treatments-xml/data/03/CC/87/03CC87C43D33520958A4FC693CD5FBE9.xml

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<mods:title id="358ACCDDA8405DB8F81616E1CAAB2889">New species of Ameletus Eaton, 1885 and redescription of Ameletus longulus Sinichenkova, 1981 from the Russian Far East (Ephemeroptera: Ameletidae)</mods:title>
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<subSubSection id="C37F65593D33520158A4FC693A1AFB7B" pageId="5" pageNumber="524" type="nomenclature">
<paragraph id="8BDA36D23D33520158A4FC693C03FBDA" blockId="5.[151,624,1005,1063]" box="[151,624,1005,1031]" pageId="5" pageNumber="524">
<heading id="D09281BE3D33520158A4FC693C03FBDA" bold="true" box="[151,624,1005,1031]" fontSize="11" level="1" pageId="5" pageNumber="524" reason="1">
<taxonomicName id="4C654D513D33520158A4FC693C03FBDA" authority="Sinichenkova, 1981" authorityName="Sinichenkova" authorityYear="1981" box="[151,624,1005,1031]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D33520158A4FC693C03FBDA" bold="true" box="[151,624,1005,1031]" pageId="5" pageNumber="524">
<emphasis id="B911EAC03D33520158A4FC693F0BFBDA" bold="true" box="[151,376,1005,1031]" italics="true" pageId="5" pageNumber="524">Ameletus longulus</emphasis>
Sinichenkova, 1981
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDA36D23D33520158A4FB8B3F2BFBFA" blockId="5.[151,624,1005,1063]" box="[151,344,1039,1063]" pageId="5" pageNumber="524">
(
<figureCitation id="135E2A573D33520158ACFB8B3E8EFBFA" box="[159,253,1039,1063]" captionStart="FIGURES 21 – 23" captionStartId="3.[151,264,1977,2000]" captionTargetBox="[165,1422,733,1939]" captionTargetId="figure@3.[151,1436,713,1955]" captionTargetPageId="3" captionText="FIGURES 21 – 23. Comb-shaped setae of maxilla: 21. Ameletus khasanensis sp. nov., 22. Ameletus longulus, 23. Ameletus costalis. The arrow shows the first seta." httpUri="https://zenodo.org/record/223957/files/figure.png" pageId="5" pageNumber="524">Figs. 22</figureCitation>
, 
<figureCitation id="135E2A573D335201593BFB8B3F23FBFA" box="[264,336,1039,1063]" captionStart-0="FIGURES 27 – 35" captionStart-1="FIGURES 36 – 46" captionStart-2="FIGURES 47 – 49" captionStartId-0="6.[151,264,1912,1935]" captionStartId-1="7.[151,264,1700,1723]" captionStartId-2="8.[151,264,1607,1630]" captionTargetBox-0="[168,1414,324,1899]" captionTargetBox-1="[159,1415,193,1649]" captionTargetBox-2="[166,1421,301,1585]" captionTargetId-0="figure@6.[161,1426,314,1902]" captionTargetId-1="figure@7.[159,1427,193,1679]" captionTargetId-2="figure@8.[151,1436,301,1586]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" captionText-1="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." captionText-2="FIGURES 47 – 49. Ameletus longulus, egg: 47. General view. 48. Structure of chorion and small knob-terminated coiled threads. 49. Large knobs. The arrow shows the micropiles (mp)." httpUri-0="https://zenodo.org/record/223959/files/figure.png" httpUri-1="https://zenodo.org/record/223960/files/figure.png" httpUri-2="https://zenodo.org/record/223961/files/figure.png" pageId="5" pageNumber="524">27–49</figureCitation>
)
</paragraph>
<paragraph id="8BDA36D23D33520158A4FBD63A1AFB7B" blockId="5.[151,1435,1106,1190]" pageId="5" pageNumber="524">
Sinitshenkova &amp; Tshernova 1976: 16, figs. 10–17 (
<taxonomicName id="4C654D513D3352015AACFBD63D3CFBB5" box="[671,847,1106,1128]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3352015AACFBD63D3CFBB5" box="[671,847,1106,1128]" italics="true" pageId="5" pageNumber="524">Ameletus costalis</emphasis>
</taxonomicName>
); Sinitshenkova 1977: 78, fig. 3, l–m (
<taxonomicName id="4C654D513D3352015CEFFBD63BFFFBB5" box="[1244,1420,1106,1128]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3352015CEFFBD63BFFFBB5" box="[1244,1420,1106,1128]" italics="true" pageId="5" pageNumber="524">Ameletus costalis</emphasis>
</taxonomicName>
); Sinichenkova 1981: 77–78, fig. 3 (
<taxonomicName id="4C654D513D3352015A0EFBF53C88FB5A" box="[573,763,1137,1159]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3352015A0EFBF53C88FB5A" box="[573,763,1137,1159]" italics="true" pageId="5" pageNumber="524">Ameletus longulus</emphasis>
</taxonomicName>
); Tshernova 
<emphasis id="B911EAC03D3352015BB9FBF63DB2FB5A" box="[906,961,1137,1159]" italics="true" pageId="5" pageNumber="524">et al.</emphasis>
1986: 126, figs. 57:1–2 (
<taxonomicName id="4C654D513D3352015CE4FBF53BFFFB5A" box="[1239,1420,1137,1159]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3352015CE4FBF53BFFFB5A" box="[1239,1420,1137,1159]" italics="true" pageId="5" pageNumber="524">Ameletus costalis</emphasis>
</taxonomicName>
); Tiunova &amp; Potikha 2005 (
<taxonomicName id="4C654D513D33520159E3FB143CF3FB7B" box="[464,640,1168,1190]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D33520159E3FB143CF3FB7B" box="[464,640,1168,1190]" italics="true" pageId="5" pageNumber="524">Ameletus costalis</emphasis>
</taxonomicName>
); Kluge 2007:257 (
<taxonomicName id="4C654D513D3352015B7BFB143D8BFB7B" box="[840,1016,1168,1190]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3352015B7BFB143D8BFB7B" box="[840,1016,1168,1190]" italics="true" pageId="5" pageNumber="524">Ameletus costalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3352015BCCFB153A2DFB7B" box="[1023,1118,1169,1190]" pageId="5" pageNumber="524">syn. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D33520158A4FB493C42F9FF" pageId="5" pageNumber="524" type="materials_examined">
<paragraph id="8BDA36D23D33520158A4FB493B46FAD5" blockId="5.[151,1437,1229,2027]" pageId="5" pageNumber="524">
<emphasis id="B911EAC03D33520158A4FB493F08FB3B" bold="true" box="[151,379,1229,1254]" pageId="5" pageNumber="524">Material examined</emphasis>
. 
<typeStatus id="54DE88703D33520159B8FB4A3F9EFB3B" box="[395,493,1230,1254]" pageId="5" pageNumber="524" type="paratype">Paratype</typeStatus>
larvae (No 2971a, collected together with the 
<typeStatus id="54DE88703D3352015C37FB493A19FB38" box="[1028,1130,1229,1253]" pageId="5" pageNumber="524" type="holotype">holotype</typeStatus>
), 
<collectingCountry id="F37276423D3352015C4CFB493ABEFB38" box="[1151,1229,1229,1253]" name="Russia" pageId="5" pageNumber="524">Russia</collectingCountry>
, Primorskiy Kray, Terneiskii region, Sikhote-Alin State Nature Biosphere Reserve, Poltkov Stream, 
<date id="FFDB10123D3352015C14FB743AD9FAD5" box="[1063,1194,1264,1288]" pageId="5" pageNumber="524" value="1974-06-17">17.VI.1974</date>
, Nadezdina.
</paragraph>
<paragraph id="8BDA36D23D33520158F4FA973C42F9FF" blockId="5.[151,1437,1229,2027]" pageId="5" pageNumber="524">
<emphasis id="B911EAC03D33520158F4FA973F8CFAF1" bold="true" box="[199,511,1299,1324]" pageId="5" pageNumber="524">Other material examined.</emphasis>
<collectingCountry id="F37276423D3352015A34FA973C26FAF6" box="[519,597,1299,1323]" name="Russia" pageId="5" pageNumber="524">Russia</collectingCountry>
, Primorskiy Kray, Terneiskii region, Sikhote-Alin State Nature Biosphere Reserve: Mashinukovskiy Stream, 
<date id="FFDB10123D3352015A02FAB33CB2FA92" box="[561,705,1335,1359]" pageId="5" pageNumber="524" value="1995-06-04" valueMax="1995-06-06" valueMin="1995-06-04">4–6.VI.1995</date>
, E Poticha, 
<specimenCount id="9D63FD5B3D3352015B69FAB33DDFFA92" box="[858,940,1335,1359]" pageId="5" pageNumber="524" type="male">4 male</specimenCount>
imagines; Zolotoi Stream, 
<date id="FFDB10123D3352015CC4FAB33B0AFA92" box="[1271,1401,1335,1359]" pageId="5" pageNumber="524" value="1995-06-21">21.VI.1995</date>
, E Poticha, 
<specimenCount id="9D63FD5B3D33520158CAFADE3F37FAAF" box="[249,324,1370,1394]" pageId="5" pageNumber="524" type="male">1 male</specimenCount>
subimago (reared), 
<specimenCount id="9D63FD5B3D3352015A15FADE3CFBFAAF" box="[550,648,1370,1394]" pageId="5" pageNumber="524" type="female">2 female</specimenCount>
imagines; Serebrynka River, 
<date id="FFDB10123D3352015BE6FADE3A25FAAF" box="[981,1110,1370,1394]" pageId="5" pageNumber="524" value="2002-06-11">11.VI.2002</date>
, E Poticha, 
<specimenCount id="9D63FD5B3D3352015CEFFADE3B54FAAF" box="[1244,1319,1370,1394]" pageId="5" pageNumber="524" type="male">3 male</specimenCount>
imagines; Lazovskiy Nature Reserve, Proselochnaya Bay, Proselochniy Stream, 
<date id="FFDB10123D3352015BF9FAF93A4DFA48" box="[970,1086,1405,1429]" pageId="5" pageNumber="524" value="2005-06-09">9.VI.2005</date>
, Y Sundukov, 
<specimenCount id="9D63FD5B3D3352015CC1FAF93B31FA48" box="[1266,1346,1405,1429]" pageId="5" pageNumber="524" type="male">1 male</specimenCount>
imago; 
<emphasis id="B911EAC03D33520158A4FA243FF6FA64" bold="true" box="[151,389,1440,1465]" pageId="5" pageNumber="524">Khabarovskiy Kray</emphasis>
, Ulchskii region, Severnaya Bay, Tatarka River, about 
<quantity id="4C9D9B373D3352015C30FA243A3EFA65" box="[1027,1101,1440,1464]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="524" unit="m" value="200.0">200 m</quantity>
above mouth, 
<date id="FFDB10123D3352015CC9FA243B0FFA65" box="[1274,1404,1440,1464]" pageId="5" pageNumber="524" value="2005-06-19">19.VI.2005</date>
, T Tiunova, 
<specimenCount id="9D63FD5B3D33520158CCFA473F35FA06" box="[255,326,1475,1499]" pageId="5" pageNumber="524" type="male">1 male</specimenCount>
imago (reared); Tatarka River, about 
<quantity id="4C9D9B373D3352015AC1FA473D4AFA06" box="[754,825,1475,1499]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="524" unit="m" value="100.0">100 m</quantity>
above mouth, 
<date id="FFDB10123D3352015BD1FA473A17FA06" box="[994,1124,1475,1499]" pageId="5" pageNumber="524" value="2007-06-22">22.VI.2007</date>
, I Tiunov, 
<specimenCount id="9D63FD5B3D3352015CECFA473B5EFA06" box="[1247,1325,1475,1499]" pageId="5" pageNumber="524" type="male">1 male</specimenCount>
, 
<specimenCount id="9D63FD5B3D3352015D09FA473BEFFA06" box="[1338,1436,1475,1499]" pageId="5" pageNumber="524" type="female">2 female</specimenCount>
imagines, 
<specimenCount id="9D63FD5B3D3352015938FA633F1AFA22" box="[267,361,1511,1535]" pageId="5" pageNumber="524" type="larva">3 larvae</specimenCount>
; same place, 
<date id="FFDB10123D3352015A31FA633CF0FA22" box="[514,643,1511,1535]" pageId="5" pageNumber="524" value="2005-06-19">19.VI.2005</date>
, I Tiunov, 8 mature larvae and 2 larval exuvia; Bolshoi Somon River, 
<date id="FFDB10123D33520158A4F98E3F6AF9FF" box="[151,281,1546,1570]" pageId="5" pageNumber="524" value="2008-06-22">22.VI.2008</date>
, I Tiunov, 
<specimenCount id="9D63FD5B3D33520159A2F98E3FAFF9FF" box="[401,476,1546,1570]" pageId="5" pageNumber="524" type="male">1 male</specimenCount>
imago.
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D33520D58F4F9A93A8FFD11" lastPageId="9" lastPageNumber="528" pageId="5" pageNumber="524" type="description">
<paragraph id="8BDA36D23D33520258F4F9A93D49FEC4" blockId="5.[151,1437,1229,2027]" lastBlockId="6.[151,1436,151,281]" lastPageId="6" lastPageNumber="525" pageId="5" pageNumber="524">
<emphasis id="B911EAC03D33520158F4F9A93F22F99B" bold="true" box="[199,337,1581,1606]" pageId="5" pageNumber="524">Description</emphasis>
. 
<emphasis id="B911EAC03D3352015952F9A93F9FF99B" bold="true" box="[353,492,1581,1606]" italics="true" pageId="5" pageNumber="524">Male imago</emphasis>
(in alcohol). Length (mm): body 14.0–16.8; forewings 13.8–15.3; cerci 20.0–23.0. 
<emphasis id="B911EAC03D33520158A4F9D53EA4F9B4" bold="true" box="[151,215,1617,1641]" pageId="5" pageNumber="524">Head</emphasis>
: upper portion of eyes gray; lower portion dark gray. Medial and lateral ocelli brown. Antennae brown. 
<emphasis id="B911EAC03D33520158A4F9F03E83F951" bold="true" box="[151,240,1652,1676]" pageId="5" pageNumber="524">Thorax</emphasis>
: Medioscutum and submedioscutum light brown; medial longitudinal suture narrow, brown. Sublateroscutum dark brown. Prelateroscutum yellowish. Medioparapsidal suture brown. Scutellum light brown; scuto-scutellar impression pale. Posterior scutal protuberance light brown or dirty yellowish; parascutellum dark (
<figureCitation id="135E2A573D33520158ACF9593E85F928" box="[159,246,1757,1781]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="5" pageNumber="524">Fig. 27</figureCitation>
). Forelegs brown; middle and hind legs yellowish; tarsal segments brownish; dorsal surface of the fore tibia without spinules. Length (mm) of foreleg segments: femora 2.8–3.0; tibia 2.7–2.9; tarsal segments 0.7–1.1, 1.7–1.9, 1.5–1.6, 1.1–1.2, and 0.5–0.6. Wings brownish, hyaline; all veins brown (
<figureCitation id="135E2A573D3352015C6CF8A73A9FF8E6" box="[1119,1260,1827,1851]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="5" pageNumber="524">Figs. 28–29</figureCitation>
), crossveins of forewings between C and R veins brown, more contrasting than others. Abdomen: terga brownish, opaque; lateral and posterior margins brown; tergum I dark brown; terga II–IX in middle portion with pair of brown oblique stripes along the length; tergum IX–X darker. Sterna whitish or brownish; sternum I brown; sterna II–VII in the middle closer to the anterior margin with a dark triangular spot and two brownish oblique stripes; sternum IX dirty brown with darkish lateral margins. Styliger light brown, lateral sides darker; forceps dark brown (
<figureCitation id="135E2A573D3352015C83F8573B75F836" box="[1200,1286,2003,2027]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="5" pageNumber="524">Fig. 30</figureCitation>
). Penis lobes light brown on inner sides and brown or dark brown on external sides; lateral penis lobes with rounded and divergent tips (
<figureCitation id="135E2A573D305202590DFF3F3FB4FF0E" box="[318,455,187,211]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="6" pageNumber="525">Figs. 32–33</figureCitation>
); lateral penis lobes densely covered with small spines; each ventral plate bears a single long and pointed denticle (
<figureCitation id="135E2A573D305202598CFF5A3C34FF2B" box="[447,583,222,246]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="6" pageNumber="525">Figs. 31, 34</figureCitation>
); distance between base of the lateral penis lobe and denticle ¼ of maximum width of denticle (
<figureCitation id="135E2A573D3052025957FE853FCBFEC4" box="[356,440,257,281]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="6" pageNumber="525">Fig. 34</figureCitation>
). Cerci brown with yellowish tips.
</paragraph>
<caption id="DF1A665A3D30520258A4F8FC3DABF811" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="6" pageNumber="525" targetBox="[168,1414,324,1899]" targetPageId="6">
<paragraph id="8BDA36D23D30520258A4F8FC3DABF811" blockId="6.[151,1436,1912,1997]" pageId="6" pageNumber="525">
<emphasis id="B911EAC03D30520258A4F8FC3F3DF853" bold="true" box="[151,334,1912,1935]" pageId="6" pageNumber="525">FIGURES 27–35</emphasis>
. 
<taxonomicName id="4C654D513D305202596FF8FD3C6BF852" box="[348,536,1913,1935]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D305202596FF8FD3C6BF852" box="[348,536,1913,1935]" italics="true" pageId="6" pageNumber="525">Ameletus longulus</emphasis>
</taxonomicName>
, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30–35=0.5; 28–29=2.0; 27=1.0
</paragraph>
</caption>
<caption id="DF1A665A3D31520358A4F9203B68F925" httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="7" pageNumber="526" targetBox="[159,1415,193,1649]" targetPageId="7">
<paragraph id="8BDA36D23D31520358A4F9203B68F925" blockId="7.[151,1437,1700,1784]" pageId="7" pageNumber="526">
<emphasis id="B911EAC03D31520358A4F9203F21F967" bold="true" box="[151,338,1700,1723]" pageId="7" pageNumber="526">FIGURES 36–46</emphasis>
. 
<taxonomicName id="4C654D513D3152035950F9213C52F966" box="[355,545,1701,1723]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="526" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3152035950F9213C52F966" box="[355,545,1701,1723]" italics="true" pageId="7" pageNumber="526">Ameletus longulus</emphasis>
</taxonomicName>
, larva: 36. Labrum, dorsal view. 37–38. Apices of mandibles, dorsal view: 37–left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41–46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41–46=0.5; 37–38=0.2. The arrow shows the group of setae.
</paragraph>
</caption>
<paragraph id="8BDA36D23D31520358F4F8B93F48F83B" blockId="7.[151,1437,1853,2022]" pageId="7" pageNumber="526">
<emphasis id="B911EAC03D31520358F4F8B93F03F88B" bold="true" box="[199,368,1853,1878]" italics="true" pageId="7" pageNumber="526">Female imago</emphasis>
. Length (mm): body 16.4–20.7; forewings 16.6–18.0; cerci 24.0. Overall color light brown. 
<emphasis id="B911EAC03D31520358A4F8E63E83F8A7" bold="true" box="[151,240,1890,1914]" pageId="7" pageNumber="526">Thorax</emphasis>
: Mesonotum brown. Forelegs brown; middle and hind legs light brown, tarsal segments brown, joints darker. Color of wings as in male. 
<emphasis id="B911EAC03D3152035A2BF8013CFFF843" bold="true" box="[536,652,1925,1950]" pageId="7" pageNumber="526">Abdomen</emphasis>
: tergum I dark brown; terga II–IX brown (abdomen with eggs). Color of sterna same as terga. Subgenital plate with deep depression (
<figureCitation id="135E2A573D3152035B4AF82E3DBCF81F" box="[889,975,1962,1986]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="7" pageNumber="526">Fig. 35</figureCitation>
). Cerci dark brown with light brown or yellowish tips.
</paragraph>
<paragraph id="8BDA36D23D3E520C58F4FF133C88FF2A" blockId="8.[151,1436,151,247]" pageId="8" pageNumber="527">
<emphasis id="B911EAC03D3E520C58F4FF133F09FF6D" bold="true" box="[199,378,151,176]" italics="true" pageId="8" pageNumber="527">Male subimago</emphasis>
. Overall color brown. Wings whitish and translucent. All crossveins of fore- and hind wings bordered by brown. Terga brown, monotonous; terga VII–X darker. Sterna brown. Forelegs brown; middle and hind legs lighter; joints dark brown. Cerci dark brown.
</paragraph>
<caption id="DF1A665A3D3E520C58A4F9C33D53F9A0" httpUri="https://zenodo.org/record/223961/files/figure.png" pageId="8" pageNumber="527" targetBox="[166,1421,301,1585]" targetPageId="8">
<paragraph id="8BDA36D23D3E520C58A4F9C33D53F9A0" blockId="8.[151,1436,1607,1661]" pageId="8" pageNumber="527">
<emphasis id="B911EAC03D3E520C58A4F9C33F2AF980" bold="true" box="[151,345,1607,1630]" pageId="8" pageNumber="527">FIGURES 47–49.</emphasis>
<taxonomicName id="4C654D513D3E520C5950F9C33C51F980" box="[355,546,1607,1629]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="527" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3E520C5950F9C33C51F980" box="[355,546,1607,1629]" italics="true" pageId="8" pageNumber="527">Ameletus longulus</emphasis>
</taxonomicName>
, egg: 47. General view. 48. Structure of chorion and small knob-terminated coiled threads. 49. Large knobs. The arrow shows the micropiles (mp).
</paragraph>
</caption>
<paragraph id="8BDA36D23D3E520D58F4F92E3C17FDCA" blockId="8.[151,1437,1705,2017]" lastBlockId="9.[151,1436,151,1040]" lastPageId="9" lastPageNumber="528" pageId="8" pageNumber="527">
<emphasis id="B911EAC03D3E520C58F4F92E3F13F91F" bold="true" box="[199,352,1705,1730]" italics="true" pageId="8" pageNumber="527">Mature larva</emphasis>
. Length (mm): body 13.5–19.0; cerci 6.0–7.5. 
<emphasis id="B911EAC03D3E520C5B49F92D3DC9F91C" bold="true" box="[890,954,1705,1729]" pageId="8" pageNumber="527">Head</emphasis>
: dark brown. Antennae light brown, basal segments brown. Labrum (length 0.8–0.93 of width) with a deep and wide apical incision (
<figureCitation id="135E2A573D3E520C5CAAF9493A9DF938" box="[1177,1262,1741,1765]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="8" pageNumber="527">Fig. 36</figureCitation>
). Incisor of left mandible with four denticles (
<figureCitation id="135E2A573D3E520C59C2F9753C3BF8D4" box="[497,584,1777,1801]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="8" pageNumber="527">Fig. 37</figureCitation>
). Incisor of right mandible with three denticles, the third somewhat larger than the first (
<figureCitation id="135E2A573D3E520C5972F8913FE9F8F0" box="[321,410,1813,1837]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="8" pageNumber="527">Fig. 38</figureCitation>
). Labium dirty whitish; paraglossae length exceeds glossae length; distal margin of the glossae and part of the paraglossae brownish; third segments of labial palpi brownish (
<figureCitation id="135E2A573D3E520C5C57F8BD3ACAF88C" box="[1124,1209,1849,1873]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="8" pageNumber="527">Fig. 39</figureCitation>
). First comb-shaped seta of maxilla with narrow and pointed denticles, their number more than ten (
<figureCitation id="135E2A573D3E520C5C1BF8D93A0CF8A8" box="[1064,1151,1885,1909]" captionStart="FIGURES 21 – 23" captionStartId="3.[151,264,1977,2000]" captionTargetBox="[165,1422,733,1939]" captionTargetId="figure@3.[151,1436,713,1955]" captionTargetPageId="3" captionText="FIGURES 21 – 23. Comb-shaped setae of maxilla: 21. Ameletus khasanensis sp. nov., 22. Ameletus longulus, 23. Ameletus costalis. The arrow shows the first seta." httpUri="https://zenodo.org/record/223957/files/figure.png" pageId="8" pageNumber="527">Fig. 22</figureCitation>
). 
<emphasis id="B911EAC03D3E520C5CA7F8D93A9FF8A8" bold="true" box="[1172,1260,1885,1909]" pageId="8" pageNumber="527">Thorax</emphasis>
: pronotum and mesonotum brown with pale median stripe. Pronotum with light oblique stripes on the sides; lateral margins darker. Legs brown; forelegs darker than middle and hind legs; distal part of femur of all legs light brown; a group of six or seven setae on the dorsal surface at the base of fore femora (
<figureCitation id="135E2A573D3E520C5B60F84D3DDAF83C" box="[851,937,1993,2017]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="8" pageNumber="527">Fig. 40</figureCitation>
). Foreleg: 2.1–2.3; tibia 1.2–1.3; and tarsus 1.4–1.5. Middle leg: femur 2.0–2.3; tibia 1.1–1.3; and tarsus 1.1–1.3. 
<collectingCountry id="F37276423D3F520D5B81FF133D99FF72" box="[946,1002,151,175]" name="India" pageId="9" pageNumber="528">Hind</collectingCountry>
leg: femur 2.1–2.5; tibia 1.1–1.3; and tarsus 1.1–1.4. 
<emphasis id="B911EAC03D3F520D5971FF3F3FC5FF09" bold="true" box="[322,438,187,212]" pageId="9" pageNumber="528">Abdomen</emphasis>
: tergum I dark brown; antero-lateral corners pale; terga II–IX with dark brown posterior margins and light brown lateral corners; maculation on terga II–IX not distinct in the form of paired brown and light stripes; tergum X brown; all terga with almost regular row of spines, which alternate between large and small. Sterna dark brown; sternum I pale; sterna II–IX with a pair of small white spots on antero-lateral corners; sterna VI–VIII with a well-defined ganglionic marking. Gills white or pale; gill I and II small, rounded in distal part; gill I with small anal rib, width 0.57 of length (
<figureCitation id="135E2A573D3F520D5A4AFEEB3CBFFE5A" box="[633,716,367,391]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="9" pageNumber="528">Fig. 41</figureCitation>
); gill II with anal rib, width 0.60 of length (
<figureCitation id="135E2A573D3F520D5C86FEEB3B7AFE5A" box="[1205,1289,367,391]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="9" pageNumber="528">Fig. 42</figureCitation>
); gills III–VI with anal rib far from anal margin; gills III–V with rounded distal margin, width 0.58 of length (
<figureCitation id="135E2A573D3F520D5CEBFE103B10FE71" box="[1240,1379,404,428]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="9" pageNumber="528">Figs. 43–44</figureCitation>
); gill VI with straight distal margin, width 0.55 of length (
<figureCitation id="135E2A573D3F520D5ADCFE333D36FE12" box="[751,837,439,463]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="9" pageNumber="528">Fig. 45</figureCitation>
); gill VII with a truncated inner distal margin, width 0.46 of length (
<figureCitation id="135E2A573D3F520D5978FE583FD1FE29" box="[331,418,476,500]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="9" pageNumber="528">Fig. 46</figureCitation>
). The basal quarter of the cerci is dark brown, more distally, brown, tips light brown or yellowish; joints of each segment lighter.
</paragraph>
<paragraph id="8BDA36D23D3F520D58F4FDA03A8FFD11" blockId="9.[151,1436,151,1040]" pageId="9" pageNumber="528">
<emphasis id="B911EAC03D3F520D58F4FDA03F7BFDE6" bold="true" box="[199,264,548,571]" italics="true" pageId="9" pageNumber="528">Eggs.</emphasis>
General form of egg flattened ellipse with a length of 282–293 μm and a width of 176–187 μm (
<figureCitation id="135E2A573D3F520D5D0FFDA03BE3FDE1" box="[1340,1424,548,572]" captionStart="FIGURES 47 – 49" captionStartId="8.[151,264,1607,1630]" captionTargetBox="[166,1421,301,1585]" captionTargetId="figure@8.[151,1436,301,1586]" captionTargetPageId="8" captionText="FIGURES 47 – 49. Ameletus longulus, egg: 47. General view. 48. Structure of chorion and small knob-terminated coiled threads. 49. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223961/files/figure.png" pageId="9" pageNumber="528">Fig. 47</figureCitation>
). Chorionic surface covered with large-mesh regular hexagonal cells; each cell with ribbed bottom; borders of cells narrow and very low; a few coiled threads, terminating in a small round knob, located in the middle of their borders, bordering higher hexagonal cells (
<figureCitation id="135E2A573D3F520D5AB0FD0B3CABFD7A" box="[643,728,655,679]" captionStart="FIGURES 47 – 49" captionStartId="8.[151,264,1607,1630]" captionTargetBox="[166,1421,301,1585]" captionTargetId="figure@8.[151,1436,301,1586]" captionTargetPageId="8" captionText="FIGURES 47 – 49. Ameletus longulus, egg: 47. General view. 48. Structure of chorion and small knob-terminated coiled threads. 49. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223961/files/figure.png" pageId="9" pageNumber="528">Fig. 48</figureCitation>
); coiled threads terminated by a large round knob, located on one pole only (
<figureCitation id="135E2A573D3F520D590DFD303FE0FD11" box="[318,403,692,716]" captionStart="FIGURES 47 – 49" captionStartId="8.[151,264,1607,1630]" captionTargetBox="[166,1421,301,1585]" captionTargetId="figure@8.[151,1436,301,1586]" captionTargetPageId="8" captionText="FIGURES 47 – 49. Ameletus longulus, egg: 47. General view. 48. Structure of chorion and small knob-terminated coiled threads. 49. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223961/files/figure.png" pageId="9" pageNumber="528">Fig. 49</figureCitation>
). Micropyles observed in the subequatorial area; sperm guide round (
<figureCitation id="135E2A573D3F520D5CA8FD303A83FD11" box="[1179,1264,692,716]" captionStart="FIGURES 47 – 49" captionStartId="8.[151,264,1607,1630]" captionTargetBox="[166,1421,301,1585]" captionTargetId="figure@8.[151,1436,301,1586]" captionTargetPageId="8" captionText="FIGURES 47 – 49. Ameletus longulus, egg: 47. General view. 48. Structure of chorion and small knob-terminated coiled threads. 49. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223961/files/figure.png" pageId="9" pageNumber="528">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D3F520D58F4FD533C31FCC9" pageId="9" pageNumber="528" type="distribution">
<paragraph id="8BDA36D23D3F520D58F4FD533C31FCC9" blockId="9.[151,1436,151,1040]" pageId="9" pageNumber="528">
<emphasis id="B911EAC03D3F520D58F4FD533F98FD2D" bold="true" box="[199,491,727,752]" pageId="9" pageNumber="528">Distribution and biology</emphasis>
. Russian Far East: Primorskiy and Khabarovskiy regions. Mature larvae and adults were collected in small foothill rivers.
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D3F520D58F4FCA43CBAFAFE" pageId="9" pageNumber="528" type="discussion">
<paragraph id="8BDA36D23D3F520D58F4FCA43C5AFBCD" blockId="9.[151,1436,151,1040]" pageId="9" pageNumber="528">
<emphasis id="B911EAC03D3F520D58F4FCA43F48FCE5" bold="true" box="[199,315,800,824]" pageId="9" pageNumber="528">Remarks.</emphasis>
<taxonomicName id="4C654D513D3F520D5971FCA43FB1FCE5" box="[322,450,800,824]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3F520D5971FCA43FB1FCE5" box="[322,450,800,824]" italics="true" pageId="9" pageNumber="528">A. longulus</emphasis>
</taxonomicName>
was described based on young larvae, which, like all young larvae, have gills narrower than in mature larvae (Sinichenkova 1981). Kluge (2007) reviewed the genus 
<taxonomicName id="4C654D513D3F520D5BC9FCC03A13FC81" box="[1018,1120,836,860]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAC03D3F520D5BC9FCC03A13FC81" box="[1018,1120,836,860]" italics="true" pageId="9" pageNumber="528">Ameletus</emphasis>
</taxonomicName>
, and considered 
<taxonomicName id="4C654D513D3F520D5D2EFCC03BEFFC81" box="[1309,1436,836,860]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3F520D5D2EFCC03BEFFC81" box="[1309,1436,836,860]" italics="true" pageId="9" pageNumber="528">A. longulus</emphasis>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="4C654D513D3F520D599DFCEC3CBFFCA2" authority="Kluge 2007" authorityName="Kluge" authorityYear="2007" box="[430,716,871,896]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D599DFCEC3C56FC5D" box="[430,549,872,896]" italics="true" pageId="9" pageNumber="528">A. costalis</emphasis>
(Kluge 2007)
</taxonomicName>
. Recently, we collected adequate new material associating the male, female, eggs, and larvae. The identity of our material was confirmed by comparison with photos of the denticles in the mandible and maxilla the 
<typeStatus id="54DE88703D3F520D5A48FC353CA8FC14" box="[635,731,945,969]" pageId="9" pageNumber="528" type="paratype">paratype</typeStatus>
(number of preparate 2971a) (kindly provided by N. Kluge). Herein, 
<taxonomicName id="4C654D513D3F520D58C6FC503F0AFC31" box="[245,377,980,1004]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3F520D58C6FC503F0AFC31" box="[245,377,980,1004]" italics="true" pageId="9" pageNumber="528">A. longulus</emphasis>
</taxonomicName>
is redescribed and new illustrations are presented, confirming that the larva illustrated by Sinichenkova (1981) is 
<emphasis id="B911EAC03D3F520D5991FC7C3C5AFBCD" box="[418,553,1016,1040]" italics="true" pageId="9" pageNumber="528">
<taxonomicName id="4C654D513D3F520D5991FC7C3C56FBCD" box="[418,549,1016,1040]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="longulus">A. longulus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDA36D23D3F520D58A4FBE73F2BFB42" blockId="9.[151,580,1123,1183]" pageId="9" pageNumber="528">
<taxonomicName id="4C654D513D3F520D58A4FBE73C37FBA1" authority="Matsumura 1931" authorityName="Matsumura" authorityYear="1931" box="[151,580,1123,1148]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D58A4FBE73C48FBA1" bold="true" box="[151,571,1123,1148]" pageId="9" pageNumber="528">
<emphasis id="B911EAC03D3F520D58A4FBE73F2DFBA1" bold="true" box="[151,350,1123,1148]" italics="true" pageId="9" pageNumber="528">Ameletus costalis</emphasis>
(Matsumura 1931
</emphasis>
)
</taxonomicName>
(
<figureCitation id="135E2A573D3F520D58ACFB033E8EFB42" box="[159,253,1159,1183]" captionStart="FIGURES 21 – 23" captionStartId="3.[151,264,1977,2000]" captionTargetBox="[165,1422,733,1939]" captionTargetId="figure@3.[151,1436,713,1955]" captionTargetPageId="3" captionText="FIGURES 21 – 23. Comb-shaped setae of maxilla: 21. Ameletus khasanensis sp. nov., 22. Ameletus longulus, 23. Ameletus costalis. The arrow shows the first seta." httpUri="https://zenodo.org/record/223957/files/figure.png" pageId="9" pageNumber="528">Figs. 23</figureCitation>
, 
<figureCitation id="135E2A573D3F520D593BFB033F56FB42" box="[264,293,1159,1183]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">50</figureCitation>
–71)
</paragraph>
<paragraph id="8BDA36D23D3F520D58A4FB553CBAFAFE" blockId="9.[151,1436,1232,1315]" pageId="9" pageNumber="528">
Matsumura 1931: 1474 (
<taxonomicName id="4C654D513D3F520D59A9FB543C39FB3B" box="[410,586,1232,1254]" class="Insecta" family="Ameletidae" genus="Chimura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D59A9FB543C39FB3B" box="[410,586,1232,1254]" italics="true" pageId="9" pageNumber="528">Chimura costalis</emphasis>
</taxonomicName>
); Imanishi 1932: 526–527, Pl 31 fig. 1, Pl 32 figs. 2–3 (
<taxonomicName id="4C654D513D3F520D5C95FB543BF6FB3B" box="[1190,1413,1232,1254]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="B911EAC03D3F520D5C95FB543BF6FB3B" box="[1190,1413,1232,1254]" italics="true" pageId="9" pageNumber="528">Ameletus sapporensis</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C654D513D3F520D58F4FB6B3F07FAD8" box="[199,372,1263,1285]" class="Insecta" family="Ameletidae" genus="Chimura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D58F4FB6B3F07FAD8" box="[199,372,1263,1285]" italics="true" pageId="9" pageNumber="528">Chimura costalis</emphasis>
</taxonomicName>
); Imanishi 1933: 65–66, figs. 3,5 (
<taxonomicName id="4C654D513D3F520D5AE4FB6B3DFBFAD8" box="[727,904,1263,1285]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D5AE4FB6B3DFBFAD8" box="[727,904,1263,1285]" italics="true" pageId="9" pageNumber="528">Ameletus costalis</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C654D513D3F520D5B97FB6B3A22FAD8" box="[932,1105,1263,1285]" class="Insecta" family="Ameletidae" genus="Chimura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D5B97FB6B3A22FAD8" box="[932,1105,1263,1285]" italics="true" pageId="9" pageNumber="528">Chimura costalis</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C654D513D3F520D5C5EFB6B3B3BFAD8" box="[1133,1352,1263,1285]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="B911EAC03D3F520D5C5EFB6B3B3BFAD8" box="[1133,1352,1263,1285]" italics="true" pageId="9" pageNumber="528">Ameletus sapporensis</emphasis>
</taxonomicName>
); Kluge 2007:257 (
<taxonomicName id="4C654D513D3F520D5906FA893F96FAFE" box="[309,485,1293,1315]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3F520D5906FA893F96FAFE" box="[309,485,1293,1315]" italics="true" pageId="9" pageNumber="528">Ameletus costalis</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C654D513D3F520D5A32FA893CC8FAFE" box="[513,699,1293,1315]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="528" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3F520D5A32FA893CC8FAFE" box="[513,699,1293,1315]" italics="true" pageId="9" pageNumber="528">Ameletus longulus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D3F520D58A4FACB3FB5FA51" pageId="9" pageNumber="528" type="materials_examined">
<paragraph id="8BDA36D23D3F520D58A4FACB3FB5FA51" blockId="9.[151,1436,1359,1996]" pageId="9" pageNumber="528">
<emphasis id="B911EAC03D3F520D58A4FACB3FF3FAB5" bold="true" box="[151,384,1359,1384]" pageId="9" pageNumber="528">Material examined.</emphasis>
<collectingCountry id="F37276423D3F520D59BBFAD43FB8FABA" box="[392,459,1360,1383]" name="Japan" pageId="9" pageNumber="528">Japan</collectingCountry>
, Nara Prefecture, Takami River, 
<date id="FFDB10123D3F520D5B08FACB3D95FABA" box="[827,998,1359,1383]" pageId="9" pageNumber="528" value="1997-03-23" valueMax="1997-03-25" valueMin="1997-03-23">23–25.III.1997</date>
, T Tiunova, 
<specimenCount id="9D63FD5B3D3F520D5C47FACB3AB1FABA" box="[1140,1218,1359,1383]" pageId="9" pageNumber="528" type="male">8 male</specimenCount>
, 
<specimenCount id="9D63FD5B3D3F520D5CFDFACB3B5CFABA" box="[1230,1327,1359,1383]" pageId="9" pageNumber="528" type="female">4 female</specimenCount>
imagines (reared), 12 larval exuviae.
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D3F520858F4FA133A21F844" lastPageId="12" lastPageNumber="531" pageId="9" pageNumber="528" type="description">
<paragraph id="8BDA36D23D3F520D58F4FA133BF3F811" blockId="9.[151,1436,1359,1996]" pageId="9" pageNumber="528">
<emphasis id="B911EAC03D3F520D58F4FA133F9FFA6D" bold="true" box="[199,492,1431,1456]" pageId="9" pageNumber="528">
Description. 
<emphasis id="B911EAC03D3F520D5952FA133F9FFA6D" bold="true" box="[353,492,1431,1456]" italics="true" pageId="9" pageNumber="528">Male imago</emphasis>
</emphasis>
(in alcohol). Length (mm): body 12.0–13.0; forewings 12.0–13.0; cerci 15.0–20.0. 
<emphasis id="B911EAC03D3F520D58A4FA383E93FA09" bold="true" box="[151,224,1468,1492]" pageId="9" pageNumber="528">Head:</emphasis>
upper portion of eyes gray; lower portion dark gray. Basal segments of antennae brown, tips pale. 
<emphasis id="B911EAC03D3F520D5D09FA383BEFFA09" bold="true" box="[1338,1436,1468,1492]" pageId="9" pageNumber="528">Thorax:</emphasis>
Medioscutum, submedioscutum and anteronotal protuberance light brown; medial longitudinal suture narrow, brown. Anteronotal transverse impression brown. Sublateroscutum from dark brown. Prelateroscutum white with brown margins. Medioparapsidal suture brown. Scutellum light brown; scuto-scutellar impression brown with pair of whitish spots. Posterior scutal protuberance light brown; parascutellum brown (
<figureCitation id="135E2A573D3F520D5C69F9C83AC0F9B9" box="[1114,1203,1612,1636]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Fig. 50</figureCitation>
). Femur of forelegs brown, tibia light brown, dorsal surface with spinules (
<figureCitation id="135E2A573D3F520D5ACAF9EB3D39F95A" box="[761,842,1647,1671]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Fig. 51</figureCitation>
), first tarsal segments light brown, other brownish or yellowish; middle and hind legs yellowish, joint brown. Length (mm) of foreleg segments: femora 2.7–3.0; tibia 2.3–2.7; tarsal segments 0.5–0.7, 1.7–2.0, 1.5–1.8, 1.1–1.3, and 0.5–0.6. Wings hyaline; near the base brown; crossveins of forewings between C and R veins dark brown and bordered by brown; other veins from brown to brownish (
<figureCitation id="135E2A573D3F520D5926F8843F03F8C5" box="[277,368,1792,1816]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Fig. 52</figureCitation>
). 
<collectingCountry id="F37276423D3F520D59B9F8843FB1F8C5" box="[394,450,1792,1816]" name="India" pageId="9" pageNumber="528">Hind</collectingCountry>
wings brownish at the base, all veins yellowish or colorless. 
<emphasis id="B911EAC03D3F520D5C89F97B3B5DF8C5" bold="true" box="[1210,1326,1791,1816]" pageId="9" pageNumber="528">Abdomen</emphasis>
: Styliger brownish, distal margin darker brownish. First segment of forceps extended distally, brown at the base; second and third segments pale yellowish (
<figureCitation id="135E2A573D3F520D59CBF8CC3C3EF8BD" box="[504,589,1864,1888]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Fig. 53</figureCitation>
). Penis lobes pale on inner margin and brown on the external sides; lateral penis lobes rounded and bent medially (
<figureCitation id="135E2A573D3F520D5A61F8E83CA8F859" box="[594,731,1900,1924]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Figs. 55–56</figureCitation>
); each ventral plate bears a single long pointed denticle parallel to the lateral lobe; denticle tips curved inside (
<figureCitation id="135E2A573D3F520D5A94F8143D74F875" box="[679,775,1936,1960]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Figs. 54</figureCitation>
); the distance between bases of the lateral penis lobes and denticles 2.5 of the maximum width of the denticles (
<figureCitation id="135E2A573D3F520D5ADFF8303D32F811" box="[748,833,1972,1996]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="9" pageNumber="528">Fig. 57</figureCitation>
). Cerci brownish at the base and yellowish distally.
</paragraph>
<paragraph id="8BDA36D23D3C520E58F4FF133C27FEE2" blockId="10.[151,1437,151,319]" pageId="10" pageNumber="529">
<emphasis id="B911EAC03D3C520E58F4FF133F03FF6D" bold="true" box="[199,368,151,176]" italics="true" pageId="10" pageNumber="529">Female imago</emphasis>
. Length (mm): body 16.5–18.5; forewings 16.5–17.4; cerci 20.0. Overall color light brown. 
<emphasis id="B911EAC03D3C520E58A4FF383E8AFF09" bold="true" box="[151,249,188,212]" pageId="10" pageNumber="529">Thorax:</emphasis>
Mesonotum light brown. Forelegs brown; middle and hind legs light brown, tarsal segments darker. Color of forewings, as in the male; hindwings colorless. 
<emphasis id="B911EAC03D3C520E5ADCFF5B3D1EFF25" bold="true" box="[751,877,223,248]" pageId="10" pageNumber="529">Abdomen:</emphasis>
tergum I dark brown; terga II–IX light brown, posterior area darker (abdomen with eggs). Sterna brownish. Subgenital plate with almost smooth margin (
<figureCitation id="135E2A573D3C520E5D0FFE803BE3FEC1" box="[1340,1424,260,284]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="10" pageNumber="529">Fig. 58</figureCitation>
). Cerci light brown, tips same but lighter.
</paragraph>
<caption id="DF1A665A3D3C520E58A4F8173A6FF83A" httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="10" pageNumber="529" targetBox="[156,1435,362,1938]" targetPageId="10">
<paragraph id="8BDA36D23D3C520E58A4F8173A6FF83A" blockId="10.[151,1436,1939,2024]" pageId="10" pageNumber="529">
<emphasis id="B911EAC03D3C520E58A4F8173F2BF874" bold="true" box="[151,344,1939,1962]" pageId="10" pageNumber="529">FIGURES 50–58.</emphasis>
<taxonomicName id="4C654D513D3C520E5951F8103C64F877" box="[354,535,1940,1962]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="529" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3C520E5951F8103C64F877" box="[354,535,1940,1962]" italics="true" pageId="10" pageNumber="529">Ameletus costalis</emphasis>
</taxonomicName>
: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50=1.0; 52=2.0; 53–58=0.5.
</paragraph>
</caption>
<caption id="DF1A665A3D3D520F58A4F9F63C8FF91B" httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="11" pageNumber="530" targetBox="[247,1340,221,1606]" targetPageId="11">
<paragraph id="8BDA36D23D3D520F58A4F9F63C8FF91B" blockId="11.[151,1437,1650,1734]" pageId="11" pageNumber="530">
<emphasis id="B911EAC03D3D520F58A4F9F63F3DF955" bold="true" box="[151,334,1650,1673]" pageId="11" pageNumber="530">FIGURES 59–67</emphasis>
. 
<taxonomicName id="4C654D513D3D520F596FF9F73C7EF954" box="[348,525,1651,1673]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="530" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3D520F596FF9F73C7EF954" box="[348,525,1651,1673]" italics="true" pageId="11" pageNumber="530">Ameletus costalis</emphasis>
</taxonomicName>
, larva: 59–60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62–67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62–68=0.5; 60–61=0.2. The arrow shows the group of setae.
</paragraph>
</caption>
<paragraph id="8BDA36D23D3D520858F4F96A3ACFFEC1" blockId="11.[151,1436,1773,2014]" lastBlockId="12.[151,1436,151,284]" lastPageId="12" lastPageNumber="531" pageId="11" pageNumber="530">
<emphasis id="B911EAC03D3D520F58F4F96A3F2DF8DB" bold="true" box="[199,350,1773,1798]" italics="true" pageId="11" pageNumber="530">Mature larva</emphasis>
. Length (mm): body 12.5–15.5; cerci 5.2–7.3. 
<emphasis id="B911EAC03D3D520F5B5FF96A3DC6F8DB" bold="true" box="[876,949,1774,1798]" pageId="11" pageNumber="530">Head:</emphasis>
Labrum (length 0.71–0.85 of width) with a shallow apical incision; first comb-shaped seta of maxilla with wide and rounded denticles, their number less than ten (
<figureCitation id="135E2A573D3D520F58F8F8B13F52F890" box="[203,289,1845,1869]" captionStart="FIGURES 21 – 23" captionStartId="3.[151,264,1977,2000]" captionTargetBox="[165,1422,733,1939]" captionTargetId="figure@3.[151,1436,713,1955]" captionTargetPageId="3" captionText="FIGURES 21 – 23. Comb-shaped setae of maxilla: 21. Ameletus khasanensis sp. nov., 22. Ameletus longulus, 23. Ameletus costalis. The arrow shows the first seta." httpUri="https://zenodo.org/record/223957/files/figure.png" pageId="11" pageNumber="530">Fig. 23</figureCitation>
). Incisor of the left mandible with four denticles; the first and second denticles of equal length, the fourth smallest (
<figureCitation id="135E2A573D3D520F597CF8DD3FD7F8AC" box="[335,420,1881,1905]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="11" pageNumber="530">Fig. 59</figureCitation>
). Incisor of the right mandible with three denticles; the first length equal to the third, both almost equal to the second (
<figureCitation id="135E2A573D3D520F59FDF8F93C51F848" box="[462,546,1917,1941]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="11" pageNumber="530">Fig. 60</figureCitation>
). 
<emphasis id="B911EAC03D3D520F5A06F8FA3CE4F84B" bold="true" box="[565,663,1918,1942]" pageId="11" pageNumber="530">Thorax:</emphasis>
Lengths (mm) of leg segments are as follows. Foreleg: 1.6–2.0; tibia 0.9–1.0; and tarsus 1.1–1.3. Middle leg: femur 1.6–2.1; tibia 0.9–1.3; and tarsus 1.1–1.2. 
<collectingCountry id="F37276423D3D520F5CB6F8253ACEF864" box="[1157,1213,1953,1977]" name="India" pageId="11" pageNumber="530">Hind</collectingCountry>
leg: femur 1.6–2.2; tibia 0.9–1.3; and tarsus 0.9–1.2. A dense group of four or five setae on the dorsal surface at the base of the fore femora (
<figureCitation id="135E2A573D3A520858C6FF133F3AFF72" box="[245,329,151,175]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="12" pageNumber="531">Fig. 61</figureCitation>
). 
<emphasis id="B911EAC03D3A5208596EFF133FA8FF6D" bold="true" box="[349,475,151,176]" pageId="12" pageNumber="531">Abdomen:</emphasis>
All terga with irregular row of spines. Gill I and II small, rounded in distal part; gill I without anal ribs, width 0.65 of length (
<figureCitation id="135E2A573D3A52085A50FF383CC9FF09" box="[611,698,188,212]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="12" pageNumber="531">Fig. 62</figureCitation>
); gill II with small anal ribs, width 0.68 of length (
<figureCitation id="135E2A573D3A52085D39FF383B2CFF09" box="[1290,1375,188,212]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="12" pageNumber="531">Fig. 63</figureCitation>
); gill III–V with rounded distal margin, width 0.51–0.53 of length (64–65); gill VI with straight distal margin, width 0.53 of length (
<figureCitation id="135E2A573D3A52085939FE803F2CFEC1" box="[266,351,260,284]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="12" pageNumber="531">Fig. 66</figureCitation>
); gill VII with a truncated inner distal margin, width 0.48 of length (
<figureCitation id="135E2A573D3A52085C6FFE803AC3FEC1" box="[1116,1200,260,284]" captionStart="FIGURES 59 – 67" captionStartId="11.[151,264,1650,1673]" captionTargetBox="[247,1340,221,1606]" captionTargetId="figure@11.[223,1363,193,1629]" captionTargetPageId="11" captionText="FIGURES 59 – 67. Ameletus costalis, larva: 59 – 60. Apices of mandibles, dorsal view: 59. Left mandible, 60. Right mandible. 61. Femur of fore leg. 62 – 67. Gills: 62. Gill I, 63. Gill II, 64. Gill III, 65. Gill IV, 66. Gill VI, 67. Gill VII. Scale (mm): 62 – 68 = 0.5; 60 – 61 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223963/files/figure.png" pageId="12" pageNumber="531">Fig. 67</figureCitation>
).
</paragraph>
<caption id="DF1A665A3D3A520858A4F9EF3CB9F97C" httpUri="https://zenodo.org/record/223964/files/figure.png" pageId="12" pageNumber="531" targetBox="[166,1422,350,1608]" targetPageId="12">
<paragraph id="8BDA36D23D3A520858A4F9EF3CB9F97C" blockId="12.[151,1437,1643,1697]" pageId="12" pageNumber="531">
<emphasis id="B911EAC03D3A520858A4F9EF3F3FF95C" bold="true" box="[151,332,1643,1666]" pageId="12" pageNumber="531">FIGURES 68–70</emphasis>
. 
<taxonomicName id="4C654D513D3A5208596AF9E83C7AF95F" box="[345,521,1644,1666]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="531" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3A5208596AF9E83C7AF95F" box="[345,521,1644,1666]" italics="true" pageId="12" pageNumber="531">Ameletus costalis</emphasis>
</taxonomicName>
, egg: 68. General view. 69. Structure of chorion and small knob-terminated coiled threads. 70. Large knobs. The arrow shows the micropiles (mp).
</paragraph>
</caption>
<paragraph id="8BDA36D23D3A520858F4F9493A21F844" blockId="12.[151,1436,1740,2018]" pageId="12" pageNumber="531">
<emphasis id="B911EAC03D3A520858F4F9493F7BF939" bold="true" box="[199,264,1741,1764]" italics="true" pageId="12" pageNumber="531">Eggs.</emphasis>
General form of egg flattened ellipse with length 180–187 μm and a width 110–123 μm (
<figureCitation id="135E2A573D3A52085D35F9493B2FF938" box="[1286,1372,1741,1765]" captionStart="FIGURES 68 – 70" captionStartId="12.[151,264,1643,1666]" captionTargetBox="[166,1422,350,1608]" captionTargetId="figure@12.[151,1436,337,1622]" captionTargetPageId="12" captionText="FIGURES 68 – 70. Ameletus costalis, egg: 68. General view. 69. Structure of chorion and small knob-terminated coiled threads. 70. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223964/files/figure.png" pageId="12" pageNumber="531">Fig. 68</figureCitation>
). The chorionic surface covered by irregularly shaped coarse cells; each cell with a ribbed bottom and well-marked protuberances, one of which is the largest; borders of cells are in contact with tubercles, relatively high (
<figureCitation id="135E2A573D3A52085D0AF8913BFCF8F0" box="[1337,1423,1813,1837]" captionStart="FIGURES 68 – 70" captionStartId="12.[151,264,1643,1666]" captionTargetBox="[166,1422,350,1608]" captionTargetId="figure@12.[151,1436,337,1622]" captionTargetPageId="12" captionText="FIGURES 68 – 70. Ameletus costalis, egg: 68. General view. 69. Structure of chorion and small knob-terminated coiled threads. 70. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223964/files/figure.png" pageId="12" pageNumber="531">Fig. 69</figureCitation>
); small round flower-like knob-terminated coiled threads, relatively rare and evenly distributed throughout the surface of the egg (
<figureCitation id="135E2A573D3A5208594AF8D93FA2F8A8" box="[377,465,1885,1909]" captionStart="FIGURES 68 – 70" captionStartId="12.[151,264,1643,1666]" captionTargetBox="[166,1422,350,1608]" captionTargetId="figure@12.[151,1436,337,1622]" captionTargetPageId="12" captionText="FIGURES 68 – 70. Ameletus costalis, egg: 68. General view. 69. Structure of chorion and small knob-terminated coiled threads. 70. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223964/files/figure.png" pageId="12" pageNumber="531">Fig. 69</figureCitation>
); large and round knobs arranged on one pole only (
<figureCitation id="135E2A573D3A52085C06F8D93AB7F8A8" box="[1077,1220,1885,1909]" captionStart="FIGURES 68 – 70" captionStartId="12.[151,264,1643,1666]" captionTargetBox="[166,1422,350,1608]" captionTargetId="figure@12.[151,1436,337,1622]" captionTargetPageId="12" captionText="FIGURES 68 – 70. Ameletus costalis, egg: 68. General view. 69. Structure of chorion and small knob-terminated coiled threads. 70. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223964/files/figure.png" pageId="12" pageNumber="531">Figs. 68, 70</figureCitation>
). Three micropiles visible in the subequatorial area; sperm guide round (
<figureCitation id="135E2A573D3A52085AD8F8053D4DF844" box="[747,830,1921,1945]" captionStart="FIGURES 68 – 70" captionStartId="12.[151,264,1643,1666]" captionTargetBox="[166,1422,350,1608]" captionTargetId="figure@12.[151,1436,337,1622]" captionTargetPageId="12" captionText="FIGURES 68 – 70. Ameletus costalis, egg: 68. General view. 69. Structure of chorion and small knob-terminated coiled threads. 70. Large knobs. The arrow shows the micropiles (mp)." httpUri="https://zenodo.org/record/223964/files/figure.png" pageId="12" pageNumber="531">Fig. 68</figureCitation>
); micropylar rim absent.
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D3A520958F4F8203A13FF09" lastPageId="13" lastPageNumber="532" pageId="12" pageNumber="531" type="distribution">
<paragraph id="8BDA36D23D3A520958F4F8203A13FF09" blockId="12.[151,1436,1740,2018]" lastBlockId="13.[151,1436,151,1077]" lastPageId="13" lastPageNumber="532" pageId="12" pageNumber="531">
<emphasis id="B911EAC03D3A520858F4F8203F9FF860" bold="true" box="[199,492,1956,1981]" pageId="12" pageNumber="531">Distribution and biology</emphasis>
. The larval exuviae and the subimago emerged from the larvae were collected in a foothill river in 
<collectingCountry id="F37276423D3A52085960F84E3FE0F83C" box="[339,403,1994,2017]" name="Japan" pageId="12" pageNumber="531">Japan</collectingCountry>
at the end of March. The subimagos emerged on a warm sunny day. The mature larvae crawled, half-submerged to large stones protruding from the water and molted to subimago. According to Imanishi (1932), 
<taxonomicName id="4C654D513D3B520958DCFF383F17FF09" box="[239,356,188,212]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B520958DCFF383F17FF09" box="[239,356,188,212]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
is one of the earliest-flying Japanese species of the genus 
<taxonomicName id="4C654D513D3B52095BC1FF383A2AFF09" box="[1010,1113,188,212]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAC03D3B52095BC1FF383A2AFF09" box="[1010,1113,188,212]" italics="true" pageId="13" pageNumber="532">Ameletus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F65593D3B520958F4FF643CD5FBE9" pageId="13" pageNumber="532" type="discussion">
<paragraph id="8BDA36D23D3B520958F4FF643DC7FE71" blockId="13.[151,1436,151,1077]" pageId="13" pageNumber="532">
<emphasis id="B911EAC03D3B520958F4FF643F48FF25" bold="true" box="[199,315,224,248]" pageId="13" pageNumber="532">Remarks.</emphasis>
Until now it was thought that 
<taxonomicName id="4C654D513D3B52095ABDFF643D70FF25" box="[654,771,224,248]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B52095ABDFF643D70FF25" box="[654,771,224,248]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
inhabited streams of the Russian Far East and 
<collectingCountry id="F37276423D3B52095D3EFF643B23FF2A" box="[1293,1360,224,247]" name="Japan" pageId="13" pageNumber="532">Japan</collectingCountry>
. Study of a new material shows that the japan-continental species 
<taxonomicName id="4C654D513D3B52095B00FE803DDAFEC1" box="[819,937,260,284]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B52095B00FE803DDAFEC1" box="[819,937,260,284]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
represents a group of species, including the Japanese species 
<taxonomicName id="4C654D513D3B52095950FEAC3FADFE9D" box="[355,478,296,320]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" sensu="stricto" species="costalis">
<emphasis id="B911EAC03D3B52095950FEAC3FADFE9D" box="[355,478,296,320]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B520959DAFEAC3C6AFEE2" box="[489,537,296,319]" pageId="13" pageNumber="532" sensu="stricto">s. s.</taxonomicNameLabel>
and two separate continental species. The new species herein described, 
<taxonomicName id="4C654D513D3B52095DB7FEAC3F6DFEB9" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095DB7FEAC3F6DFEB9" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095916FEC83F0CFEB9" box="[293,383,332,356]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095916FEC83F0CFEB9" bold="true" box="[293,383,332,356]" pageId="13" pageNumber="532">sp. nov.</emphasis>
</taxonomicNameLabel>
and the redescribed species 
<taxonomicName id="4C654D513D3B52095AF1FEC83D32FEB9" box="[706,833,332,356]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095AF1FEC83D32FEB9" box="[706,833,332,356]" italics="true" pageId="13" pageNumber="532">A. longulus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C654D513D3B52095B63FEC83A6CFEB9" box="[848,1055,332,356]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095B63FEC83A6CFEB9" box="[848,1055,332,356]" italics="true" pageId="13" pageNumber="532">Ameletus longulus</emphasis>
</taxonomicName>
was regarded to be a synonym of 
<taxonomicName id="4C654D513D3B520958A4FEF43F62FE55" box="[151,273,368,392]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B520958A4FEF43F62FE55" box="[151,273,368,392]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
, when 
<taxonomicName id="4C654D513D3B52095942FEF43F9FFE55" box="[369,492,368,392]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B52095942FEF43F9FFE55" box="[369,492,368,392]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
was accepted in wide sense, including the continental form (Sinitshenkova &amp; Tshernova 1976; Tiunova &amp; Potikha 2005; Tiunova 2006, 2007, 2009).
</paragraph>
<paragraph id="8BDA36D23D3B520958F4FE333CD5FBE9" blockId="13.[151,1436,151,1077]" pageId="13" pageNumber="532">
<emphasis id="B911EAC03D3B520958F4FE333F3FFE0D" bold="true" box="[199,332,439,464]" pageId="13" pageNumber="532">Discussion.</emphasis>
The male imago of 
<taxonomicName id="4C654D513D3B52095A00FE3C3CA9FE0D" box="[563,730,440,464]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095A00FE3C3CA9FE0D" box="[563,730,440,464]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095AD1FE3C3D44FE0D" box="[738,823,440,464]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095AD1FE3C3D44FE0D" bold="true" box="[738,823,440,464]" pageId="13" pageNumber="532">sp. nov</emphasis>
</taxonomicNameLabel>
is similar to 
<taxonomicName id="4C654D513D3B52095BFEFE3C3A30FE0D" box="[973,1091,440,464]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B52095BFEFE3C3A30FE0D" box="[973,1091,440,464]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
by its forewings, in which the crossveins between the C and R veins are dark brown and are bordered by brown (
<figureCitation id="135E2A573D3B52095C0FFE583AFFFE29" box="[1084,1164,476,500]" captionStart="FIGURES 1 – 9" captionStartId="1.[151,264,1877,1900]" captionTargetBox="[153,1406,270,1852]" captionTargetId="figure@1.[151,1436,266,1854]" captionTargetPageId="1" captionText="FIGURES 1 – 9. Ameletus khasanensis sp. nov., male imago: 1. Mesonotum, dorsal view. 2. Fore wing. 3. Hind wing. 4. Sternum IX, styliger and gonostyli. 5. Genitalia, lateral view. 6. Penis, ventral view. 7. Penis, dorsal view. 8. Penis, lateral view. 9. Subgenital plate, female, dorsal view. Scale (mm): 4 – 9 = 0.5; 1, 3 = 1.0; 2 = 2.0." httpUri="https://zenodo.org/record/223955/files/figure.png" pageId="13" pageNumber="532">Figs. 2</figureCitation>
, 
<figureCitation id="135E2A573D3B52095CAAFE583ACAFE29" box="[1177,1209,476,500]" captionStart="FIGURES 50 – 58" captionStartId="10.[151,264,1939,1962]" captionTargetBox="[156,1435,362,1938]" captionTargetId="figure@10.[151,1436,357,1945]" captionTargetPageId="10" captionText="FIGURES 50 – 58. Ameletus costalis: male imago: 50. Mesonotum, dorsal view. 51. Tibia of foreleg. 52. Fore wing. 53. Sternum IX, styliger and gonostyli. 54. Genitalia, lateral view. 55. Penis, ventral view. 56. Penis, dorsal view. 57. Penis, lateral view. 58. Subgenital plate, female, dorsal view. Scale (mm): 50 = 1.0; 52 = 2.0; 53 – 58 = 0.5." httpUri="https://zenodo.org/record/223962/files/figure.png" pageId="13" pageNumber="532">52</figureCitation>
); however, the male imago can be distinguished from that of 
<taxonomicName id="4C654D513D3B52095A69FD843CBCFDC5" box="[602,719,512,536]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B52095A69FD843CBCFDC5" box="[602,719,512,536]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
by the absence of spinules on the dorsal surface of the fore tibia and the structure of the penis. The male imago of 
<taxonomicName id="4C654D513D3B52095AE5FDA03D24FDE1" box="[726,855,548,572]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095AE5FDA03D24FDE1" box="[726,855,548,572]" italics="true" pageId="13" pageNumber="532">A. longulus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C654D513D3B52095CA3FDA03B4BFDE1" box="[1168,1336,548,572]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095CA3FDA03B4BFDE1" box="[1168,1336,548,572]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095D72FDA03BEFFDE1" box="[1345,1436,548,572]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095D72FDA03BE6FDE1" bold="true" box="[1345,1429,548,572]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
and 
<taxonomicName id="4C654D513D3B520958F5FDCC3F4AFDBD" box="[198,313,584,608]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B520958F5FDCC3F4AFDBD" box="[198,313,584,608]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
by the color of forewings, which are brownish with all veins brown in 
<taxonomicName id="4C654D513D3B52095C62FDCC3AA2FDBD" box="[1105,1233,584,608]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095C62FDCC3AA2FDBD" box="[1105,1233,584,608]" italics="true" pageId="13" pageNumber="532">A. longulus</emphasis>
</taxonomicName>
(
<figureCitation id="135E2A573D3B52095CECFDC33B1BFD82" box="[1247,1384,583,607]" captionStart="FIGURES 27 – 35" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[168,1414,324,1899]" captionTargetId="figure@6.[161,1426,314,1902]" captionTargetPageId="6" captionText="FIGURES 27 – 35. Ameletus longulus, male imago: 27. Mesonotum, dorsal view. 28. Fore wing. 29. Hind wing. 30. Sternum IX, styliger and gonostyli. 31. Genitalia, lateral view. 32. Penis, ventral view. 33. Penis, dorsal view. 34. Penis, lateral view. 35. Subgenital plate, female, dorsal view. Scale (mm): 30 – 35 = 0.5; 28 – 29 = 2.0; 27 = 1.0" httpUri="https://zenodo.org/record/223959/files/figure.png" pageId="13" pageNumber="532">Figs. 28–29</figureCitation>
). By the structure of the gills, 
<taxonomicName id="4C654D513D3B5209598CFDE83C1BFD59" box="[447,616,620,644]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B5209598CFDE83C1BFD59" box="[447,616,620,644]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095A40FDE83CBDFD59" box="[627,718,620,644]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095A40FDE83CBBFD59" bold="true" box="[627,712,620,644]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
, 
<taxonomicName id="4C654D513D3B52095AD3FDE83D10FD59" box="[736,867,620,644]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095AD3FDE83C82FD59" box="[736,753,620,644]" italics="true" pageId="13" pageNumber="532">A</emphasis>
. 
<emphasis id="B911EAC03D3B52095B31FDE83D10FD59" box="[770,867,620,644]" italics="true" pageId="13" pageNumber="532">longulus</emphasis>
</taxonomicName>
, 
<emphasis id="B911EAC03D3B52095B47FDE83AF5FD59" box="[884,1158,620,644]" italics="true" pageId="13" pageNumber="532">
<taxonomicName id="4C654D513D3B52095B47FDE83D9DFD59" box="[884,1006,620,644]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">A. costalis</taxonomicName>
, 
<taxonomicName id="4C654D513D3B52095BCFFDE83AF5FD59" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[1020,1158,620,644]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="formosus">A. formosus</taxonomicName>
</emphasis>
Kang &amp; Yang 1994, 
<taxonomicName id="4C654D513D3B52095DB7FDE83FB9FD7A" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="B911EAC03D3B52095DB7FDE83E95FD75" italics="true" pageId="13" pageNumber="532">A. atratus</emphasis>
Kang &amp; Yang 1994
</taxonomicName>
and 
<taxonomicName id="4C654D513D3B52095A32FD143DF5FD7A" authority="Kang &amp; Yang 1994" authorityName="Kang &amp; Yang" authorityYear="1994" box="[513,902,655,680]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B911EAC03D3B52095A32FD143CD1FD75" box="[513,674,656,680]" italics="true" pageId="13" pageNumber="532">A. montivagus</emphasis>
Kang &amp; Yang 1994
</taxonomicName>
can be assigned to a group of species in which gills III–VI bear an anal rib far from the anal margin. The larvae of 
<taxonomicName id="4C654D513D3B52095BA2FD303A44FD11" box="[913,1079,692,716]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095BA2FD303A44FD11" box="[913,1079,692,716]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095C0CFD303AEAFD11" box="[1087,1177,692,716]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095C0CFD303AE1FD11" bold="true" box="[1087,1170,692,716]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
differ from those of 
<taxonomicName id="4C654D513D3B52095DB7FD303E84FD2D" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095DB7FD303BE6FD11" box="[1412,1429,692,716]" italics="true" pageId="13" pageNumber="532">A</emphasis>
. 
<emphasis id="B911EAC03D3B520958A4FD5C3E84FD2D" box="[151,247,728,752]" italics="true" pageId="13" pageNumber="532">longulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C654D513D3B52095936FD5C3FFAFD2D" box="[261,393,728,752]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B911EAC03D3B52095936FD5C3FFAFD2D" box="[261,393,728,752]" italics="true" pageId="13" pageNumber="532">A. formosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C654D513D3B520959F3FD5C3C5EFD2D" box="[448,557,728,752]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="B911EAC03D3B520959F3FD5C3C5EFD2D" box="[448,557,728,752]" italics="true" pageId="13" pageNumber="532">A. atratus</emphasis>
</taxonomicName>
in the structure of gills I. In 
<taxonomicName id="4C654D513D3B52095B58FD5C3A62FD2D" box="[875,1041,728,752]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095B58FD5C3A62FD2D" box="[875,1041,728,752]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095C24FD5C3A1CFD32" box="[1047,1135,728,752]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095C24FD5C3A1AFD2D" bold="true" box="[1047,1129,728,752]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
, gill I has no vestigial anal ribs (
<figureCitation id="135E2A573D3B520958E3FD783F50FCC9" box="[208,291,764,788]" captionStart="FIGURES 10 – 20" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[154,1429,209,1616]" captionTargetId="figure@2.[151,1436,193,1667]" captionTargetPageId="2" captionText="FIGURES 10 – 20. Ameletus khasanensis sp. nov., larva: 10. Labrum, dorsal view. 11 – 12. Apices of mandibles, dorsal view: 11 – left mandible, 12 right mandible. 13. Labium, dorsal view. 14. Femur of foreleg. 15 – 20. Gills: 15. Gill I, 16. Gill II, 17. Gill III, 18. Gill IV, 19 Gill VI, 20. Gill VII. Scale (mm): 10, 13, 15 – 20 = 0.5; 11 – 12 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223956/files/figure.png" pageId="13" pageNumber="532">Fig. 15</figureCitation>
), in contrast to 
<emphasis id="B911EAC03D3B520959E1FD783CA7FCC9" box="[466,724,764,788]" italics="true" pageId="13" pageNumber="532">
<taxonomicName id="4C654D513D3B520959E1FD783C28FCC9" box="[466,603,764,788]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="formosus">A. formosus</taxonomicName>
, 
<taxonomicName id="4C654D513D3B52095A55FD783CA7FCC9" box="[614,724,764,788]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="atratus">A. atratus</taxonomicName>
</emphasis>
[
<figureCitation id="135E2A573D3B52095AD7FD783D4FFCC9" box="[740,828,764,788]" captionStart="FIGURES 1 – 9" captionStartId="1.[151,264,1877,1900]" captionTargetBox="[153,1406,270,1852]" captionTargetId="figure@1.[151,1436,266,1854]" captionTargetPageId="1" captionText="FIGURES 1 – 9. Ameletus khasanensis sp. nov., male imago: 1. Mesonotum, dorsal view. 2. Fore wing. 3. Hind wing. 4. Sternum IX, styliger and gonostyli. 5. Genitalia, lateral view. 6. Penis, ventral view. 7. Penis, dorsal view. 8. Penis, lateral view. 9. Subgenital plate, female, dorsal view. Scale (mm): 4 – 9 = 0.5; 1, 3 = 1.0; 2 = 2.0." httpUri="https://zenodo.org/record/223955/files/figure.png" pageId="13" pageNumber="532">Fig.1, 3</figureCitation>
(Kang &amp; Yang 1994)], and 
<taxonomicName id="4C654D513D3B52095C44FD783A84FCC9" box="[1143,1271,764,788]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095C44FD783A84FCC9" box="[1143,1271,764,788]" italics="true" pageId="13" pageNumber="532">A. longulus</emphasis>
</taxonomicName>
(
<figureCitation id="135E2A573D3B52095D35FD783BFCFCC9" box="[1286,1423,764,788]" captionStart="FIGURES 36 – 46" captionStartId="7.[151,264,1700,1723]" captionTargetBox="[159,1415,193,1649]" captionTargetId="figure@7.[159,1427,193,1679]" captionTargetPageId="7" captionText="FIGURES 36 – 46. Ameletus longulus, larva: 36. Labrum, dorsal view. 37 – 38. Apices of mandibles, dorsal view: 37 – left mandible, 38 right mandible. 39. Labium, dorsal view. 40. Femur of foreleg. 41 – 46. Gills: 41. Gill I, 42. Gill II, 43. Gill III, 44. Gill IV, 45. Gill VI, 46. Gill VII. Scale (mm): 36, 39, 41 – 46 = 0.5; 37 – 38 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223960/files/figure.png" pageId="13" pageNumber="532">Figs. 41–42</figureCitation>
), in which gill I has vestigial anal ribs. The larvae of 
<taxonomicName id="4C654D513D3B52095AE7FCA43D08FCE5" box="[724,891,800,824]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095AE7FCA43D08FCE5" box="[724,891,800,824]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095BB2FCA43DA9FCE5" box="[897,986,800,824]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095BB2FCA43DA0FCE5" bold="true" box="[897,979,800,824]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
differ from 
<taxonomicName id="4C654D513D3B52095C51FCA43B71FCE5" box="[1122,1282,800,824]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B911EAC03D3B52095C51FCA43B71FCE5" box="[1122,1282,800,824]" italics="true" pageId="13" pageNumber="532">A. montivagus</emphasis>
</taxonomicName>
by the width/ length ratio of gill IV. In 
<taxonomicName id="4C654D513D3B520959F6FCC03C03FC81" box="[453,624,836,860]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B520959F6FCC03C03FC81" box="[453,624,836,860]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095A49FCC03CA5FC81" box="[634,726,836,860]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095A49FCC03CA3FC81" bold="true" box="[634,720,836,860]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
gill IV width/length ratio is 0.54 (
<figureCitation id="135E2A573D3B52095C43FCC03AB4FC81" box="[1136,1223,836,860]" captionStart="FIGURES 10 – 20" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[154,1429,209,1616]" captionTargetId="figure@2.[151,1436,193,1667]" captionTargetPageId="2" captionText="FIGURES 10 – 20. Ameletus khasanensis sp. nov., larva: 10. Labrum, dorsal view. 11 – 12. Apices of mandibles, dorsal view: 11 – left mandible, 12 right mandible. 13. Labium, dorsal view. 14. Femur of foreleg. 15 – 20. Gills: 15. Gill I, 16. Gill II, 17. Gill III, 18. Gill IV, 19 Gill VI, 20. Gill VII. Scale (mm): 10, 13, 15 – 20 = 0.5; 11 – 12 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223956/files/figure.png" pageId="13" pageNumber="532">Fig. 18</figureCitation>
), in contrast to 
<taxonomicName id="4C654D513D3B52095DB7FCC03F6BFC5D" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B911EAC03D3B52095DB7FCC03F6BFC5D" italics="true" pageId="13" pageNumber="532">A. montivagus</emphasis>
</taxonomicName>
, in which this ratio is 0.55–0.58 [
<figureCitation id="135E2A573D3B52095AAAFCE33CACFCA2" box="[665,735,871,895]" captionStart="FIGURES 1 – 9" captionStartId="1.[151,264,1877,1900]" captionTargetBox="[153,1406,270,1852]" captionTargetId="figure@1.[151,1436,266,1854]" captionTargetPageId="1" captionText="FIGURES 1 – 9. Ameletus khasanensis sp. nov., male imago: 1. Mesonotum, dorsal view. 2. Fore wing. 3. Hind wing. 4. Sternum IX, styliger and gonostyli. 5. Genitalia, lateral view. 6. Penis, ventral view. 7. Penis, dorsal view. 8. Penis, lateral view. 9. Subgenital plate, female, dorsal view. Scale (mm): 4 – 9 = 0.5; 1, 3 = 1.0; 2 = 2.0." httpUri="https://zenodo.org/record/223955/files/figure.png" pageId="13" pageNumber="532">Fig. 2</figureCitation>
(Kang &amp; Yang 1994)]. The larvae of 
<taxonomicName id="4C654D513D3B52095CA2FCEC3B4AFC5D" box="[1169,1337,872,896]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="khasanensis" status="sp. nov.">
<emphasis id="B911EAC03D3B52095CA2FCEC3B4AFC5D" box="[1169,1337,872,896]" italics="true" pageId="13" pageNumber="532">A. khasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257BB3D3B52095D72FCEC3BEFFC5D" box="[1345,1436,872,896]" pageId="13" pageNumber="532" rank="species">
<emphasis id="B911EAC03D3B52095D72FCEC3BE6FC5D" bold="true" box="[1345,1429,872,896]" pageId="13" pageNumber="532">sp. nov</emphasis>
.
</taxonomicNameLabel>
differ from those of 
<taxonomicName id="4C654D513D3B5209594EFC083F81FC79" box="[381,498,908,932]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B911EAC03D3B5209594EFC083F81FC79" box="[381,498,908,932]" italics="true" pageId="13" pageNumber="532">A. costalis</emphasis>
</taxonomicName>
in the length of the denticles on the right mandible (
<figureCitation id="135E2A573D3B52095C7AFC083AEFFC79" box="[1097,1180,908,932]" captionStart="FIGURES 10 – 20" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[154,1429,209,1616]" captionTargetId="figure@2.[151,1436,193,1667]" captionTargetPageId="2" captionText="FIGURES 10 – 20. Ameletus khasanensis sp. nov., larva: 10. Labrum, dorsal view. 11 – 12. Apices of mandibles, dorsal view: 11 – left mandible, 12 right mandible. 13. Labium, dorsal view. 14. Femur of foreleg. 15 – 20. Gills: 15. Gill I, 16. Gill II, 17. Gill III, 18. Gill IV, 19 Gill VI, 20. Gill VII. Scale (mm): 10, 13, 15 – 20 = 0.5; 11 – 12 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223956/files/figure.png" pageId="13" pageNumber="532">Fig. 12</figureCitation>
), in the absence of the group of setae on the dorsal surface at the base of the fore femora (
<figureCitation id="135E2A573D3B52095BA4FC2B3D99FC1A" box="[919,1002,943,967]" captionStart="FIGURES 10 – 20" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[154,1429,209,1616]" captionTargetId="figure@2.[151,1436,193,1667]" captionTargetPageId="2" captionText="FIGURES 10 – 20. Ameletus khasanensis sp. nov., larva: 10. Labrum, dorsal view. 11 – 12. Apices of mandibles, dorsal view: 11 – left mandible, 12 right mandible. 13. Labium, dorsal view. 14. Femur of foreleg. 15 – 20. Gills: 15. Gill I, 16. Gill II, 17. Gill III, 18. Gill IV, 19 Gill VI, 20. Gill VII. Scale (mm): 10, 13, 15 – 20 = 0.5; 11 – 12 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223956/files/figure.png" pageId="13" pageNumber="532">Fig. 14</figureCitation>
) and the width/length ratio of gill VII (
<figureCitation id="135E2A573D3B520958ACFC503E86FC31" box="[159,245,980,1004]" captionStart="FIGURES 10 – 20" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[154,1429,209,1616]" captionTargetId="figure@2.[151,1436,193,1667]" captionTargetPageId="2" captionText="FIGURES 10 – 20. Ameletus khasanensis sp. nov., larva: 10. Labrum, dorsal view. 11 – 12. Apices of mandibles, dorsal view: 11 – left mandible, 12 right mandible. 13. Labium, dorsal view. 14. Femur of foreleg. 15 – 20. Gills: 15. Gill I, 16. Gill II, 17. Gill III, 18. Gill IV, 19 Gill VI, 20. Gill VII. Scale (mm): 10, 13, 15 – 20 = 0.5; 11 – 12 = 0.2. The arrow shows the group of setae." httpUri="https://zenodo.org/record/223956/files/figure.png" pageId="13" pageNumber="532">Fig. 20</figureCitation>
). 
<taxonomicName id="4C654D513D3B5209593AFC503FABFC31" box="[265,472,980,1004]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B5209593AFC503FABFC31" box="[265,472,980,1004]" italics="true" pageId="13" pageNumber="532">Ameletus longulus</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C654D513D3B52095A5DFC503C86FC31" box="[622,757,980,1004]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B911EAC03D3B52095A5DFC503C86FC31" box="[622,757,980,1004]" italics="true" pageId="13" pageNumber="532">A. formosus</emphasis>
</taxonomicName>
in the presence of vestigial anal ribs on gills I–II, but differ from this species based on the width/length ratio of gill IV. In 
<taxonomicName id="4C654D513D3B52095B6BFC7C3DABFBCD" box="[856,984,1016,1040]" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B911EAC03D3B52095B6BFC7C3DABFBCD" box="[856,984,1016,1040]" italics="true" pageId="13" pageNumber="532">A. longulus</emphasis>
</taxonomicName>
, gill IV width/length ratio is 0.55; in 
<taxonomicName id="4C654D513D3B52095DB7FC7C3E8DFBE9" class="Insecta" family="Ameletidae" genus="Ameletus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="532" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B911EAC03D3B52095DB7FC7C3E8DFBE9" italics="true" pageId="13" pageNumber="532">A. formosus</emphasis>
</taxonomicName>
it is 0.47– 0.49 (Kang &amp; Yang 1994).
</paragraph>
</subSubSection>
</treatment>
</document>