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<document id="32D5E45FFED271DF59B0DF2DE3A67D8E" ID-CLB-Dataset="7809" ID-DOI="10.5281/zenodo.4503519" ID-GBIF-Dataset="8713f3de-67ef-4823-929b-7b1b5d37e26a" ID-ISSN="0374-1036" ID-Zenodo-Dep="4503519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612461365170" checkinUser="felipe" docAuthor="Kočárek, Petr" docDate="2006" docId="A1441D06BF470302FE1BFD64FD3CFAFF" docLanguage="en" docName="ActaEntMusNatPra.46.1-6.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 46" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Nala alenae Kočárek 2006, sp. nov." docType="treatment" docUuid="183A10DA-1A4C-4AD6-AD47-FA9D76FB1634" docUuidSource="ZooBank" docVersion="10" lastPageNumber="4" masterDocId="5D7D657EBF460301FFF2FFDEFFBFFFEA" masterDocTitle="A new species of Nala (Dermaptera: Labiduridae) from Cambodia" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1698927097416" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="4782022BDF968551A8F40C49A4501CFC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FFC8F0854390D8540795C96D2946C153">
<mods:title id="AEAD51B9558691D971CA144DFF7CEFEA">A new species of Nala (Dermaptera: Labiduridae) from Cambodia</mods:title>
</mods:titleInfo>
<mods:name id="0E5E68C3DD6853DE69DCE39F4A96E61B" type="personal">
<mods:role id="DB5370B13D1A1E260B30CD1E16481B20">
<mods:roleTerm id="A2C25B798CB3EEE184C25160491A8C6F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="316C6178F34BE165EEB848C2CD3443C4">Kočárek, Petr</mods:namePart>
</mods:name>
<mods:typeOfResource id="E70AF12422ABBB4079EE3734A13F7A41">text</mods:typeOfResource>
<mods:relatedItem id="B1E83CBB8D85D03B2BB29EA10886F868" type="host">
<mods:titleInfo id="9D7BD9DC8903E622B312C237FED9D6D0">
<mods:title id="ADEFA5640BCB49546865AED069EC8FD1">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="3A0C63FE25D693853300C73B3F2A6692">
<mods:date id="13DF66349DAFBE2FE4398536FA32843B">2006</mods:date>
<mods:detail id="9816CCFA58EEFA8D922C9A3E4D84D01D" type="pubDate">
<mods:number id="2AF2EA00572071379FDC1D9EEC09742C">2006-11-06</mods:number>
</mods:detail>
<mods:detail id="C13B9D400F557B9F6BADFF52A77D3317" type="volume">
<mods:number id="D2A57F94F20B221A105489ACE4E931C5">46</mods:number>
</mods:detail>
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<mods:start id="9AC69A3B79D0C166020CB9B278FF8994">1</mods:start>
<mods:end id="8CA04A0EB65B7C82103B5F45663610FC">6</mods:end>
</mods:extent>
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<mods:classification id="49407FE4FD88846890CF6968C736A9F2">journal article</mods:classification>
<mods:identifier id="FC8199E730422ED33DC541FD355C672B" type="CLB-Dataset">7809</mods:identifier>
<mods:identifier id="A29715F4A626F25159B9F192624D53DC" type="DOI">10.5281/zenodo.4503519</mods:identifier>
<mods:identifier id="A8FACD86538170B667C3A9F41263E94A" type="GBIF-Dataset">8713f3de-67ef-4823-929b-7b1b5d37e26a</mods:identifier>
<mods:identifier id="8A19BDB8CD65C3A14C1DFF7EF40B3E58" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="02B3736030C2F7E35702276C8C7798E8" type="Zenodo-Dep">4503519</mods:identifier>
<mods:identifier id="CEBF717BF28EACD2BE60736C4778555B" type="ZooBank">1C364620-9300-4F01-9662-34A5564229F7</mods:identifier>
</mods:mods>
<treatment id="A1441D06BF470302FE1BFD64FD3CFAFF" ID-DOI="http://doi.org/10.5281/zenodo.4594241" ID-GBIF-Taxon="180302136" ID-Zenodo-Dep="4594241" LSID="urn:lsid:zoobank.org:act:183A10DA-1A4C-4AD6-AD47-FA9D76FB1634" httpUri="http://treatment.plazi.org/id/A1441D06BF470302FE1BFD64FD3CFAFF" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection id="61F7FF9BBF470300FE1BFD64FD75FD38" box="[489,714,698,722]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="2952AC10BF470300FE1BFD64FD75FD38" blockId="1.[489,714,698,722]" box="[489,714,698,722]" pageId="1" pageNumber="2">
<heading id="721A1B7CBF470300FE1BFD64FD75FD38" bold="true" box="[489,714,698,722]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="2" reason="3">
<emphasis id="1B997002BF470300FE1BFD64FD75FD38" bold="true" box="[489,714,698,722]" pageId="1" pageNumber="2">
<taxonomicName id="EEEDD793BF470300FE1BFD64FDD2FD38" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[489,621,698,722]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF470300FE1BFD64FDD2FD38" bold="true" box="[489,621,698,722]" italics="true" pageId="1" pageNumber="2">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF470300FD86FD64FD75FD38" box="[628,714,698,722]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF470300FDDEFD04FD39FD07" box="[556,646,730,749]" pageId="1" pageNumber="2" type="description">
<paragraph id="2952AC10BF470300FDDEFD04FD39FD07" blockId="1.[556,646,730,749]" box="[556,646,730,749]" pageId="1" pageNumber="2">
(
<figureCitation id="B1D6B095BF470300FDC0FD04FDC0FD07" box="[562,639,730,749]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="1" pageNumber="2">Figs. 1-5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF470300FF83FCCEFD12FC99" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="2952AC10BF470300FF83FCCEFDD3FCC9" blockId="1.[113,1089,784,883]" box="[113,620,784,803]" pageId="1" pageNumber="2">
<emphasis id="1B997002BF470300FF83FCCEFF54FCC9" bold="true" box="[113,235,784,803]" pageId="1" pageNumber="2">
<typeStatus id="F65612B2BF470300FF83FCCEFF22FCC9" box="[113,157,784,803]" pageId="1" pageNumber="2">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="51FAEC80BF470300FF03FCCEFEF5FCC9" box="[241,330,784,803]" name="Cambodia" pageId="1" pageNumber="2">Cambodia</collectingCountry>
,
<collectingRegion id="EB2962F2BF470300FEA6FCCEFE0CFCC9" box="[340,435,784,803]" country="Cambodia" name="Siem Reap" pageId="1" pageNumber="2">Siem Reap</collectingRegion>
, Angkor Wat temple.
</paragraph>
<paragraph id="2952AC10BF470300FF83FCF4FD12FC99" blockId="1.[113,1089,784,883]" pageId="1" pageNumber="2">
<emphasis id="1B997002BF470300FF83FCF4FF4BFCD7" bold="true" box="[113,244,810,829]" pageId="1" pageNumber="2">Type material.</emphasis>
<materialsCitation id="9985A64DBF470300FF0BFCF5FDCAFCB2" ID-GBIF-Occurrence="3047787301" collectingDate="2002-09-07" collectingDateMax="2002-09-11" collectingDateMin="2002-09-07" collectionCode="MMBC" collectorName="P. Kocarek" country="Cambodia" elevation="40" latitude="13.41173" location="Angkor Wat" longLatPrecision="1" longitude="103.863846" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Siem Reap" typeStatus="holotype">
<typeStatus id="F65612B2BF470300FF0BFCF5FEF4FCD6" box="[249,331,811,830]" pageId="1" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
: ♁, ‘NW
<collectingCountry id="51FAEC80BF470300FE6CFCF4FDACFCD7" box="[414,531,810,829]" name="Cambodia" pageId="1" pageNumber="2">CAMBODIA</collectingCountry>
,
<collectingRegion id="EB2962F2BF470300FDE9FCF4FDC8FCD4" box="[539,631,810,830]" country="Cambodia" name="Siem Reap" pageId="1" pageNumber="2">Siem Reap</collectingRegion>
env. /
<location id="2C32FACBBF470300FD5EFCF4FCACFCD4" LSID="urn:lsid:plazi:treatment:A1441D06BF470302FE1BFD64FD3CFAFF:2C32FACBBF470300FD5EFCF4FCACFCD4" box="[684,787,810,830]" country="Cambodia" latitude="13.41173" longLatPrecision="1" longitude="103.863846" name="Angkor Wat" pageId="1" pageNumber="2" stateProvince="Siem Reap">Angkor Wat</location>
temple;
<elevation id="A2C04B23BF470300FCAEFCF4FC0DFCD7" box="[860,946,810,830]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="2" unit="m" value="40.0">
<quantity id="EE1501F5BF470300FCAEFCF4FC39FCD4" box="[860,902,810,830]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="2" unit="m" value="40.0">40 m</quantity>
a.s.l.
</elevation>
/
<geoCoordinate id="4CD9CAD7BF470300FC4EFCF4FB82FCD4" box="[956,1085,809,830]" degrees="13" direction="north" minutes="24" orientation="latitude" pageId="1" pageNumber="2" precision="1" seconds="42.23" value="13.41173">13°24′42.23″N</geoCoordinate>
,
<geoCoordinate id="4CD9CAD7BF470300FF83FC9BFF45FCB3" box="[113,250,836,857]" degrees="103" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="2" precision="1" seconds="49.84" value="103.863846">103°51′49.84″E</geoCoordinate>
/
<collectorName id="8418C9C6BF470300FEFDFC98FED0FCB2" box="[271,367,837,857]" pageId="1" pageNumber="2">P. Kočárek</collectorName>
leg.;
<date id="5D538AD0BF470300FE51FC98FDA8FCB2" box="[419,535,837,857]" pageId="1" pageNumber="2" value="2002-09-07" valueMax="2002-09-11" valueMin="2002-09-07">
<collectingDate id="4D177338BF470300FE51FC98FDA8FCB2" box="[419,535,837,857]" pageId="1" pageNumber="2" value="2002-09-07" valueMax="2002-09-11" valueMin="2002-09-07">7.-11.ix.2002</collectingDate>
</date>
’ (
<collectionCode id="4FFC34D5BF470300FDDAFC9BFDD0FCB2" box="[552,623,837,856]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="2">MMBC</collectionCode>
)
</materialsCitation>
;
<typeStatus id="F65612B2BF470300FD72FC98FD69FCB3" box="[640,726,838,857]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FEB6799BF470300FD12FC9BFCB4FCB3" box="[736,779,833,857]" pageId="1" pageNumber="2" type="female">5 ♀♀</specimenCount>
, same label data as
<typeStatus id="F65612B2BF470300FC3BFC9BFBACFCB2" box="[969,1043,837,856]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
(one specimen
<collectionCode id="4FFC34D5BF470300FF3AFCBEFEB3FC99" box="[200,268,864,883]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="2">MMBC</collectionCode>
, one specimen
<collectionCode id="4FFC34D5BF470300FE63FCBEFE71FC99" box="[401,462,864,883]" pageId="1" pageNumber="2">ZSMG</collectionCode>
, three specimens
<collectionCode id="4FFC34D5BF470300FD95FCBEFD1CFC99" box="[615,675,864,883]" pageId="1" pageNumber="2">PKOC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF470302FF83FC57FBABFE5F" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="2952AC10BF470300FF83FC57FD31FBCB" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
<emphasis id="1B997002BF470300FF83FC57FEE8FC48" bold="true" box="[113,343,905,930]" pageId="1" pageNumber="2">Description of male.</emphasis>
Body dark reddish brown; head entirely black, antennae dark brown; pronotum and elytra reddish brown to brown with paler sides; legs yellowish brown, femora and tibiae with dark brown basal parts; ultimate abdominal tergite dark brown; forceps dark brown to reddish brown at apex. Cuticle punctured, dull, pubescent; tegmina rugose. Length without forceps
<quantity id="EE1501F5BF470300FEEFFBD4FECEFBCB" box="[285,369,1034,1058]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="2" unit="mm" value="7.3">7.3 mm</quantity>
, length of forceps
<quantity id="EE1501F5BF470300FDC7FBD4FD35FBCB" box="[565,650,1034,1058]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="2" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
<paragraph id="2952AC10BF470300FF62FBF7FC02FB0B" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
Head globose, as wide as long, postfrontal and coronal sutures indistinct, posterior margin straight, with short and dense pubescence. Eyes small, less than 0.5 times as long as length of head behind eyes. Antennae of
<typeStatus id="F65612B2BF470300FE4DFBB7FDA3FB6B" box="[447,540,1129,1153]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
male with 17 antennomeres (incomplete); antennomere 1 short, narrowed basally, widened terminally, half as long as the distance between antennal bases; antennomere 2 transverse, only little longer than wide; antennomere 3 longer than antennomere 4. Antennomeres 1-3 cylindrical, antennomeres 4-17 conical.
</paragraph>
<paragraph id="2952AC10BF470300FF62FB34FE3AFA4B" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
Pronotum transverse, widened posteriorly.Anterior margin straight, posterior margin broadly rounded. Pronotal disc rugose, with smoother anterior half, median suture distinct. Tegmina as long as pronotum, rugose, parallel-sided; each with well marked lateral longitudinal ridge, tegminal disc flattened; posterior margin convex, with broadly rounded interior angle. Wings entirely concealed under tegmina, reduced to very small, longitudinally oriented flaps (
<quantity id="EE1501F5BF470300FBE3FAB7FF25FA4B" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="2" unit="mm" value="0.25">0.25 mm</quantity>
long,
<quantity id="EE1501F5BF470300FF2FFA57FE84FA4B" box="[221,315,1417,1441]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="mm" value="0.05">0.05 mm</quantity>
wide).
</paragraph>
<paragraph id="2952AC10BF470300FF62FA77FED6F98B" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
Abdomen convex, somewhat widened posteriorly, densely punctured, shortly pubescent with long setae along hind margin of each tergite except the ultimate one, lateral glandular folds invisible. Penultimate sternite punctulate, sides scarcely concave, hind margin obtusely produced in middle (
<figureCitation id="B1D6B095BF470300FEA4F9D7FE25F9C8" box="[342,410,1545,1570]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
). Ultimate tergite transverse, slightly depressed medially, with invisible median line, little oblique laterally, hind margin straight with protuberance at each base of forceps branch.
</paragraph>
<paragraph id="2952AC10BF440303FF44FF01FBF2FEBD" blockId="2.[151,1127,223,343]" pageId="2" pageNumber="3">
Pygidium invisible in dorsal view. Forceps (
<figureCitation id="B1D6B095BF440303FD66FF01FD44FF1D" box="[660,763,223,247]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
) with branches remote, gradually incurved, each tapering to pointed and gently hooked apex, triangular in basal half and dorsoventrally flattened in distal half. Inner and upper margins of forceps serrated in proximal third, middle of forceps with distinct tapering, distal third widened into flat protuberance.
</paragraph>
<caption id="7D92FC98BF440303FF65F9C5FED6F98E" ID-DOI="http://doi.org/10.5281/zenodo.4503521" ID-Zenodo-Dep="4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,187,1563,1582]" targetBox="[170,1103,390,1539]" targetPageId="2">
<paragraph id="2952AC10BF440303FF65F9C5FED6F98E" blockId="2.[151,1127,1563,1636]" pageId="2" pageNumber="3">
Figs 1-5.
<taxonomicName id="EEEDD793BF440303FF1BF9C5FEEFF9C4" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[233,336,1563,1582]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF440303FF1BF9C5FEEFF9C4" box="[233,336,1563,1582]" italics="true" pageId="2" pageNumber="3">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF440303FEA5F9C5FE25F9C4" box="[343,410,1563,1582]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="2952AC10BF450302FF62FF03FC5EFEDF" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
Genitalia (
<figureCitation id="B1D6B095BF450302FF0DFF03FE80FF1E" box="[255,319,220,245]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
) typical for the genus
<taxonomicName id="EEEDD793BF450302FDDCFF03FDDEFF1F" authorityName="Zacher" authorityYear="1910" box="[558,609,221,245]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FDDCFF03FDDEFF1F" box="[558,609,221,245]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
, with parameres a little over six times longer than broad, narrowed apically, tip acuminate; genital lobes widening to the apex, with straight apical part; virga tubular with undulated central part and sclerotized basal vesicles.
</paragraph>
<paragraph id="2952AC10BF450302FF83FEE2FBABFE5F" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FEE2FED3FEBE" bold="true" box="[113,364,316,340]" pageId="3" pageNumber="4">Description of female.</emphasis>
Body length without forceps
<quantity id="EE1501F5BF450302FD5FFEE3FC93FEBE" box="[685,812,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.9" metricValueMin="6.8" pageId="3" pageNumber="4" unit="mm" value="7.35" valueMax="7.9" valueMin="6.8">6.8-7.9 mm</quantity>
, length of forceps
<quantity id="EE1501F5BF450302FC04FEE3FF21FE9E" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="3" pageNumber="4" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4-1.5 mm</quantity>
. Body colouration similar to
<typeStatus id="F65612B2BF450302FE12FE83FD82FE9F" box="[480,573,349,373]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
male, abdomen wider in middle part. Ultimate tergite with tapering sides, hind margin sinuate in middle. Forceps straight, simple, tapering, inner margin serrated, triangular in basal third, the rest dorsoventrally flattened (
<figureCitation id="B1D6B095BF450302FC34FE43FBB9FE5F" box="[966,1030,413,437]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FE62FC7AFCDF" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="2952AC10BF450302FF83FE62FC7AFCDF" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FE62FED2FE3E" bold="true" box="[113,365,444,468]" pageId="3" pageNumber="4">Differential diagnosis.</emphasis>
<taxonomicName id="EEEDD793BF450302FE84FE63FE45FE3F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[374,506,445,469]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FE84FE63FE45FE3F" box="[374,506,445,469]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FDF6FE63FDE6FE3E" box="[516,601,445,468]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="EEEDD793BF450302FD61FE63FC04FE3F" authority="(Dufour, 1829)" baseAuthorityName="Dufour" baseAuthorityYear="1829" box="[659,955,445,469]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lividipes">
<emphasis id="1B997002BF450302FD61FE63FCB0FE3F" box="[659,783,445,469]" italics="true" pageId="3" pageNumber="4">N. lividipes</emphasis>
(Dufour, 1829)
</taxonomicName>
are the two smallest species of this genus known from South-East Asia, with total body length including forceps ranging between
<quantity id="EE1501F5BF450302FE74FE22FDBBFDFE" box="[390,516,508,532]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="3" pageNumber="4" unit="mm" value="8.0" valueMax="8.5" valueMin="7.5">7.5-8.5 mm</quantity>
.
<taxonomicName id="EEEDD793BF450302FDE7FE23FD26FDFF" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[533,665,509,533]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FDE7FE23FD26FDFF" box="[533,665,509,533]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FD51FE23FD46FDFE" box="[675,761,509,532]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="EEEDD793BF450302FBDAFE23FF75FDDF" baseAuthorityName="Dufour" baseAuthorityYear="1829" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lividipes">
<emphasis id="1B997002BF450302FBDAFE23FF75FDDF" italics="true" pageId="3" pageNumber="4">N. lividipes</emphasis>
</taxonomicName>
by the very short tegmina, which are only as long as the pronotum, and the wings entirely concealed under the tegmina.
<taxonomicName id="EEEDD793BF450302FDE0FDE3FCE4FDBF" authority="Brindle, 1967" authorityName="Brindle" authorityYear="1967" box="[530,859,573,597]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="timorensis">
<emphasis id="1B997002BF450302FDE0FDE3FD04FDBE" box="[530,699,573,597]" italics="true" pageId="3" pageNumber="4">Nala timorensis</emphasis>
Brindle, 1967
</taxonomicName>
, and
<taxonomicName id="EEEDD793BF450302FC65FDE3FBBBFDBF" authorityName="Bey-Bienko" authorityYear="1970" box="[919,1028,573,597]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="1B997002BF450302FC65FDE3FBBBFDBF" box="[919,1028,573,597]" italics="true" pageId="3" pageNumber="4">N. basalis</emphasis>
</taxonomicName>
Bey-Bienko, 1970, also have concealed wings; however, winged specimens were described for the latter species (
<bibRefCitation id="4D7CD1E1BF450302FEE2FDA3FE7BFD7F" author="SRIVASTAVA G. K." box="[272,452,637,661]" pageId="3" pageNumber="4" refId="ref2425" refString="SRIVASTAVA G. K. 2003: Dermaptera. Part II. Superfamily: Anisolaboidea. Fauna of India and adjacent countries. Zoological Survey of India, Kolkata, 235 pp." type="book" year="2003">SRIVASTAVA 2003</bibRefCitation>
). The forceps of
<taxonomicName id="EEEDD793BF450302FD70FDA3FD54FD7F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[642,747,637,661]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FD70FDA3FD54FD7F" box="[642,747,637,661]" italics="true" pageId="3" pageNumber="4">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FD06FDA3FCF5FD7E" box="[756,842,637,660]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
are unusually short in males, being only as long as the width of the ultimate tergite.
<taxonomicName id="EEEDD793BF450302FCF2FD43FC3EFD5F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[768,897,669,693]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FCF2FD43FC3EFD5F" box="[768,897,669,693]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FC7AFD43FC65FD5E" box="[904,986,669,692]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is further characterized by the entirely black head. Although the females of
<taxonomicName id="EEEDD793BF450302FCDDFD63FCDDFD3F" authorityName="Zacher" authorityYear="1910" box="[815,866,701,725]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FCDDFD63FCDDFD3F" box="[815,866,701,725]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
are morphologically very similar and cannot be reliably identified, the females of
<taxonomicName id="EEEDD793BF450302FCE2FD03FCC8FD1F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[784,887,733,757]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FCE2FD03FCC8FD1F" box="[784,887,733,757]" italics="true" pageId="3" pageNumber="4">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FC73FD03FC69FD1E" box="[897,982,733,756]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
are fairly easily recognized by their exceptionally short tegmina and the absence of wings (see above). The males of South-East Asia
<taxonomicName id="EEEDD793BF450302FE41FCC3FE59FCDF" authorityName="Zacher" authorityYear="1910" box="[435,486,797,821]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FE41FCC3FE59FCDF" box="[435,486,797,821]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
can be separated by the key provided below.
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FCE3FDA0FC3F" pageId="3" pageNumber="4" type="etymology">
<paragraph id="2952AC10BF450302FF83FCE3FF58FC9F" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FCE3FF4EFCBF" bold="true" box="[113,241,829,853]" pageId="3" pageNumber="4">Etymology.</emphasis>
The new species is dedicated to my wife Alena, who helped me collect the
<typeStatus id="F65612B2BF450302FBE1FCE3FBFEFCBF" box="[1043,1089,829,853]" pageId="3" pageNumber="4">type</typeStatus>
specimens.
</paragraph>
<paragraph id="2952AC10BF450302FF83FCA2FDA0FC3F" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FCA2FF51FC7E" bold="true" box="[113,238,892,916]" pageId="3" pageNumber="4">Bionomics.</emphasis>
Unknown.
<typeStatus id="F65612B2BF450302FE9FFCA3FE1DFC7F" box="[365,418,893,917]" pageId="3" pageNumber="4">Type</typeStatus>
specimens have been collected in the surroundings of Angkor Wat temples in a stony habitat surrounded by lowland rain forest. The earwigs were observed to run on stony ground in late afternoon.
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FC02FEC1FC1F" box="[113,382,988,1013]" pageId="3" pageNumber="4" type="distribution">
<paragraph id="2952AC10BF450302FF83FC02FEC1FC1F" blockId="3.[113,1090,220,1301]" box="[113,382,988,1013]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FC02FEBDFC1E" bold="true" box="[113,258,988,1012]" pageId="3" pageNumber="4">Distribution.</emphasis>
<collectingCountry id="51FAEC80BF450302FEF8FC03FEC5FC1F" box="[266,378,989,1013]" name="Cambodia" pageId="3" pageNumber="4">Cambodia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FC22FD3CFAFF" pageId="3" pageNumber="4" type="discussion">
<paragraph id="2952AC10BF450302FF83FC22FD3CFAFF" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FC22FF4DFBFE" bold="true" box="[113,242,1020,1044]" pageId="3" pageNumber="4">Comments.</emphasis>
<taxonomicName id="EEEDD793BF450302FF09FC23FE3FFBFF" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[251,384,1021,1045]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alanae" status="sp. nov.">
<emphasis id="1B997002BF450302FF09FC23FE3FFBFF" box="[251,384,1021,1045]" italics="true" pageId="3" pageNumber="4">Nala alanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FE78FC23FE60FBFE" box="[394,479,1021,1044]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is a very characteristic species, differing from all other Asian species in the shape of the reduced tegmina with well marked longitudinal ridges (see
<figureCitation id="B1D6B095BF450302FF83FBE3FF0FFBBF" box="[113,176,1085,1109]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
). Other
<taxonomicName id="EEEDD793BF450302FEF9FBE3FE81FBBF" authorityName="Zacher" authorityYear="1910" box="[267,318,1085,1109]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FEF9FBE3FE81FBBF" box="[267,318,1085,1109]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
species with similar tegmina, reduced wings and short stout forceps are known only from Central and
<collectingCountry id="51FAEC80BF450302FE41FB83FD84FB9F" box="[435,571,1117,1141]" name="South Africa" pageId="3" pageNumber="4">South Africa</collectingCountry>
(
<taxonomicName id="EEEDD793BF450302FDB8FB83FCE8FB9F" authority="Menozzi, 1929" authorityName="Menozzi" authorityYear="1929" box="[586,855,1117,1141]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caprai">
<emphasis id="1B997002BF450302FDB8FB83FD11FB9E" box="[586,686,1117,1140]" italics="true" pageId="3" pageNumber="4">N. caprai</emphasis>
Menozzi, 1929
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FC90FB83FB83FB9F" authority="Burr, 1908" authorityName="Burr" authorityYear="1908" box="[866,1084,1117,1141]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="figinii">
<emphasis id="1B997002BF450302FC90FB83FC7EFB9F" box="[866,961,1117,1141]" italics="true" pageId="3" pageNumber="4">N. figinii</emphasis>
Burr, 1908
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FF83FBA3FE14FB7F" authority="Menozzi, 1937" authorityName="Menozzi" authorityYear="1937" box="[113,427,1149,1173]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="1B997002BF450302FF83FBA3FEBCFB7F" box="[113,259,1149,1173]" italics="true" pageId="3" pageNumber="4">N. intermedia</emphasis>
Menozzi, 1937
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FE44FBA3FD02FB7F" authority="Brindle, 1968" authorityName="Brindle" authorityYear="1968" box="[438,701,1149,1173]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="1B997002BF450302FE44FBA3FD9CFB7E" box="[438,547,1149,1172]" italics="true" pageId="3" pageNumber="4">N. saegeri</emphasis>
Brindle, 1968
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FD3AFBA3FBB0FB7F" authority="Vigna Taglianti, 1994" authorityName="Vigna Taglianti" authorityYear="1994" box="[712,1039,1149,1173]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rossii">
<emphasis id="1B997002BF450302FD3AFBA3FC9EFB7E" box="[712,801,1149,1172]" italics="true" pageId="3" pageNumber="4">N. rossii</emphasis>
<bibRefCitation id="4D7CD1E1BF450302FCDAFBA3FBB0FB7F" author="VIGNA TAGLIANTI A." box="[808,1039,1149,1173]" pageId="3" pageNumber="4" pagination="199 - 212" refId="ref2505" refString="VIGNA TAGLIANTI A. 1994: Further notes on Dermaptera from Sierra Leone. Quaderni dell'Accademia Nazionale dei Lincei 267: 199 - 212." type="journal article" year="1994">Vigna Taglianti, 1994</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="EEEDD793BF450302FF83FB43FE6BFB5F" authority="Bivar de Sousa, 1999" authorityName="Bivar de Sousa" authorityYear="1999" box="[113,468,1181,1205]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mendesi">
<emphasis id="1B997002BF450302FF83FB43FF58FB5F" box="[113,231,1181,1205]" italics="true" pageId="3" pageNumber="4">N. mendesi</emphasis>
<bibRefCitation id="4D7CD1E1BF450302FF1FFB43FE6BFB5F" author="BIVAR DE SOUSA A." box="[237,468,1181,1205]" pageId="3" pageNumber="4" pagination="141 - 146" refId="ref2285" refString="BIVAR DE SOUSA A. 1999: Descricao de uma nova especie de dermaptero de Guine Bissau. Nala mendesi sp. nov. (Insecta, Dermaptera). (Description of a new species of Dermaptera from Guinea-Bissau. Nala mendesi n. sp. (Insecta, Dermaptera )). Boletim da Sociedade Portuguesa de Entomologia 194: 141 - 146 (in Portugese, English summary)." type="journal article" year="1999">Bivar de Sousa, 1999</bibRefCitation>
</taxonomicName>
).
<taxonomicName id="EEEDD793BF450302FE14FB43FDD9FB5F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[486,614,1181,1205]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FE14FB43FDD9FB5F" box="[486,614,1181,1205]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FD9FFB43FD00FB5E" box="[621,703,1181,1204]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is the only known Asian representative with a noticeable morphological similarity to this African group of species, which may indicate some affinity to this group. From the known African species,
<taxonomicName id="EEEDD793BF450302FC88FB03FC5CFB1F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[890,995,1245,1269]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FC88FB03FC5CFB1F" box="[890,995,1245,1269]" italics="true" pageId="3" pageNumber="4">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FC1EFB03FBFEFB1E" box="[1004,1089,1245,1268]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
differs in the characteristic shape of male forceps.
</paragraph>
</subSubSection>
</treatment>
</document>
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