treatments-xml/data/03/C2/87/03C287E8FFCDFFFCFF4BFF42FD52E667.xml

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<mods:title id="B7305D6E96070CB5A1CD7F1EB8610CD1">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<mods:namePart id="679646796FDB7AC657C136BE0F4F26EB">Calvo-Revuelta, Marta</mods:namePart>
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<treatment id="03C287E8FFCDFFFCFF4BFF42FD52E667" ID-DOI="http://doi.org/10.5281/zenodo.7293509" ID-GBIF-Taxon="204305439" ID-Zenodo-Dep="7293509" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667" httpUri="http://treatment.plazi.org/id/03C287E8FFCDFFFCFF4BFF42FD52E667" lastPageNumber="112" pageId="111" pageNumber="112" scope_family="Lamprophiidae" scope_order="Squamata">
<subSubSection id="C3716575FFCDFFFCFF4BFF42FD17E2CB" box="[151,674,223,250]" pageId="111" pageNumber="112" type="nomenclature">
<paragraph id="8BD436FEFFCDFFFCFF4BFF42FD17E2CB" blockId="111.[151,674,223,285]" box="[151,674,223,250]" pageId="111" pageNumber="112">
<heading id="D09C8192FFCDFFFCFF4BFF42FD17E2CB" bold="true" box="[151,674,223,250]" fontSize="11" level="2" pageId="111" pageNumber="112" reason="4">
<taxonomicName id="4C6B4D7DFFCDFFFCFF4BFF42FD17E2CB" ID-CoL="692K7" authority="Gunther, 1863" authorityName="Gunther" authorityYear="1863" box="[151,674,223,250]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B91FEAECFFCDFFFCFF4BFF42FD17E2CB" bold="true" box="[151,674,223,250]" pageId="111" pageNumber="112">
<emphasis id="B91FEAECFFCDFFFCFF4BFF42FE5DE2C8" bold="true" box="[151,488,223,250]" italics="true" pageId="111" pageNumber="112">Bothrophthalmus brunneus</emphasis>
Günther, 1863
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3716575FFCDFFFCFF4BFE9EFB52E352" pageId="111" pageNumber="112" type="description">
<paragraph id="8BD436FEFFCDFFFCFF4BFE9EFEADE32F" blockId="111.[151,674,223,285]" box="[151,280,259,285]" pageId="111" pageNumber="112">
<figureCitation id="13502A7BFFCDFFFCFF4BFE9EFEADE32F" box="[151,280,259,285]" captionStart="FIGURE 28" captionStartId="114.[151,250,1909,1933]" captionTargetBox="[163,1399,197,1879]" captionTargetId="figure-19@114.[154,1432,181,1885]" captionTargetPageId="114" captionText="FIGURE 28. Bothrophthalmus brunneus (A) (EBD 20922): from left to right: lateral, dorsal and ventral views of the head; Chamaelycus fasciatus (B) (EBD 27765) from left to right: complete specimen and dorsal view of the head, respectively; Gonionotophis brussauxi (C) (without EBD number, from Río Muni). Photos: AS-V." figureDoi="http://doi.org/10.5281/zenodo.7285709" httpUri="https://zenodo.org/record/7285709/files/figure.png" pageId="111" pageNumber="112">Figure 28A</figureCitation>
</paragraph>
<paragraph id="8BD436FEFFCDFFFCFF4BFED5FB52E352" blockId="111.[151,1255,327,352]" box="[151,1255,327,352]" pageId="111" pageNumber="112">
<taxonomicName id="4C6B4D7DFFCDFFFCFF4BFED5FD29E36D" authority="Gunther, 1863" authorityName="Gunther" authorityYear="1863" box="[151,668,327,352]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="subSpecies" species="lineatus" subSpecies="brunneus">
<emphasis id="B91FEAECFFCDFFFCFF4BFED5FDB4E352" box="[151,513,328,352]" italics="true" pageId="111" pageNumber="112">Bothrophthalmus lineatus brunneus</emphasis>
Günther, 1863
</taxonomicName>
. 
<typeStatus id="54D0885CFFCDFFFCFD7AFED5FD6DE352" box="[678,728,328,352]" pageId="111" pageNumber="112">Type</typeStatus>
locality: “Fernando Po”. 
<location id="8EB46025FFCDFFFCFC00FED5FBA9E352" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFC00FED5FBA9E352" box="[988,1052,328,352]" country="Equatorial Guinea" name="Bioko" pageId="111" pageNumber="112">Bioko</location>
, 
<collectingCountry id="F37C766EFFCDFFFCFBFAFED5FB57E352" box="[1062,1250,328,352]" name="Equatorial Guinea" pageId="111" pageNumber="112">Equatorial Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFCDFFFCFF4BFE10FD67E3C2" pageId="111" pageNumber="112" type="distribution">
<paragraph id="8BD436FEFFCDFFFCFF4BFE10FD67E3C2" blockId="111.[151,1437,397,1109]" pageId="111" pageNumber="112">
<emphasis id="B91FEAECFFCDFFFCFF4BFE10FE9CE395" bold="true" box="[151,297,397,423]" pageId="111" pageNumber="112">Distribution</emphasis>
. 
<collectingCountry id="F37C766EFFCDFFFCFEE4FE13FE05E39A" box="[312,432,398,424]" name="Cameroon" pageId="111" pageNumber="112">Cameroon</collectingCountry>
, 
<collectingCountry id="F37C766EFFCDFFFCFE67FE13FDBDE39A" box="[443,520,398,424]" name="Gabon" pageId="111" pageNumber="112">Gabon</collectingCountry>
, 
<collectingCountry id="F37C766EFFCDFFFCFDC9FE13FCA2E39A" box="[533,791,398,424]" name="Republic of the Congo" pageId="111" pageNumber="112">Republic of the Congo</collectingCountry>
, and 
<collectingCountry id="F37C766EFFCDFFFCFC88FE13FB90E39A" box="[852,1061,398,424]" name="Equatorial Guinea" pageId="111" pageNumber="112">Equatorial Guinea</collectingCountry>
. In 
<collectingCountry id="F37C766EFFCDFFFCFB92FE13FAA9E39A" box="[1102,1308,398,424]" name="Equatorial Guinea" pageId="111" pageNumber="112">Equatorial Guinea</collectingCountry>
it has been recorded both in Bioko (
<bibRefCitation id="EFFA4B0FFFCDFFFCFE73FE2FFDE8E3FE" author="Gunther" box="[431,605,434,460]" pageId="111" pageNumber="112" pagination="348 - 365" refId="ref78893" refString="Gunther. A. (1863 a) Third account of new species of snakes in the collection of the British Museum. Annals &amp; Magazine of Natural History, Series 3, 12 (71), 348 - 365. https: // doi. org / 10.1080 / 00222936308681536" type="journal article" year="1863">Günther 1863a</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFDB6FE2FFC97E3FE" author="Boulenger, G. A." box="[618,802,434,460]" pageId="111" pageNumber="112" refId="ref71723" refString="Boulenger, G. A. (1893) Catalogue of the Snakes in the British Museum (Natural History). Vol. 1. Trustees, London, xiii + 448 pp., 28 pls." type="book" year="1893">Boulenger 1893</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFCF3FE2FFC72E3FE" author="Bocage, J. V. B." box="[815,967,434,460]" pageId="111" pageNumber="112" pagination="25 - 59" refId="ref71106" refString="Bocage, J. V. B. (1903 a) Contribution a la faune des quatre iles du Golfe de Guinee. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 7, 25 - 59." type="journal article" year="1903">Bocage 1903</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFC08FE2FFBCBE3FE" author="Mertens, R." box="[980,1150,434,460]" pageId="111" pageNumber="112" pagination="211 - 238" refId="ref84765" refString="Mertens, R. (1964 a) Die Reptilien von Fernando Poo. Bonner zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens 1964a</bibRefCitation>
) and 
<collectingCountry id="F37C766EFFCDFFFCFB1CFE2FFA9BE3FE" box="[1216,1326,434,460]" name="Equatorial Guinea" pageId="111" pageNumber="112">Río Muni</collectingCountry>
at Monte Alén National Park (
<bibRefCitation id="EFFA4B0FFFCDFFFCFEA2FE4BFDF4E3C2" author="Lasso, C. A. &amp; Rial, A. I. &amp; Castroviejo, J. &amp; De la Riva, I." box="[382,577,470,496]" pageId="111" pageNumber="112" pagination="21 - 34" refId="ref81678" refString="Lasso, C. A., Rial, A. I., Castroviejo, J. &amp; De la Riva, I. (2002) Herpetofauna del Parque Nacional de Monte Alen (Rio Muni, Guinea Ecuatorial). Graellsia, 58, 21 - 34. https: // doi. org / 10.3989 / graellsia. 2002. v 58. i 2.276" type="journal article" year="2002">
Lasso 
<emphasis id="B91FEAECFFCDFFFCFE14FE4AFDB5E3C2" box="[456,512,470,496]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2002
</bibRefCitation>
) (
<figureCitation id="13502A7BFFCDFFFCFD8BFE4BFD70E3C2" box="[599,709,470,496]" captionStart="MAPS 31" captionStartId="110.[151,223,1928,1952]" captionTargetBox="[207,1379,194,1894]" captionTargetId="figure-19@110.[165,1422,181,1905]" captionTargetPageId="110" captionText="MAPS 31 A–C. Distribution maps for Equatorial Guinean records of (A) Boaedon virgatus; (B) Bothrolycus ater (question mark represents an imprecise record); (C) Bothrophthalmus brunneus (question mark represents an imprecise record)." figureDoi="http://doi.org/10.5281/zenodo.7305189" httpUri="https://zenodo.org/record/7305189/files/figure.png" pageId="111" pageNumber="112">Map 31C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFCDFFFCFF1BFE64FD52E667" pageId="111" pageNumber="112" type="discussion">
<paragraph id="8BD436FEFFCDFFFCFF1BFE64FC8FE1F6" blockId="111.[151,1437,397,1109]" pageId="111" pageNumber="112">
<emphasis id="B91FEAECFFCDFFFCFF1BFE64FEFAE021" bold="true" box="[199,335,505,531]" pageId="111" pageNumber="112">Comments.</emphasis>
There are two species of 
<taxonomicName id="4C6B4D7DFFCDFFFCFDB4FE67FCAEE026" box="[616,795,506,532]" class="Reptilia" family="Lamprophiidae" genus="Bothropthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="genus">
<emphasis id="B91FEAECFFCDFFFCFDB4FE67FCAEE026" box="[616,795,506,532]" italics="true" pageId="111" pageNumber="112">Bothropthalmus</emphasis>
</taxonomicName>
currently recognized, namely 
<taxonomicName id="4C6B4D7DFFCDFFFCFBB0FE66FB44E026" authorityName="Gunther" authorityYear="1863" box="[1132,1265,506,532]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B91FEAECFFCDFFFCFBB0FE66FB44E026" box="[1132,1265,506,532]" italics="true" pageId="111" pageNumber="112">B. brunneus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6B4D7DFFCDFFFCFAFAFE66FE93E00A" authority="Peters, 1863" authorityName="Peters" authorityYear="1863" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91FEAECFFCDFFFCFAFAFE66FA28E026" box="[1318,1437,506,532]" italics="true" pageId="111" pageNumber="112">B. lineatus</emphasis>
Peters, 1863
</taxonomicName>
. The taxonomic status of 
<taxonomicName id="4C6B4D7DFFCDFFFCFD91FD82FD63E00A" authorityName="Gunther" authorityYear="1863" box="[589,726,542,568]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B91FEAECFFCDFFFCFD91FD82FD63E00A" box="[589,726,542,568]" italics="true" pageId="111" pageNumber="112">B. brunneus</emphasis>
</taxonomicName>
is controversial. The two species are distinguishable based on colouration only: 
<taxonomicName id="4C6B4D7DFFCDFFFCFE80FDDEFE57E06E" authorityName="Gunther" authorityYear="1863" box="[348,482,578,604]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B91FEAECFFCDFFFCFE80FDDEFE57E06E" box="[348,482,578,604]" italics="true" pageId="111" pageNumber="112">B. brunneus</emphasis>
</taxonomicName>
presents a uniformly dark body pattern, whereas 
<taxonomicName id="4C6B4D7DFFCDFFFCFBD9FDDEFBC9E06E" authorityName="Peters" authorityYear="1863" box="[1029,1148,578,604]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91FEAECFFCDFFFCFBD9FDDEFBC9E06E" box="[1029,1148,578,604]" italics="true" pageId="111" pageNumber="112">B. lineatus</emphasis>
</taxonomicName>
shows a red-black striped pattern (
<bibRefCitation id="EFFA4B0FFFCDFFFCFF2FFDFBFE20E0B2" author="Schmidt, K. P." box="[243,405,614,640]" pageId="111" pageNumber="112" pagination="1 - 148" refId="ref88767" refString="Schmidt, K. P. (1923) Contributions to the herpetology of Belgian Congo based on the collections of the American Museum Congo Expedition, 1905 - 1915. Part II. Snakes; with field notes by Herbert Lang and James P. Chapin. Bulletin of the American Museum of Natural History, 49, 1 - 148." type="journal article" year="1923">Schmidt 1923</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFE7DFDFBFD7BE0B2" author="Chippaux, J. P. &amp; Jackson, K." box="[417,718,614,640]" pageId="111" pageNumber="112" refId="ref74358" refString="Chippaux, J. P. &amp; Jackson, K. (2019) Snakes of Central and Western Africa. Johns Hopkins University Press, Baltimore, Maryland, 429 pp." type="book" year="2019">Chippaux &amp; Jackson 2019</bibRefCitation>
). 
<bibRefCitation id="EFFA4B0FFFCDFFFCFD3CFDFBFC12E0B2" author="Boulenger, G. A." box="[736,935,614,640]" pageId="111" pageNumber="112" refId="ref71723" refString="Boulenger, G. A. (1893) Catalogue of the Snakes in the British Museum (Natural History). Vol. 1. Trustees, London, xiii + 448 pp., 28 pls." type="book" year="1893">Boulenger (1893)</bibRefCitation>
included 
<taxonomicName id="4C6B4D7DFFCDFFFCFBC9FDFAFB28E0B2" authorityName="Gunther" authorityYear="1863" box="[1045,1181,614,640]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B91FEAECFFCDFFFCFBC9FDFAFB28E0B2" box="[1045,1181,614,640]" italics="true" pageId="111" pageNumber="112">B. brunneus</emphasis>
</taxonomicName>
in the synonymy of 
<taxonomicName id="4C6B4D7DFFCDFFFCFA58FDFAFF45E096" authorityName="Peters" authorityYear="1863" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91FEAECFFCDFFFCFA58FDFAFF45E096" italics="true" pageId="111" pageNumber="112">B. lineatus</emphasis>
</taxonomicName>
, treating Günther’s taxon as a variety of the latter. Other authors have considered this taxon as a subspecies of 
<taxonomicName id="4C6B4D7DFFCDFFFCFF69FD32FE99E0FA" authorityName="Peters" authorityYear="1863" box="[181,300,686,712]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91FEAECFFCDFFFCFF69FD32FE99E0FA" box="[181,300,686,712]" italics="true" pageId="111" pageNumber="112">B. lineatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFA4B0FFFCDFFFCFEE5FD33FE6CE0FA" author="Schmidt, K. P." box="[313,473,686,712]" pageId="111" pageNumber="112" pagination="1 - 148" refId="ref88767" refString="Schmidt, K. P. (1923) Contributions to the herpetology of Belgian Congo based on the collections of the American Museum Congo Expedition, 1905 - 1915. Part II. Snakes; with field notes by Herbert Lang and James P. Chapin. Bulletin of the American Museum of Natural History, 49, 1 - 148." type="journal article" year="1923">Schmidt 1923</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFE38FD33FD74E0FA" author="Pauwels, O. S. G. &amp; Kamdem Toham, A. &amp; Chimsunchart, C." box="[484,705,686,712]" pageId="111" pageNumber="112" pagination="47 - 57" refId="ref85753" refString="Pauwels, O. S. G., Kamdem Toham, A. &amp; Chimsunchart, C. (2002) Recherches sur l'herpetofaune du Massif du Chaillu, Gabon. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 72, 47 - 57." type="journal article" year="2002">
Pauwels 
<emphasis id="B91FEAECFFCDFFFCFD94FD32FDCAE0FA" box="[584,639,686,712]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2002
</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFD17FD33FCCDE0FA" author="Chippaux, J. - P." box="[715,888,686,712]" pageId="111" pageNumber="112" refId="ref74326" refString="Chippaux, J. - P. (2006) Les Serpents d'Afrique Occidentale et Centrale. Institut de Recherche pour le Developpement (IRD), Paris, 311 pp." type="book" year="2006">Chippaux 2006</bibRefCitation>
) or treated it with full species status (
<bibRefCitation id="EFFA4B0FFFCDFFFCFAC8FD33FA22E0FB" author="Trape, J. - F." box="[1300,1431,686,713]" pageId="111" pageNumber="112" pagination="135 - 140" refId="ref90979" refString="Trape, J. - F. (1985) Les serpents de la region de Dimonika (Mayombe), Republique Populaire du Congo. Revue de Zoologie Africaine, 99 (2), 135 - 140." type="journal article" year="1985">Trape 1985</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFF4BFD4FFE96E0DE" author="Meirte, D." box="[151,291,722,748]" pageId="111" pageNumber="112" pagination="1 - 161" refId="ref84512" refString="Meirte, D. (1992) Cles de determination des serpents d'Afrique. Annales du Musee Royal de l'Afrique Centrale. Sciences zoologiques, 267, 1 - 161." type="journal article" year="1992">Meirte 1992</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFEF2FD4FFDF4E0DE" author="Pauwels, O. S. G. &amp; David, P." box="[302,577,722,748]" pageId="111" pageNumber="112" pagination="13 - 18" refId="ref85642" refString="Pauwels, O. S. G. &amp; David, P. (2008 a) Miscellanea herpetologica Gabonica I. Hamadryad, 32 (1), 13 - 18." type="journal article" year="2008">Pauwels &amp; David 2008a</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFD97FD4FFC92E0DE" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[587,807,722,748]" pageId="111" pageNumber="112" refId="ref91877" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the world: a catalogue of living and extinct species. CRC Press, Boca Raton, Florida, 1237 pp. https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
Wallach 
<emphasis id="B91FEAECFFCDFFFCFD70FD4EFD51E0DE" box="[684,740,722,748]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2014
</bibRefCitation>
). Recent molecular works included a single sample of 
<taxonomicName id="4C6B4D7DFFCDFFFCFA58FD4EFF45E122" authorityName="Peters" authorityYear="1863" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91FEAECFFCDFFFCFA58FD4EFF45E122" italics="true" pageId="111" pageNumber="112">B. lineatus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F37C766EFFCDFFFCFEE9FD6BFE39E122" box="[309,396,758,784]" name="Uganda" pageId="111" pageNumber="112">Uganda</collectingCountry>
and another one of 
<taxonomicName id="4C6B4D7DFFCDFFFCFDB0FD6AFD46E122" authorityName="Gunther" authorityYear="1863" box="[620,755,758,784]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B91FEAECFFCDFFFCFDB0FD6AFD46E122" box="[620,755,758,784]" italics="true" pageId="111" pageNumber="112">B. brunneus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F37C766EFFCDFFFCFCE4FD6BFC30E122" box="[824,901,758,784]" name="Gabon" pageId="111" pageNumber="112">Gabon</collectingCountry>
, supporting them with full species status (
<bibRefCitation id="EFFA4B0FFFCDFFFCFA80FD6BFEA5E106" author="Kelly, C. M. &amp; Branch, W. R. &amp; Broadley, D. G. &amp; Barker, N. P. &amp; Villet, M. H." pageId="111" pageNumber="112" pagination="415 - 426" refId="ref81125" refString="Kelly, C. M., Branch, W. R., Broadley, D. G., Barker, N. P. &amp; Villet, M. H. (2011) Molecular systematics of the African snake family Lamprophiidae Fitzinger, 1843 (Serpentes: Elapoidea), with particular focus on the genera Lamprophis Fitzinger 1843 and Mehelya Csiki 1903. Molecular Phylogenetics and Evolution, 58 (3), 415 - 426. https: // doi. org / 10.1016 / j. ympev. 2010.11.010" type="journal article" year="2011">
Kelly 
<emphasis id="B91FEAECFFCDFFFCFF4BFC86FF65E106" box="[151,208,794,820]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2011
</bibRefCitation>
; 
<bibRefCitation id="EFFA4B0FFFCDFFFCFEC0FC87FE54E106" author="Zaher, H. &amp; Murphy, R. W. &amp; Arredondo, J. C. &amp; Graboski, R. &amp; Machado-Filho, P. R. &amp; Mahlow, K. &amp; Montingelli, G. G. &amp; Quadros, A. B. &amp; Orlov, N. L. &amp; Wilkinson, M. &amp; Zhang, Y. P. &amp; Grazziotin, F. G." box="[284,481,794,820]" pageId="111" pageNumber="112" pagination="0216148" refId="ref92693" refString="Zaher, H., Murphy, R. W., Arredondo, J. C., Graboski, R., Machado-Filho, P. R., Mahlow, K., Montingelli, G. G., Quadros, A. B., Orlov, N. L., Wilkinson, M., Zhang, Y. P. &amp; Grazziotin, F. G. (2019) Large-scale molecular phylogeny, morphology, divergence-time estimation, and the fossil record of advanced caenophidian snakes (Squamata: Serpentes). PLoS ONE, 14 (5), e 0216148. https: // doi. org / 10.1371 / journal. pone. 0216148" type="journal article" year="2019">
Zaher 
<emphasis id="B91FEAECFFCDFFFCFEB9FC86FE28E106" box="[357,413,794,820]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2019
</bibRefCitation>
). However, in order to resolve this taxonomic problem, it will be required to improve the sampling for molecular analyses across most of the range of both species, including their 
<typeStatus id="54D0885CFFCDFFFCFB15FCA3FB4CE16A" box="[1225,1273,830,856]" pageId="111" pageNumber="112">type</typeStatus>
localities [the 
<typeStatus id="54D0885CFFCDFFFCFF4BFCFFFF72E14E" box="[151,199,866,892]" pageId="111" pageNumber="112">type</typeStatus>
locality of 
<taxonomicName id="4C6B4D7DFFCDFFFCFE9BFCFEFE0BE14E" authorityName="Peters" authorityYear="1863" box="[327,446,866,892]" class="Reptilia" family="Lamprophiidae" genus="Bothrophthalmus" kingdom="Animalia" order="Squamata" pageId="111" pageNumber="112" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91FEAECFFCDFFFCFE9BFCFEFE0BE14E" box="[327,446,866,892]" italics="true" pageId="111" pageNumber="112">B. lineatus</emphasis>
</taxonomicName>
is the Coast of the Gulf of 
<collectingCountry id="F37C766EFFCDFFFCFD30FCFFFCF5E14E" box="[748,832,866,892]" name="Guinea-Bissau" pageId="111" pageNumber="112">Guinea</collectingCountry>
, but 
<emphasis id="B91FEAECFFCDFFFCFCA9FCFFFC2AE14E" box="[885,927,866,892]" italics="true" pageId="111" pageNumber="112">fide</emphasis>
<bibRefCitation id="EFFA4B0FFFCDFFFCFC7AFCFFFB05E14E" author="Hughes, B. &amp; Barry, D. H." box="[934,1200,866,892]" pageId="111" pageNumber="112" pagination="1004 - 1041" refId="ref80167" refString="Hughes, B. &amp; Barry, D. H. (1969) The snakes of Ghana, a checklist and key. Bulletin de l'Institut Fondamental d'Afrique Noire, 31 (3), 1004 - 1041." type="journal article" year="1969">Hughes &amp; Barry (1969)</bibRefCitation>
the true 
<typeStatus id="54D0885CFFCDFFFCFACDFCFFFAF4E14E" box="[1297,1345,866,892]" pageId="111" pageNumber="112">type</typeStatus>
locality is likely in 
<collectingCountry id="F37C766EFFCDFFFCFEC8FC1BFED6E192" box="[276,355,902,928]" name="Ghana" pageId="111" pageNumber="112">Ghana</collectingCountry>
]. There are no precise records of this species in Bioko (
<bibRefCitation id="EFFA4B0FFFCDFFFCFC11FC1BFBCFE192" author="Mertens, R." box="[973,1146,902,928]" pageId="111" pageNumber="112" pagination="211 - 238" refId="ref84765" refString="Mertens, R. (1964 a) Die Reptilien von Fernando Poo. Bonner zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens 1964a</bibRefCitation>
). The examined specimen EBD 20925 contained an unidentified mouse in its stomach.
</paragraph>
<paragraph id="8BD436FEFFCDFFFCFF1BFC50FD52E667" blockId="111.[151,1437,397,1109]" pageId="111" pageNumber="112">
<emphasis id="B91FEAECFFCDFFFCFF1BFC50FE70E1D5" bold="true" box="[199,453,973,999]" pageId="111" pageNumber="112">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FFCDFFFCFE17FC53FB09E1DA" ID-GBIF-Occurrence="3962324640" accessionNumber="EBD 20923" box="[459,1212,974,1000]" collectingDate="1986-04-16" collectionCode="EBD" country="Equatorial Guinea" location="Eight" municipality="Miboman" pageId="111" pageNumber="112" specimenCount="1">
<location id="8EB46025FFCDFFFCFE17FC53FDBDE1DA" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFE17FC53FDBDE1DA" box="[459,520,974,1000]" country="Equatorial Guinea" municipality="Miboman" name="Eight" pageId="111" pageNumber="112">Eight</location>
specimens. 
<collectingCountry id="F37C766EFFCDFFFCFD4CFC53FD4AE1DA" box="[656,767,974,1000]" name="Equatorial Guinea" pageId="111" pageNumber="112">Río Muni</collectingCountry>
: 
<collectingMunicipality id="6BB0AC84FFCDFFFCFCD5FC53FCCCE1DA" box="[777,889,974,1000]" pageId="111" pageNumber="112">Miboman</collectingMunicipality>
, 
<date id="FFD5103EFFCDFFFCFC5FFC53FBA9E1DA" box="[899,1052,974,1000]" pageId="111" pageNumber="112" value="1986-04-16">
<collectingDate id="EF91E9D6FFCDFFFCFC5FFC53FBA9E1DA" box="[899,1052,974,1000]" pageId="111" pageNumber="112" value="1986-04-16">16 April 1986</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFCDFFFCFBF6FC53FBD1E1DA" box="[1066,1124,974,1000]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="111" pageNumber="112">EBD</collectionCode>
20923)
</materialsCitation>
, 
<materialsCitation id="3B033CA3FFCDFFFCFB1AFC53FE8AE63F" ID-GBIF-Occurrence="3962324329" accessionNumber="EBD 20924" collectingDate="1986-07-08" collectionCode="EBD" country="Equatorial Guinea" isTailEnumeration="true" location="Eight" municipality="Miboman" pageId="111" pageNumber="112" specimenCode="EBD 20924-20925" specimenCount="1">
<date id="FFD5103EFFCDFFFCFB1AFC53FAE0E1DA" box="[1222,1365,974,1000]" pageId="111" pageNumber="112" value="1986-07-08">
<collectingDate id="EF91E9D6FFCDFFFCFB1AFC53FAE0E1DA" box="[1222,1365,974,1000]" pageId="111" pageNumber="112" value="1986-07-08">08 July 1986</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FFCDFFFCFABFFC53FE82E63F" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="111" pageNumber="112">EBD 20924–20925</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B033CA3FFCDFFFCFE94FC6FFC56E63E" ID-GBIF-Occurrence="3962324378" accessionNumber="EBD 20922" box="[328,995,1010,1036]" collectingDate="1986-10-14" collectionCode="EBD" country="Equatorial Guinea" location="Miboman" municipality="Bata" pageId="111" pageNumber="112" specimenCount="1">
<location id="8EB46025FFCDFFFCFE94FC6FFE0DE63E" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFE94FC6FFE0DE63E" box="[328,440,1010,1036]" country="Equatorial Guinea" municipality="Bata" name="Miboman" pageId="111" pageNumber="112">Miboman</location>
, 
<quantity id="4C939B1BFFCDFFFCFE1EFC6FFDBDE63E" box="[450,520,1010,1036]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="111" pageNumber="112" unit="km" value="30.0">30 km</quantity>
from 
<collectingMunicipality id="6BB0AC84FFCDFFFCFD95FC6FFDCAE63E" box="[585,639,1010,1036]" pageId="111" pageNumber="112">Bata</collectingMunicipality>
, 
<date id="FFD5103EFFCDFFFCFD55FC6FFCF6E63E" box="[649,835,1010,1036]" pageId="111" pageNumber="112" value="1986-10-14">
<collectingDate id="EF91E9D6FFCDFFFCFD55FC6FFCF6E63E" box="[649,835,1010,1036]" pageId="111" pageNumber="112" value="1986-10-14">14 October 1986</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFCDFFFCFC8DFC6FFC3EE63E" box="[849,907,1010,1036]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="111" pageNumber="112">EBD</collectionCode>
20922)
</materialsCitation>
; 
<materialsCitation id="3B033CA3FFCDFFFCFC31FC6FFE72E603" ID-GBIF-Occurrence="3962324616" accessionNumber="EBD 27514" collectingDate="1987-08" collectionCode="EBD" country="Equatorial Guinea" county="Niefang" location="Miboman" municipality="Bata" pageId="111" pageNumber="112" specimenCount="1">
<location id="8EB46025FFCDFFFCFC31FC6FFBEFE63E" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFC31FC6FFBEFE63E" box="[1005,1114,1010,1036]" country="Equatorial Guinea" county="Niefang" municipality="Bata" name="Miboman" pageId="111" pageNumber="112">Miboman</location>
(road from 
<collectingMunicipality id="6BB0AC84FFCDFFFCFB01FC6FFAA5E63E" box="[1245,1296,1010,1036]" pageId="111" pageNumber="112">Bata</collectingMunicipality>
to 
<collectingCounty id="62B54E72FFCDFFFCFAEEFC6FFA24E63E" box="[1330,1425,1010,1036]" pageId="111" pageNumber="112">Niefang</collectingCounty>
), 
<date id="FFD5103EFFCDFFFCFF4BFB8BFE93E602" box="[151,294,1046,1072]" pageId="111" pageNumber="112" value="1987-08">
<collectingDate id="EF91E9D6FFCDFFFCFF4BFB8BFE93E602" box="[151,294,1046,1072]" pageId="111" pageNumber="112" value="1987-08">August 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFCDFFFCFEE8FB8BFEDBE602" box="[308,366,1046,1072]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="111" pageNumber="112">EBD</collectionCode>
27514)
</materialsCitation>
; 
<materialsCitation id="3B033CA3FFCDFFFCFE0DFB8BFABFE602" ID-GBIF-Occurrence="3962324392" accessionNumber="EBD 18600" box="[465,1290,1046,1072]" collectingDate="1984-06-19" collectionCode="EBD" country="Equatorial Guinea" county="Km" location="Ncoeakien" municipality="Movo" pageId="111" pageNumber="112" specimenCount="1">
<location id="8EB46025FFCDFFFCFE0DFB8BFDFCE602" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFE0DFB8BFDFCE602" box="[465,585,1046,1072]" country="Equatorial Guinea" county="Km" municipality="Movo" name="Ncoeakien" pageId="111" pageNumber="112">Ncoeakieñ</location>
(road from 
<location id="8EB46025FFCDFFFCFD11FB8BFCB5E602" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFD11FB8BFCB5E602" box="[717,768,1046,1072]" country="Equatorial Guinea" county="Km" municipality="Movo" name="Bata" pageId="111" pageNumber="112">Bata</location>
to 
<collectingMunicipality id="6BB0AC84FFCDFFFCFCFFFB8BFCDCE602" box="[803,873,1046,1072]" pageId="111" pageNumber="112">Movo</collectingMunicipality>
, 
<collectingCounty id="62B54E72FFCDFFFCFCAFFB8BFC28E602" box="[883,925,1046,1072]" pageId="111" pageNumber="112">Km</collectingCounty>
23), 
<date id="FFD5103EFFCDFFFCFC0AFB8BFBDFE602" box="[982,1130,1046,1072]" pageId="111" pageNumber="112" value="1984-06-19">
<collectingDate id="EF91E9D6FFCDFFFCFC0AFB8BFBDFE602" box="[982,1130,1046,1072]" pageId="111" pageNumber="112" value="1984-06-19">19 June 1984</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFCDFFFCFBA4FB8BFB07E602" box="[1144,1202,1046,1072]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="111" pageNumber="112">EBD</collectionCode>
18600)
</materialsCitation>
; 
<materialsCitation id="3B033CA3FFCDFFFCFAC9FB8BFD56E667" ID-GBIF-Occurrence="3962324315" accessionNumber="EBD 25132" collectingDate="1986-08-01" collectionCode="EBD" country="Equatorial Guinea" location="San Joaquin de Nydiacom" pageId="111" pageNumber="112" specimenCode="EBD 25132-25133" specimenCount="1">
<location id="8EB46025FFCDFFFCFAC9FB8BFE84E666" LSID="urn:lsid:plazi:treatment:03C287E8FFCDFFFCFF4BFF42FD52E667:8EB46025FFCDFFFCFAC9FB8BFE84E666" country="Equatorial Guinea" name="San Joaquin de Nydiacom" pageId="111" pageNumber="112">San Joaquín de Nydiacom</location>
, 
<date id="FFD5103EFFCDFFFCFEE7FBA7FE58E666" box="[315,493,1082,1108]" pageId="111" pageNumber="112" value="1986-08-01">
<collectingDate id="EF91E9D6FFCDFFFCFEE7FBA7FE58E666" box="[315,493,1082,1108]" pageId="111" pageNumber="112" value="1986-08-01">01 August 1986</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FFCDFFFCFE27FBA7FD69E667" box="[507,732,1082,1109]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="111" pageNumber="112">EBD 25132–25133</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>