C1DC9960DD">2024</mods:date>
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<paragraph id="8BC3D85EFFA5403F4CA9FF47FC866C13" blockId="5.[136,855,244,271]" box="[136,855,244,271]" pageId="5" pageNumber="55">
<heading id="D08B6F32FFA5403F4CA9FF47FC866C13" box="[136,855,244,271]" fontSize="11" level="2" pageId="5" pageNumber="55" reason="3">
<taxonomicName id="4C7CA3DDFFA5403F4CA9FF47FDA76C13" authority="S. M. Tang &amp; S. H. Li" authorityName="S. M. Tang &amp; S. H. Li" box="[136,630,244,271]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="5" pageNumber="55" phylum="Basidiomycota" rank="species" species="lixivium" status="sp. nov.">
<emphasis id="B908044CFFA5403F4CA9FF47FEA66C12" bold="true" box="[136,375,244,271]" italics="true" pageId="5" pageNumber="55">Lyophyllum lixivium</emphasis>
S. M. Tang &amp; S. H. Li
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<figureCitation id="1347C4DBFFA5403F4EF8FF47FCF46C13" box="[729,805,244,270]" captionStart="FIGURE 2" captionStartId="5.[136,229,1542,1564]" captionTargetBox="[152,1436,583,1518]" captionTargetId="figure-121@5.[152,1436,583,1518]" captionTargetPageId="5" captionText="FIGURE 2. Lyophyllum lixivium. a Basidiomata (HKAS129929). b: Hymenophoral trama.c: Pileipellis.d: Stipitipellis (presence fusiform caulocystidia). Bars: a = 5 mm, b–f = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13214163" httpUri="https://zenodo.org/record/13214163/files/figure.png" pageId="5" pageNumber="55">Figs. 2</figureCitation>
&amp; 
<figureCitation id="1347C4DBFFA5403F4F68FF47FC866C13" box="[841,855,244,270]" captionStart="FIGURE 3" captionStartId="6.[136,229,1834,1856]" captionTargetBox="[151,1436,281,1809]" captionTargetId="figure-65@6.[151,1436,281,1809]" captionTargetPageId="6" captionText="FIGURE 3. Lyophyllum lixivium (HKAS129929). a: Cheilocystidia. b: Pleurocystidia. c: Basidia. d: Basidiospores as seen under the microscope. e: Clamp connections. f. Scanning electron microscope photo of basidiospores. Bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13214165" httpUri="https://zenodo.org/record/13214165/files/figure.png" pageId="5" pageNumber="55">3</figureCitation>
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</paragraph>
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<emphasis id="B908044CFFA5403F4CA9FEAFFF296C2C" bold="true" box="[136,248,284,305]" pageId="5" pageNumber="55">MycoBank</emphasis>
: MB849904
</paragraph>
</subSubSection>
<subSubSection id="C3668BD5FFA5403F4CA9FED3FC696C67" box="[136,952,352,379]" pageId="5" pageNumber="55" type="etymology">
<paragraph id="8BC3D85EFFA5403F4CA9FED3FC696C67" blockId="5.[136,1452,352,559]" box="[136,952,352,379]" pageId="5" pageNumber="55">
<emphasis id="B908044CFFA5403F4CA9FED3FED86C67" bold="true" box="[136,265,352,378]" pageId="5" pageNumber="55">Etymology</emphasis>
: 
<taxonomicName id="4C7CA3DDFFA5403F4D39FED3FEAA6C67" box="[280,379,352,378]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="5" pageNumber="55" phylum="Basidiomycota" rank="species" species="lixivium">
‘ 
<emphasis id="B908044CFFA5403F4D00FED3FEAA6C67" box="[289,379,352,378]" italics="true" pageId="5" pageNumber="55">lixivium</emphasis>
</taxonomicName>
’ refers to the light brown colouration of the pileus.
</paragraph>
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<paragraph id="8BC3D85EFFA5403F4C9CFE37FCFC6CDF" blockId="5.[136,1452,352,559]" pageId="5" pageNumber="55">
<materialsCitation id="3B14D203FFA5403F4C9CFE37FCFC6CDF" collectingDate="2020-08-11" collectionCode="HKAS" country="China" county="Cangshan Mountain" elevation="2371" latitude="25.698332" location="Yunanan" longLatPrecision="15" longitude="100.04095" municipality="Dali City" pageId="5" pageNumber="55" specimenCode="HKAS129929" specimenCount="2" typeStatus="holotype">
<emphasis id="B908044CFFA5403F4C9CFE37FEFB6C83" bold="true" box="[189,298,388,414]" pageId="5" pageNumber="55">
<typeStatus id="54C766FCFFA5403F4C9CFE37FEFB6C83" box="[189,298,388,414]" pageId="5" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectingCountry id="F36B98CEFFA5403F4D1CFE37FE526C83" box="[317,387,388,414]" name="China" pageId="5" pageNumber="55">China</collectingCountry>
, 
<location id="8EA38E85FFA5403F4DB0FE36FE276C82" LSID="urn:lsid:plazi:treatment:03D56948FFA5403D4CA9FF47FECA6F91:8EA38E85FFA5403F4DB0FE36FE276C82" box="[401,502,389,415]" country="China" county="Cangshan Mountain" latitude="25.698332" longLatPrecision="15" longitude="100.04095" municipality="Dali City" name="Yunanan" pageId="5" pageNumber="55">Yunanan</location>
, 
<collectingMunicipality id="6BA74224FFA5403F4E24FE37FDA36C83" box="[517,626,388,414]" pageId="5" pageNumber="55">Dali City</collectingMunicipality>
, 
<collectingCounty id="62A2A0D2FFA5403F4EA0FE37FCBA6C83" box="[641,875,388,414]" pageId="5" pageNumber="55">Cangshan Mountain</collectingCounty>
, 
<geoCoordinate id="EE48BE99FFA5403F4F5BFE37FBC06C82" box="[890,1041,388,415]" degrees="25" direction="north" minutes="41" orientation="latitude" pageId="5" pageNumber="55" precision="15" seconds="54" value="25.698332">25°41’54” N</geoCoordinate>
, 
<geoCoordinate id="EE48BE99FFA5403F4801FE37FB076C82" box="[1056,1238,388,415]" degrees="100" direction="east" minutes="2" orientation="longitude" pageId="5" pageNumber="55" precision="1" seconds="27.42" value="100.04095">100°2’27.42” E</geoCoordinate>
, elev. 
<quantity id="4C8475BBFFA5403F4904FE37FA516C82" box="[1317,1408,388,415]" metricMagnitude="3" metricUnit="m" metricValue="2.371" pageId="5" pageNumber="55" unit="m" value="2371.0">
<elevation id="00513F6DFFA5403F4904FE37FA516C82" box="[1317,1408,388,415]" metricMagnitude="3" metricUnit="m" metricValue="2.371" pageId="5" pageNumber="55" unit="m" value="2371.0">2371 m</elevation>
</quantity>
, 
<date id="FFC2FE9EFFA5403F49B1FE37FECA6CDF" pageId="5" pageNumber="55" value="2020-08-11">
<collectingDate id="EF860776FFA5403F49B1FE37FECA6CDF" pageId="5" pageNumber="55" value="2020-08-11">11 August 2020</collectingDate>
</date>
, Zhang Zhengquan (
<specimenCode id="DBDA7025FFA5403F4E20FE1BFD776CDF" box="[513,678,424,450]" collectionCode="HKAS" country="China" httpUri="http://grbio.org/cool/di2a-40jb" name="Cryptogamic Herbarium of Kunming Institute of Botany" pageId="5" pageNumber="55">HKAS129929</specimenCode>
, 
<typeStatus id="54C766FCFFA5403F4E90FE1BFCCF6CDF" box="[689,798,424,450]" pageId="5" pageNumber="55" type="holotype">holotype</typeStatus>
!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3668BD5FFA5403F4C9CFE7FFD3A6F33" pageId="5" pageNumber="55" type="diagnosis">
<paragraph id="8BC3D85EFFA5403F4C9CFE7FFD3A6F33" blockId="5.[136,1452,352,559]" pageId="5" pageNumber="55">
<emphasis id="B908044CFFA5403F4C9CFE7FFEE16CFB" bold="true" box="[189,304,460,486]" pageId="5" pageNumber="55">Diagnosis</emphasis>
: 
<taxonomicName id="4C7CA3DDFFA5403F4D1EFE7FFDF36CFB" box="[319,546,460,486]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="5" pageNumber="55" phylum="Basidiomycota" rank="species" species="lixivium">
<emphasis id="B908044CFFA5403F4D1EFE7FFDF36CFB" box="[319,546,460,486]" italics="true" pageId="5" pageNumber="55">Lyophyllum lixivium</emphasis>
</taxonomicName>
is characterized by having an umbonate pileus and abundant white floccus on the stipe base, stipe, and lamellae becoming dark brown when bruised, broadly globose basidiospores (4.4–6.5 × 3.7–5.8 µm), and presence a fusiform caulocystidia (
<figureCitation id="1347C4DBFFA5403F4E5AFDA7FD0F6F33" box="[635,734,532,558]" captionStart="FIGURE 2" captionStartId="5.[136,229,1542,1564]" captionTargetBox="[152,1436,583,1518]" captionTargetId="figure-121@5.[152,1436,583,1518]" captionTargetPageId="5" captionText="FIGURE 2. Lyophyllum lixivium. a Basidiomata (HKAS129929). b: Hymenophoral trama.c: Pileipellis.d: Stipitipellis (presence fusiform caulocystidia). Bars: a = 5 mm, b–f = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13214163" httpUri="https://zenodo.org/record/13214163/files/figure.png" pageId="5" pageNumber="55">Figure 2</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="8BC3D85EFFA5403F4CA9F9B5FDE86B5D" blockId="5.[136,1452,1542,1601]" pageId="5" pageNumber="55">
<emphasis id="B908044CFFA5403F4CA9F9B5FF2D6B01" bold="true" box="[136,252,1542,1564]" pageId="5" pageNumber="55">FIGURE 2.</emphasis>
<taxonomicName id="4C7CA3DDFFA5403F4D20F9B4FE126B01" box="[257,451,1543,1564]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="5" pageNumber="55" phylum="Basidiomycota" rank="species" species="lixivium">
<emphasis id="B908044CFFA5403F4D20F9B4FE126B01" box="[257,451,1543,1564]" italics="true" pageId="5" pageNumber="55">Lyophyllum lixivium</emphasis>
</taxonomicName>
. a Basidiomata (HKAS129929). b: Hymenophoral trama. c: Pileipellis. d: Stipitipellis (presence fusiform caulocystidia). Bars: a = 5 mm, b–f = 10 μm.
</paragraph>
</caption>
<subSubSection id="C3668BD5FFA5403C4C9CF9DCFBE26AB0" lastPageId="6" lastPageNumber="56" pageId="5" pageNumber="55" type="description">
<paragraph id="8BC3D85EFFA5403F4C9CF9DCFDC86A20" blockId="5.[136,1452,1647,2070]" pageId="5" pageNumber="55">
<emphasis id="B908044CFFA5403F4C9CF9DCFE966B94" bold="true" box="[189,327,1647,1673]" pageId="5" pageNumber="55">Description</emphasis>
: Pileus 
<quantity id="4C8475BBFFA5403F4D8BF9DCFDF66B97" box="[426,551,1647,1674]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="7.3" metricValueMin="2.1" pageId="5" pageNumber="55" unit="mm" value="47.0" valueMax="73.0" valueMin="21.0">21–73 mm</quantity>
diam, hemispherical to campanulate when young, becoming umbonate when mature, light brown (5D4–5D6), pileus margin inflexed. Context, 
<quantity id="4C8475BBFFA5403F4FA5F920FBDF6BB3" box="[900,1038,1683,1710]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.4" metricValueMin="1.6" pageId="5" pageNumber="55" unit="mm" value="2.5" valueMax="3.4" valueMin="1.6">1.6–3.4 mm</quantity>
wide, white, becoming light brown when injured. Lamellae adnate to slightly decurrent, white to creamy, 
<quantity id="4C8475BBFFA5403F4FBBF904FBF06BCF" box="[922,1057,1719,1746]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.2" metricValueMin="3.6" pageId="5" pageNumber="55" unit="mm" value="4.4" valueMax="5.2" valueMin="3.6">3.6–5.2 mm</quantity>
wide, with 1–4 unequal lamellulae between two entire, usually discoloring to dark at the edge when touched or injured. Stipe 91–115 × 
<quantity id="4C8475BBFFA5403F4909F968FA766BEB" box="[1320,1447,1755,1782]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="5" pageNumber="55" unit="mm" value="12.0" valueMax="14.0" valueMin="10.0">10–14 mm</quantity>
, greyish yellow (4C4–4C5), cylindrical slightly enlarged towards the base, centrally solid, abundant floccus on the base. Odor and taste not distinctive.
</paragraph>
<paragraph id="8BC3D85EFFA5403C4C9CF8F4FB0B6C1D" blockId="5.[136,1452,1647,2070]" lastBlockId="6.[136,1451,158,257]" lastPageId="6" lastPageNumber="56" pageId="5" pageNumber="55">
Basidiospores [80/2/2] (3.5–) 4.4–6.5 (–6.7) × (2.9–) 3.7–5.8 (–6.1) µm, av. 5.7 × 4.9 µm, Q = 1.0–1.4, Q 
<subScript id="17F8DA1BFFA5403F497FF8E9FABA6A75" attach="left" box="[1374,1387,1882,1896]" fontSize="6" pageId="5" pageNumber="55">m</subScript>
= 1.1 ± 0.09, mostly broadly lacrymoid or amygdaliform with acute apex in side view, ovoid in frontal view, occasionally fusiform, hyaline, thin-walled, always with oil drops. Spore deposit white (1A1). Basidia 26–38 × 7–10 µm, av. 32.1 × 9.4 µm, hyaline, clavate; sterigmata 2.4–4.6 × 1.3–2.5 µm, mostly 4–spored, rarely 2–spored. Hymenophoral trama is regular, hyaline hyphae, thin-walled, 4–15 µm wide. Cheilocystidia 14–36 × 2–6 µm, av. 20.6 × 4.8 µm, clavate. Pleurocystidia 16–33 × 3–5 µm, av. 21.3 × 4.2 µm, clavate. Pileipellis composed of 3.8–9.0 µm wide hyphae, cylindrical hyphae parallel or slightly crossed, smooth. Stipitipellis arranged regular, hyphae parallel, smooth, composed of 7–14 µm wide hyphae. Caulocystidia present on the stipe, 20–40 × 6–8 µm, av. 30.3 × 6.7 µm, fusiform, irregular, with subacute apex, thin-walled. Clamp connections present on the lamellae, pileipellis and stipitipellis.
</paragraph>
<caption id="DF0388D6FFA6403C4CA9F899FB5B6A79" ID-DOI="http://doi.org/10.5281/zenodo.13214165" ID-Zenodo-Dep="13214165" httpUri="https://zenodo.org/record/13214165/files/figure.png" pageId="6" pageNumber="56" startId="6.[136,229,1834,1856]" targetBox="[151,1436,281,1809]" targetPageId="6" targetType="figure">
<paragraph id="8BC3D85EFFA6403C4CA9F899FB5B6A79" blockId="6.[136,1451,1834,1892]" pageId="6" pageNumber="56">
<emphasis id="B908044CFFA6403C4CA9F899FED16A22" bold="true" box="[136,256,1834,1856]" pageId="6" pageNumber="56">FIGURE 3.</emphasis>
<taxonomicName id="4C7CA3DDFFA6403C4D29F899FE1F6A22" box="[264,462,1834,1855]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="6" pageNumber="56" phylum="Basidiomycota" rank="species" species="lixivium">
<emphasis id="B908044CFFA6403C4D29F899FE1F6A22" box="[264,462,1834,1855]" italics="true" pageId="6" pageNumber="56">Lyophyllum lixivium</emphasis>
</taxonomicName>
(HKAS129929). a: Cheilocystidia. b: Pleurocystidia. c: Basidia. d: Basidiospores as seen under the microscope. e: Clamp connections. f. Scanning electron microscope photo of basidiospores. Bars = 10 µm.
</paragraph>
</caption>
<paragraph id="8BC3D85EFFA6403C4C9CF821FBE26AB0" blockId="6.[136,1451,1938,2073]" box="[189,1075,1938,1965]" pageId="6" pageNumber="56">
<emphasis id="B908044CFFA6403C4C9CF821FE426AB1" bold="true" box="[189,403,1938,1965]" pageId="6" pageNumber="56">Habit and habitat</emphasis>
: Growing in moss-covered soil under 
<taxonomicName id="4C7CA3DDFFA6403C4F60F820FBFA6AB1" box="[833,1067,1938,1965]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="56" phylum="Tracheophyta" rank="species" species="mollissima">
<emphasis id="B908044CFFA6403C4F60F820FBFA6AB1" box="[833,1067,1938,1965]" italics="true" pageId="6" pageNumber="56">Castanea mollissima</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3668BD5FFA6403D4C9CF805FECA6F91" lastPageId="7" lastPageNumber="57" pageId="6" pageNumber="56" type="distribution">
<paragraph id="8BC3D85EFFA6403C4C9CF805FC206ACD" blockId="6.[136,1451,1938,2073]" box="[189,1009,1974,2001]" pageId="6" pageNumber="56">
<emphasis id="B908044CFFA6403C4C9CF805FE9E6ACD" bold="true" box="[189,335,1974,2000]" pageId="6" pageNumber="56">Distribution</emphasis>
: Found only in the 
<typeStatus id="54C766FCFFA6403C4E0DF804FD8D6ACC" box="[556,604,1975,2001]" pageId="6" pageNumber="56">type</typeStatus>
locality in 
<collectingRegion id="49B816BCFFA6403C4EFBF804FC4A6ACD" box="[730,923,1974,2001]" country="China" name="Yunnan" pageId="6" pageNumber="56">Yunnan province</collectingRegion>
, 
<collectingCountry id="F36B98CEFFA6403C4F87F805FC3C6ACD" box="[934,1005,1974,2000]" name="China" pageId="6" pageNumber="56">China</collectingCountry>
.
</paragraph>
<paragraph id="8BC3D85EFFA6403C4C9CF869FDFC6504" blockId="6.[136,1451,1938,2073]" pageId="6" pageNumber="56">
<emphasis id="B908044CFFA6403C4C9CF869FDCE6AE9" bold="true" box="[189,543,2010,2036]" pageId="6" pageNumber="56">Additional material examined</emphasis>
: 
<collectingCountry id="F36B98CEFFA6403C4E0DF869FDA36AE9" box="[556,626,2010,2036]" name="China" pageId="6" pageNumber="56">China</collectingCountry>
, 
<collectingRegion id="49B816BCFFA6403C4E5BF868FCEB6AE9" box="[634,826,2010,2037]" country="China" name="Yunnan" pageId="6" pageNumber="56">Yunnan Province</collectingRegion>
, Dali Cangshan Mountain, 
<date id="FFC2FE9EFFA6403C4844F868FAC46AE8" box="[1125,1301,2011,2037]" pageId="6" pageNumber="56" value="2020-08-11">11 August 2020</date>
, elev. 
<quantity id="4C8475BBFFA6403C4976F868FA766AE8" box="[1367,1447,2011,2037]" metricMagnitude="3" metricUnit="m" metricValue="2.371" pageId="6" pageNumber="56" unit="m" value="2371.0">2371m</quantity>
, S.H. Li 2020081139 (HKAS129930).
</paragraph>
<paragraph id="8BC3D85EFFA7403D4C9CFF2DFE756C55" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="57">
<emphasis id="B908044CFFA7403D4C9CFF2DFED16DA5" bold="true" box="[189,256,158,184]" pageId="7" pageNumber="57">Notes</emphasis>
: In multi-locus phylogenetic analyses, 
<taxonomicName id="4C7CA3DDFFA7403D4E8EFF2CFCF46DA5" box="[687,805,158,184]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="lixivium">
<emphasis id="B908044CFFA7403D4E8EFF2CFCF46DA5" box="[687,805,158,184]" italics="true" pageId="7" pageNumber="57">L. lixivium</emphasis>
</taxonomicName>
is related to 
<taxonomicName id="4C7CA3DDFFA7403D4F8FFF2CFBB66DA5" box="[942,1127,158,184]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="moncalvonum">
<emphasis id="B908044CFFA7403D4F8FFF2CFBB66DA5" box="[942,1127,158,184]" italics="true" pageId="7" pageNumber="57">L. moncalvonum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7CA3DDFFA7403D48BBFF2CFAE26DA5" box="[1178,1331,158,184]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="atrofuscum">
<emphasis id="B908044CFFA7403D48BBFF2CFAE26DA5" box="[1178,1331,158,184]" italics="true" pageId="7" pageNumber="57">L. atrofuscum</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="4C7CA3DDFFA7403D4CA9FF70FE936DC1" box="[136,322,194,220]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="moncalvonum">
<emphasis id="B908044CFFA7403D4CA9FF70FE936DC1" box="[136,322,194,220]" italics="true" pageId="7" pageNumber="57">L. moncalvonum</emphasis>
</taxonomicName>
originally described from 
<collectingCountry id="F36B98CEFFA7403D4E48FF70FCD16DC1" box="[617,768,194,221]" name="New Zealand" pageId="7" pageNumber="57">New Zealand</collectingCountry>
, has olivaceous black pileus and globose basidiospores (5.0 ± 0.5 μm; 
<bibRefCitation id="EFEDA5AFFFA7403D4CC2FF55FE646C1D" author="Cooper, J." box="[227,437,230,256]" pageId="7" pageNumber="57" pagination="263 - 288" refId="ref5062" refString="Cooper, J. (2014) New species and combinations of some New Zealand agarics belonging to Clitopilus, Lyophyllum, Gerhardtia, Clitocybe, Hydnangium, Mycena, Rhodocollydia and Gerronema. Mycosphere 5 (2): 263 - 288. https: // doi. org / 10.5943 / mycosphere / 5 / 2 / 2" type="journal article" year="2014">
Cooper 
<emphasis id="B908044CFFA7403D4D1DFF54FEA26C1D" box="[316,371,230,256]" italics="true" pageId="7" pageNumber="57">et al.</emphasis>
2014
</bibRefCitation>
). 
<taxonomicName id="4C7CA3DDFFA7403D4DE7FF55FD1C6C1D" box="[454,717,230,256]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="atrofuscum">
<emphasis id="B908044CFFA7403D4DE7FF55FD1C6C1D" box="[454,717,230,256]" italics="true" pageId="7" pageNumber="57">Lyophyllum atrofuscum</emphasis>
</taxonomicName>
has ellipsoid, subrhomboid, and irregular rhomboid basidiospores (5.5–7 × 3.9–5.0 µm), basidia mostly 2-spored, rarely 4-spored, absent cheilocystidia and caulocystidia on the stipe surface (
<bibRefCitation id="EFEDA5AFFFA7403D4CC6FE9DFE486C55" author="Wei, S. W. &amp; Lu, B. Y. &amp; Wang, Y. &amp; Dou, W. J. &amp; Wang, Q. &amp; Li, Y." box="[231,409,302,328]" pageId="7" pageNumber="57" refId="ref6927" refString="Wei, S. W., Lu, B. Y., Wang, Y., Dou, W. J., Wang, Q. &amp; Li, Y. (2023) Morphology and phylogeny of Lyophylloid mushroom in China with description of four new species. Journal of Fungi 9 (1): 77. https: // doi. org / 10.3390 / jof 9010077" type="journal volume" year="2023">
Wei 
<emphasis id="B908044CFFA7403D4D3CFE9CFE846C55" box="[285,341,302,328]" italics="true" pageId="7" pageNumber="57">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC3D85EFFA7403D4C9CFEE1FECA6F91" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="57">
<taxonomicName id="4C7CA3DDFFA7403D4C9CFEE1FE726C71" box="[189,419,338,364]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="lixivium">
<emphasis id="B908044CFFA7403D4C9CFEE1FE726C71" box="[189,419,338,364]" italics="true" pageId="7" pageNumber="57">Lyophyllum lixivium</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C7CA3DDFFA7403D4E19FEE0FD416C71" box="[568,656,338,364]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="bonii">
<emphasis id="B908044CFFA7403D4E19FEE0FD416C71" box="[568,656,338,364]" italics="true" pageId="7" pageNumber="57">L. bonii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7CA3DDFFA7403D4EBEFEE0FCBB6C71" box="[671,874,338,364]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="fuscobrunneum">
<emphasis id="B908044CFFA7403D4EBEFEE0FCBB6C71" box="[671,874,338,364]" italics="true" pageId="7" pageNumber="57">L. fuscobrunneum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7CA3DDFFA7403D4F58FEE0FBEA6C71" box="[889,1083,338,364]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="rhombisporum">
<emphasis id="B908044CFFA7403D4F58FEE0FBEA6C71" box="[889,1083,338,364]" italics="true" pageId="7" pageNumber="57">L. rhombisporum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7CA3DDFFA7403D485BFEE0FB2C6C71" box="[1146,1277,338,364]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="solidipes">
<emphasis id="B908044CFFA7403D485BFEE0FB2C6C71" box="[1146,1277,338,364]" italics="true" pageId="7" pageNumber="57">L. solidipes</emphasis>
</taxonomicName>
in context, and the stipe becomes black when bruised. The original description of 
<taxonomicName id="4C7CA3DDFFA7403D4F41FEC5FBF26C8D" box="[864,1059,374,400]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="bonii">
<emphasis id="B908044CFFA7403D4F41FEC5FBF26C8D" box="[864,1059,374,400]" italics="true" pageId="7" pageNumber="57">Lyophyllum bonii</emphasis>
</taxonomicName>
from the Canary Islands has a paler pileus with no umbo (
<bibRefCitation id="EFEDA5AFFFA7403D4D5EFE29FD4D6CA9" author="Consiglio, G. &amp; Contu, M." box="[383,668,410,436]" pageId="7" pageNumber="57" pagination="99 - 181" refId="ref5025" refString="Consiglio, G. &amp; Contu, M. (2002) II genere Lyophyllum P. Karst. emend. Kuhner, in Italia. River Micology 45 (2): 99 - 181." type="journal article" year="2002">Consiglio &amp; Contu 2002</bibRefCitation>
). According to its original description of 
<taxonomicName id="4C7CA3DDFFA7403D484CFE29FA746CA9" box="[1133,1445,410,436]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="fuscobrunneum">
<emphasis id="B908044CFFA7403D484CFE29FA746CA9" box="[1133,1445,410,436]" italics="true" pageId="7" pageNumber="57">Lyophyllum fuscobrunneum</emphasis>
</taxonomicName>
, which was described from the Canary Islands, has broadly ellipsoid to ellipsoid basidiospores (5.2–6.7 × 3.7–4.5 µm), relatively large basidia (30–50 × 7–9 µm), and repent to loosely interwoven pileipellis (
<bibRefCitation id="EFEDA5AFFFA7403D4892FE51FA716CE1" author="Dahncke, R. M. &amp; Contu, M. &amp; Vizzini, A." box="[1203,1440,482,508]" pageId="7" pageNumber="57" pagination="323 - 330" refId="ref5183" refString="Dahncke, R. M., Contu, M. &amp; Vizzini, A. (2011) Two new species of Lyophyllum s. l. (Basidiomycota, Agaricomycetes) from La Palma (Canary Islands, Spain). Mycotaxon 111: 323 - 330. https: // doi. org / 10.5248 / 111.323" type="journal article" year="2011">
Dähncke 
<emphasis id="B908044CFFA7403D4900FE50FA8A6CE1" box="[1313,1371,482,508]" italics="true" pageId="7" pageNumber="57">et al.</emphasis>
2011
</bibRefCitation>
). 
<taxonomicName id="4C7CA3DDFFA7403D4CA9FDB5FE686F3D" box="[136,441,518,544]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="rhombisporum">
<emphasis id="B908044CFFA7403D4CA9FDB5FE686F3D" box="[136,441,518,544]" italics="true" pageId="7" pageNumber="57">Lyophyllum rhombisporum</emphasis>
</taxonomicName>
has a relatively large basidia (36.5–46.0 × 10.0–11.3 μm) and basidiospores (14.5–17.0 × 10.0–11.5 μm; 
<bibRefCitation id="EFEDA5AFFFA7403D4D6DFD98FDC56F59" author="Wang, X. Q. &amp; Zhou, D. Q. &amp; Zhao, Y. C. &amp; Zhang, X. L. &amp; Li, L. &amp; Li, S. H." box="[332,532,554,581]" pageId="7" pageNumber="57" pagination="473 - 477" refId="ref6859" refString="Wang, X. Q., Zhou, D. Q., Zhao, Y. C., Zhang, X. L., Li, L. &amp; Li, S. H. (2013) Lyophyllum rhombisporum sp. nov. from China. Mycotaxon 123: 473 - 477. https: // doi. org / 10.5248 / 123.473" type="journal article" year="2013">
Wang 
<emphasis id="B908044CFFA7403D4DB4FD98FE1E6F59" box="[405,463,554,580]" italics="true" pageId="7" pageNumber="57">et al.</emphasis>
2013
</bibRefCitation>
). 
<taxonomicName id="4C7CA3DDFFA7403D4E09FD99FCC66F59" box="[552,791,554,580]" class="Agaricomycetes" family="Lyophyllaceae" genus="Lyophyllum" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="57" phylum="Basidiomycota" rank="species" species="solidipes">
<emphasis id="B908044CFFA7403D4E09FD99FCC66F59" box="[552,791,554,580]" italics="true" pageId="7" pageNumber="57">Lyophyllum solidipes</emphasis>
</taxonomicName>
collected from Hood National Forest of the 
<collectingCountry id="F36B98CEFFA7403D4935FD99FA7A6F59" box="[1300,1451,554,580]" name="United States of America" pageId="7" pageNumber="57">United States</collectingCountry>
has narrower hyphae of stipitipellis (3–5 μm) and relatively large basidiospores (6.5–8.1 × 3.7–4.7 μm; 
<bibRefCitation id="EFEDA5AFFFA7403D492EFDFDFEDF6F91" author="Clemencon, H. &amp; Smith, A." pageId="7" pageNumber="57" pagination="379 - 437" refId="ref4987" refString="Clemencon, H. &amp; Smith, A. (1983) New species of Lyophyllum (Agaricales) from North America and a key to the known staining species. Mycotaxon 18: 379 - 437." type="journal article" year="1983">Clémençon &amp; Smith 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>