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<mods:title id="A24C154E20CBD500994707BBA4B83634">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="F46C6D8FE2FE4430E6CD5EE7CCD558D0">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="0412FC70FC5D85F54D71B3D8B8E57517">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:namePart id="5D2B1C1341FD1D29C770EEC9CFCE00F1">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="F12D4C17AC4FE9FFE362BFEFF664E8DB">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<heading id="862F81A8FFF589059D6CFA62FC3EFA67" box="[621,967,1491,1517]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
Genus 
<taxonomicName id="1AD84D47FFF589059DBEFA62FC3EFA67" ID-CoL="68C6" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[703,967,1491,1517]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EFACEAD6FFF589059DBEFA62FCF1FA67" bold="true" box="[703,776,1491,1517]" italics="true" pageId="3" pageNumber="4">Onitis</emphasis>
<bibRefCitation id="B9494B35FFF589059C11FA62FC3EFA67" author="Fabricius J. C." box="[784,967,1491,1517]" pageId="3" pageNumber="4" refId="ref29024" refString="Fabricius J. C. 1798. Supplementum Entomologiae Systematicae. Proft et Storch, Copenhagen [Hafniae]. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1798">Fabricius,1798</bibRefCitation>
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<paragraph id="DD6736C4FFF589059FBCF9A9FB93F9BE" blockId="3.[189,1130,1559,1588]" box="[189,1130,1559,1588]" pageId="3" pageNumber="4">
<emphasis id="EFACEAD6FFF589059FBCF9A9FB93F9BE" bold="true" box="[189,1130,1559,1588]" pageId="3" pageNumber="4">
Key to species of 
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<emphasis id="EFACEAD6FFF589059EA9F9A6FE01F9B9" bold="true" box="[424,504,1559,1587]" italics="true" pageId="3" pageNumber="4">Onitis</emphasis>
<bibRefCitation id="B9494B35FFF589059D01F9A6FD28F9BE" author="Fabricius J. C." box="[512,721,1559,1588]" pageId="3" pageNumber="4" refId="ref29024" refString="Fabricius J. C. 1798. Supplementum Entomologiae Systematicae. Proft et Storch, Copenhagen [Hafniae]. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1798">Fabricius, 1798</bibRefCitation>
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from the Indian subcontinent
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The following key is based on external morphological characters and can be used to identify both male and female adults of 
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<emphasis id="EFACEAD6FFF589059EACF9C0FE0CF901" box="[429,501,1649,1675]" italics="true" pageId="3" pageNumber="4">Onitis</emphasis>
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. For the convenience of identification, we have retained the groups suggested by 
<bibRefCitation id="B9494B35FFF589059FE0F925FE67F924" author="Janssens A." box="[225,414,1684,1710]" pageId="3" pageNumber="4" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens (1937)</bibRefCitation>
. The keys have been modified from the identification keys to the species provided by 
<bibRefCitation id="B9494B35FFF589059FBCF906FEA6F95B" author="Arrow G. J." box="[189,351,1719,1745]" pageId="3" pageNumber="4" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow (1931)</bibRefCitation>
and 
<bibRefCitation id="B9494B35FFF589059E99F906FD99F95B" author="Balthasar V." box="[408,608,1719,1745]" pageId="3" pageNumber="4" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar (1963)</bibRefCitation>
.
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<keyLead id="662995F1FFF589059FBCF94BFA8EF8B2" pageId="3" pageNumber="4">1. Head with frontal carina interrupted medially, with tubercle placed adjoining the carina; clypeus always with a short or long transverse carina.................................................................................... 2</keyLead>
</paragraph>
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<keyLead id="662995F1FFF589059FBCF8F2FA8FF80A" pageId="3" pageNumber="4">– Head with frontal carina uninterrupted or entire, with tubercle behind or absent; clypeal carina feebly expressed.......................................................................................................................................... 13</keyLead>
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2. Body completely black or dark brown (
<emphasis id="EFACEAD6FFF589059DB8F818FCCDF849" bold="true" box="[697,820,1961,1987]" pageId="3" pageNumber="4">GROUP I</emphasis>
) .......................................................................... 3
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– Body metallic, with coppery or green tinge....................................................................................... 6 
<keyStep id="662C2E61FFF289029FBCFEB8FA8EFEA9" box="[189,1399,265,292]" pageId="4" pageNumber="5">
3. Pronotum minutely punctured (
<figureCitation id="45E32A41FFF289029D5BFEB8FD2BFEAE" box="[602,722,265,292]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="4" pageNumber="5">Fig. 5E–F</figureCitation>
)........................................................................................ 4
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</paragraph>
<paragraph id="DD6736C4FFF289029FBCFE9FFA8EFEC2" blockId="4.[189,1399,265,328]" box="[189,1399,301,328]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFE9FFA8EFEC2" box="[189,1399,301,328]" pageId="4" pageNumber="5">
– Pronotum strongly punctured (
<figureCitation id="45E32A41FFF289029D53FE9CFD26FECD" box="[594,735,301,327]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">Fig. 14A–B</figureCitation>
)...................................................................................... 5
</keyLead>
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<paragraph id="DD6736C4FFF289029FBCFEC1FA8FFE27" blockId="4.[189,1399,368,501]" pageId="4" pageNumber="5">
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4. Metasternal shield in male with deep transverse excavation in the middle; without an anterior longitudinal groove ................................................... 
<taxonomicName id="1AD84D47FFF289029C4BFE22FB86FE27" ID-CoL="49KG2" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[842,1151,403,429]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFF289029C4BFE22FC1CFE27" bold="true" box="[842,997,403,429]" italics="true" pageId="4" pageNumber="5">O. excavatus</emphasis>
<bibRefCitation id="B9494B35FFF289029CEAFE22FB86FE27" author="Arrow G. J." box="[1003,1151,403,429]" pageId="4" pageNumber="5" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B8FFE22FB14FE24" box="[1166,1261,403,430]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="4" pageNumber="5">Figs 5E</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BF8FE22FABDFE27" box="[1273,1348,403,429]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="4" pageNumber="5">6A–D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFE22FA94FE27" box="[1359,1389,403,429]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="4" pageNumber="5">18</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFE09FA8FFE7F" blockId="4.[189,1399,368,501]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFE09FA8FFE7F" pageId="4" pageNumber="5">
– Metasternal shield without transverse excavation in the middle, with an anterior longitudinal groove ...................................................................... 
<taxonomicName id="1AD84D47FFF289029C46FE6BFB7FFE7E" ID-CoL="49KG5" authority="(Wulfen, 1786)" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[839,1158,474,501]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFF289029C46FE6BFC32FE7F" bold="true" box="[839,971,474,501]" italics="true" pageId="4" pageNumber="5">O. falcatus</emphasis>
(
<bibRefCitation id="B9494B35FFF289029CDBFE6BFB87FE7E" author="Wulfen X." box="[986,1150,474,500]" pageId="4" pageNumber="5" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B94FE6BFB09FE7F" box="[1173,1264,474,501]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="4" pageNumber="5">Figs 5F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BFDFE6BFABAFE7E" box="[1276,1347,474,500]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="4" pageNumber="5">6E–H</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFE6AFA94FE7F" box="[1359,1389,475,501]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="4" pageNumber="5">18</figureCitation>
)
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<paragraph id="DD6736C4FFF289029FBCFDACFA8FFDD0" blockId="4.[189,1398,541,674]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFDACFA8FFDD0" pageId="4" pageNumber="5">
5. Pygidium with a median longitudinal groove; protibia with a rudimentary spur before the extremity (
<figureCitation id="45E32A41FFF289029FF2FDF1FEA7FDD0" box="[243,350,576,602]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">Fig. 16A</figureCitation>
) ..................................... 
<taxonomicName id="1AD84D47FFF289029D88FDF1FCB1FDD1" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[649,840,576,603]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFF289029D88FDF1FCB1FDD1" bold="true" box="[649,840,576,603]" italics="true" pageId="4" pageNumber="5">O. bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFF289029C4EFDF0FC53FDD1" box="[847,938,577,603]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="45E32A41FFF289029CB8FDF1FBFCFDD0" box="[953,1029,576,602]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742855" httpUri="https://zenodo.org/record/13742855/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B13FDF1FBB4FDD0" box="[1042,1101,576,602]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="4" pageNumber="5">6I–L</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B58FDF1FB79FDD0" box="[1113,1152,576,602]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">9A</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B8DFDF1FB38FDD0" box="[1164,1217,576,602]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14A</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BCCFDF1FAFBFDD1" box="[1229,1282,576,603]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="4" pageNumber="5">15A</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A0FFDF1FABDFDD0" box="[1294,1348,576,602]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">16A</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFDF1FA94FDD0" box="[1359,1389,576,602]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="4" pageNumber="5">17</figureCitation>
)
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</paragraph>
<paragraph id="DD6736C4FFF289029FBCFDD4FA8FFD28" blockId="4.[189,1398,541,674]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFDD4FA8FFD28" pageId="4" pageNumber="5">
– Pygidium without a median longitudinal groove; protibia without a rudimentary spur (
<figureCitation id="45E32A41FFF289029BDFFDD5FABEFDF4" box="[1246,1351,612,638]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">Fig. 16B</figureCitation>
).... ................................. 
<taxonomicName id="1AD84D47FFF289029EEBFD39FC71FD28" ID-CoL="49KKQ" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[490,904,648,674]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFF289029EEBFD39FD2BFD28" bold="true" box="[490,722,648,674]" italics="true" pageId="4" pageNumber="5">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFF289029DD8FD39FC71FD28" author="Janssens A." box="[729,904,648,674]" pageId="4" pageNumber="5" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029C96FD39FBE3FD28" box="[919,1050,648,674]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="4" pageNumber="5">Figs 6M–P</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B27FD39FBB3FD28" box="[1062,1098,648,674]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">9B</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B56FD39FB7CFD28" box="[1111,1157,648,674]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="4" pageNumber="5">10F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B90FD39FB3DFD28" box="[1169,1220,648,674]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14B</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BD1FD36FAFAFD28" box="[1232,1283,647,674]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="4" pageNumber="5">15B</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A11FD39FABAFD28" box="[1296,1347,648,674]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">16B</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFD39FA94FD28" box="[1359,1389,648,674]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="4" pageNumber="5">19</figureCitation>
)
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<keyStep id="662C2E61FFF289029FBCFD7BFA8FFCC5" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFD7BFA8EFC82" blockId="4.[189,1399,714,847]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFD7BFA8EFC82" pageId="4" pageNumber="5">
6. Vertex covered with granules or highly raspy punctuation; clypeal carina closer to the frontal carina than to the clypeal margin (
<figureCitation id="45E32A41FFF289029D2DFD5CFD51FC82" box="[556,680,749,776]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">Fig. 9A–F</figureCitation>
) (
<emphasis id="EFACEAD6FFF289029DC0FD5CFCBEFC82" bold="true" box="[705,839,749,776]" pageId="4" pageNumber="5">GROUP II</emphasis>
)........................................................................ 7
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</paragraph>
<paragraph id="DD6736C4FFF289029FBCFCA3FA8FFCC5" blockId="4.[189,1399,714,847]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFCA3FA8FFCC5" pageId="4" pageNumber="5">
– Vertex covered with punctures and never rough or granular; clypeal carina midway between clypeal margin and frontal carina (
<figureCitation id="45E32A41FFF289029D22FC84FD7DFCC5" box="[547,644,821,847]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" figureDoi="http://doi.org/10.5281/zenodo.13742852" httpUri="https://zenodo.org/record/13742852/files/figure.png" pageId="4" pageNumber="5">Figs 1A</figureCitation>
, 
<figureCitation id="45E32A41FFF289029D90FC84FD3CFCC5" box="[657,709,821,847]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="4" pageNumber="5">10D</figureCitation>
) (
<emphasis id="EFACEAD6FFF289029DDFFC84FC89FCC5" bold="true" box="[734,880,821,847]" pageId="4" pageNumber="5">GROUP III</emphasis>
)................................................................ 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFCC9FA8FFC3C" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFCC9FA8EFC18" blockId="4.[189,1399,887,950]" box="[189,1399,887,914]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFCC9FA8EFC18" box="[189,1399,887,914]" pageId="4" pageNumber="5">
7. Metasternum longitudinally grooved anteriorly (
<figureCitation id="45E32A41FFF289029C23FCC6FC71FC18" box="[802,904,887,914]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="4" pageNumber="5">Fig. 12F</figureCitation>
)............................................................... 8
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFC2DFA8FFC3C" blockId="4.[189,1399,887,950]" box="[189,1398,924,950]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFC2DFA8FFC3C" box="[189,1398,924,950]" pageId="4" pageNumber="5">
– Metasternum not grooved anteriorly (
<figureCitation id="45E32A41FFF289029DA2FC2DFD05FC3C" box="[675,764,924,950]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" figureDoi="http://doi.org/10.5281/zenodo.13742852" httpUri="https://zenodo.org/record/13742852/files/figure.png" pageId="4" pageNumber="5">Fig. 1B</figureCitation>
).................................................................................11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFC6FFA8FFBE9" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFC6FFA8EFB96" blockId="4.[189,1399,990,1123]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFC6FFA8EFB96" pageId="4" pageNumber="5">
8. Pronotum without a smooth median longitudinal line; genae smooth, inconspicuously punctured (
<figureCitation id="45E32A41FFF289029FF2FBB0FE7DFB96" box="[243,388,1025,1052]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">Fig. 14C–D</figureCitation>
); male profemur not toothed......................................................................................... 9
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFB97FA8FFBE9" blockId="4.[189,1399,990,1123]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFB97FA8FFBE9" pageId="4" pageNumber="5">
– Pronotum with a smooth median longitudinal line; genae sparingly granular (
<figureCitation id="45E32A41FFF289029B99FB97FADCFBCA" box="[1176,1317,1062,1088]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">Fig. 14E–F</figureCitation>
); male profemur toothed ............................................................................................................................. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFB3DFA8FFA9A" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFB3DFA8FFB43" blockId="4.[189,1399,1163,1297]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFB3DFA8FFB43" pageId="4" pageNumber="5">
9. Clypeus granular in male and rugose in female (
<figureCitation id="45E32A41FFF289029C30FB3AFC6CFB2C" box="[817,917,1163,1190]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">Figs 9D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029CA5FB3DFC23FB2C" box="[932,986,1164,1190]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14D</figureCitation>
), pronotum strongly and unevenly punctured .......................... 
<taxonomicName id="1AD84D47FFF289029D2BFB1EFC87FB43" ID-CoL="49KK8" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[554,894,1198,1225]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFF289029D2BFB1EFD46FB43" bold="true" box="[554,703,1198,1225]" italics="true" pageId="4" pageNumber="5">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFF289029DC6FB1EFC87FB43" author="Fabricius J. C." box="[711,894,1199,1225]" pageId="4" pageNumber="5" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029C8CFB1EFBE9FB43" box="[909,1040,1199,1225]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="4" pageNumber="5">Figs 8E–H</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B1AFB1EFBB8FB43" box="[1051,1089,1199,1225]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">9D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B4FFB1EFB79FB43" box="[1102,1152,1199,1225]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="4" pageNumber="5">10E</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B8DFB1EFB38FB43" box="[1164,1217,1199,1225]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BCCFB1FFAFBFB43" box="[1229,1282,1198,1225]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="4" pageNumber="5">15D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A0FFB1EFABAFB43" box="[1294,1347,1199,1225]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">16D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFB1EFA94FB43" box="[1359,1389,1199,1225]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="4" pageNumber="5">19</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFB62FA8FFA9A" blockId="4.[189,1399,1163,1297]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFB62FA8FFA9A" pageId="4" pageNumber="5">
– Clypeus rugulose in both sexes; pronotum strongly and closely punctured (
<figureCitation id="45E32A41FFF289029B58FB62FB41FB67" box="[1113,1208,1235,1261]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">Figs 9C</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BC4FB62FB0EFB67" box="[1221,1271,1235,1261]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14C</figureCitation>
)............... ................................................................. 
<taxonomicName id="1AD84D47FFF289029DD0FB47FCB9FA9B" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[721,832,1270,1297]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFF289029DD0FB47FCB9FA9B" bold="true" box="[721,832,1270,1297]" italics="true" pageId="4" pageNumber="5">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFF289029C46FB46FC5BFA9B" box="[839,930,1271,1297]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="45E32A41FFF289029CB0FB47FC04FA9B" box="[945,1021,1270,1297]" captionStart="Fig" captionStartId="26.[189,232,1923,1949]" captionTargetBox="[299,1295,368,1851]" captionTargetId="figure-42@26.[191,1400,269,1864]" captionTargetPageId="26" captionText="Fig. 7. Onitis kethai sp. nov. A–E. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. E. Elytra. F. Paratype, ♀, habitus, dorsal view. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742869" httpUri="https://zenodo.org/record/13742869/files/figure.png" pageId="4" pageNumber="5">Figs 7</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B08FB47FBADFA9A" box="[1033,1108,1270,1296]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="4" pageNumber="5">8A–D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B61FB47FB7DFA9A" box="[1120,1156,1270,1296]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">9C</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B90FB47FB3DFA9A" box="[1169,1220,1270,1296]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14C</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BD1FB47FAFDFA9B" box="[1232,1284,1270,1297]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="4" pageNumber="5">15C</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A11FB47FABAFA9A" box="[1296,1347,1270,1296]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">16C</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFB47FA94FA9A" box="[1359,1389,1270,1296]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="4" pageNumber="5">18</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFA88FA8FF9AD" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFA88FA8FFA13" blockId="4.[189,1399,1337,1576]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFA88FA8FFA13" pageId="4" pageNumber="5">
10. Lateral margin of pronotum feebly curved; front angles of pronotum blunt (
<figureCitation id="45E32A41FFF289029B51FA88FB50FAD9" box="[1104,1193,1337,1363]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">Figs 9F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BB2FA88FB1CFAD9" box="[1203,1253,1337,1363]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14F</figureCitation>
); mid-femur in male with a tooth near the middle of its posterior edge, another at the extremity (
<figureCitation id="45E32A41FFF289029BBBFAEDFAD9FAFC" box="[1210,1312,1372,1398]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">Fig. 16F</figureCitation>
) ......... ......................................... 
<taxonomicName id="1AD84D47FFF289029D1FFACEFC5AFA10" ID-CoL="49KL7" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[542,931,1406,1434]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFF289029D1FFACEFD21FA13" bold="true" box="[542,728,1406,1433]" italics="true" pageId="4" pageNumber="5">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFF289029DDEFACEFC5AFA10" author="Lansberge G." box="[735,931,1407,1434]" pageId="4" pageNumber="5" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029CB3FACEFBD0FA13" box="[946,1065,1407,1433]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="4" pageNumber="5">Figs 8I–L</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B34FACEFBADFA13" box="[1077,1108,1407,1433]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">9F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B61FACEFB6FFA13" box="[1120,1174,1407,1433]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="4" pageNumber="5">12A</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BA0FACEFB29FA13" box="[1185,1232,1407,1433]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BDAFACEFAF0FA10" box="[1243,1289,1407,1434]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="4" pageNumber="5">15F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A14FACEFABDFA13" box="[1301,1348,1407,1433]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">16F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EFACEFA94FA13" box="[1359,1389,1407,1433]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="4" pageNumber="5">19</figureCitation>
)
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<paragraph id="DD6736C4FFF289029FBCFA15FA8FF9AD" blockId="4.[189,1399,1337,1576]" pageId="4" pageNumber="5">
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– Lateral margin of pronotum strongly curved; front angles of pronotum sharp (
<figureCitation id="45E32A41FFF289029B91FA12FB09FA37" box="[1168,1264,1443,1469]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">Figs 9E</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BFFFA12FACBFA37" box="[1278,1330,1443,1469]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14E</figureCitation>
); mid femur in male without a tooth near the middle of the posterior edge, the extremity with a sharp tooth on the lower edge and a blunt one above (
<figureCitation id="45E32A41FFF289029DD4FA5BFCC5F98E" box="[725,828,1514,1540]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">Fig. 16E</figureCitation>
) ........................................................................... ....................................................... 
<taxonomicName id="1AD84D47FFF289029D86F9BCFCF5F9AD" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[647,780,1548,1575]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFF289029D86F9BCFCF5F9AD" bold="true" box="[647,780,1548,1575]" italics="true" pageId="4" pageNumber="5">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFF289029C12F9BCFC97F9AD" box="[787,878,1549,1575]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="45E32A41FFF289029C7CF9BCFBF8F9AD" box="[893,1025,1549,1575]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="4" pageNumber="5">Figs 8M–P</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B0DF9BCFBD6F9AD" box="[1036,1071,1549,1575]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="4" pageNumber="5">9E</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B3AF9BCFB73F9AD" box="[1083,1162,1549,1575]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="4" pageNumber="5">12E–F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029B97F9BCFB3EF9AD" box="[1174,1223,1549,1575]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="4" pageNumber="5">14E</figureCitation>
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<figureCitation id="45E32A41FFF289029BD5F9BCFAFFF9A2" box="[1236,1286,1549,1576]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="4" pageNumber="5">15E</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A10F9BCFABAF9AD" box="[1297,1347,1549,1575]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="4" pageNumber="5">16E</figureCitation>
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<figureCitation id="45E32A41FFF289029A4EF9BCFA97F9AD" box="[1359,1390,1549,1575]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="4" pageNumber="5">20</figureCitation>
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11. Frontal carina narrowly interrupted; pygidium smooth with minute punctures.................................. ............................................................................. 
<taxonomicName id="1AD84D47FFF289029C2BF9C2FB9BF907" ID-CoL="49KMB" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[810,1122,1650,1677]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFF289029C2BF9C2FC66F907" bold="true" box="[810,927,1651,1677]" italics="true" pageId="4" pageNumber="5">O. virens</emphasis>
<bibRefCitation id="B9494B35FFF289029C9FF9C2FB9BF907" author="Lansberge G." box="[926,1122,1650,1677]" pageId="4" pageNumber="5" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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<figureCitation id="45E32A41FFF289029B70F9C2FB18F907" box="[1137,1249,1651,1677]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="4" pageNumber="5">Figs 12D</figureCitation>
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<figureCitation id="45E32A41FFF289029BEFF9C2FABDF907" box="[1262,1348,1651,1677]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. E–H. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742887" httpUri="https://zenodo.org/record/13742887/files/figure.png" pageId="4" pageNumber="5">13E–H</figureCitation>
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<figureCitation id="45E32A41FFF289029A4EF9C2FA97F907" box="[1359,1390,1651,1677]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="4" pageNumber="5">20</figureCitation>
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– Frontal carina widely interrupted; pygidium opaque with imperceptible punctures........................... ............................................................................ 
<taxonomicName id="1AD84D47FFF289029C22F90BFB99F95E" ID-CoL="49KLJ" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[803,1120,1722,1749]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFF289029C22F90BFC3FF95F" bold="true" box="[803,966,1722,1749]" italics="true" pageId="4" pageNumber="5">O. subopacus</emphasis>
<bibRefCitation id="B9494B35FFF289029CCDF90BFB99F95E" author="Arrow G. J." box="[972,1120,1722,1748]" pageId="4" pageNumber="5" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
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<figureCitation id="45E32A41FFF289029B6EF90BFB27F95E" box="[1135,1246,1722,1748]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="4" pageNumber="5">Figs 12C</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BEBF90BFABAF95E" box="[1258,1347,1722,1748]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. E–H. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742887" httpUri="https://zenodo.org/record/13742887/files/figure.png" pageId="4" pageNumber="5">13A–D</figureCitation>
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<figureCitation id="45E32A41FFF289029A4EF90BFA97F95E" box="[1359,1390,1722,1748]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="4" pageNumber="5">20</figureCitation>
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<paragraph id="DD6736C4FFF289029FBCF94CFA8FF8B0" blockId="4.[189,1399,1789,1922]" pageId="4" pageNumber="5">
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12. Pronotal anterior angles not extended, blunt; male protibia with 3–4 teeth on the inferomedial ridge ........................................................................... 
<taxonomicName id="1AD84D47FFF289029C1AF891FB81F8B0" ID-CoL="49KJD" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[795,1144,1824,1850]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFF289029C1AF891FC50F8B0" bold="true" box="[795,937,1824,1850]" italics="true" pageId="4" pageNumber="5">O. naviauxi</emphasis>
<bibRefCitation id="B9494B35FFF289029CB0F891FB81F8B0" author="Cambefort Y." box="[945,1144,1824,1850]" pageId="4" pageNumber="5" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
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<figureCitation id="45E32A41FFF289029B86F891FAFBF8B0" box="[1159,1282,1824,1850]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="4" pageNumber="5">Figs 4E–F</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A0FF891FABAF8B0" box="[1294,1347,1824,1850]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="4" pageNumber="5">10D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EF891FA94F8B0" box="[1359,1389,1824,1850]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="4" pageNumber="5">19</figureCitation>
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– Pronotal anterior angles slightly extended, sharp; male protibia with 6–7 teeth on the inferomedial ridge (
<figureCitation id="45E32A41FFF289029E36F8D6FE76F80B" box="[311,399,1895,1921]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" figureDoi="http://doi.org/10.5281/zenodo.13742852" httpUri="https://zenodo.org/record/13742852/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
) ........................................................ 
<taxonomicName id="1AD84D47FFF289029C41F8D6FB68F80B" ID-CoL="6SMW9" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[832,1169,1895,1921]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF289029C41F8D6FC12F80B" bold="true" box="[832,1003,1895,1921]" italics="true" pageId="4" pageNumber="5">O. assamensis</emphasis>
<bibRefCitation id="B9494B35FFF289029CF2F8D6FB68F80B" author="Biswas S." box="[1011,1169,1895,1921]" pageId="4" pageNumber="5" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
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(
<figureCitation id="45E32A41FFF289029BA1F8D6FB12F80B" box="[1184,1259,1895,1921]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" figureDoi="http://doi.org/10.5281/zenodo.13742852" httpUri="https://zenodo.org/record/13742852/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
, 
<figureCitation id="45E32A41FFF289029BF8F8D6FABDF80B" box="[1273,1348,1895,1921]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="4" pageNumber="5">4A–D</figureCitation>
, 
<figureCitation id="45E32A41FFF289029A4EF8D6FA94F80B" box="[1359,1389,1895,1921]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="4" pageNumber="5">17</figureCitation>
)
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</paragraph>
</keyStep>
<paragraph id="DD6736C4FFF289029FBCF81BFA8FF84E" blockId="4.[189,1398,1962,2025]" box="[189,1398,1962,1988]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCF81BFA8FF84E" box="[189,1398,1962,1988]" pageId="4" pageNumber="5">
<keyStep id="662C2E61FFF289029FBCF81BFA8FF84E" box="[189,1398,1962,1988]" pageId="4" pageNumber="5">
13. Pygidium covered with short or long hair (
<emphasis id="EFACEAD6FFF289029DDEF81BFC83F84E" bold="true" box="[735,890,1962,1988]" pageId="4" pageNumber="5">GROUP VII</emphasis>
)............................................................... 14
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</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289039FBCF87EFA8FFECC" blockId="4.[189,1398,1962,2025]" lastBlockId="5.[189,1398,265,362]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289039FBCF87EFA8FFECC" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
– Pygidium not covered with hair or setae ......................................................................................... 17 
<keyStep id="662C2E61FFF389039FBCFEB8FA8FFECC" pageId="5" pageNumber="6">
14. Clypeus elliptical; head not tuberculate; elytral intervals slightly carinate in the middle................... ..................................................................................... 
<taxonomicName id="1AD84D47FFF389039C63FE9DFB9AFECC" ID-CoL="49KG7" authority="Felsche, 1907" authorityName="Felsche" authorityYear="1907" box="[866,1123,300,326]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFF389039C63FE9DFC41FECC" bold="true" box="[866,952,300,326]" italics="true" pageId="5" pageNumber="6">O. feae</emphasis>
<bibRefCitation id="B9494B35FFF389039CC1FE9DFB9AFECC" author="Felsche C." box="[960,1123,300,326]" pageId="5" pageNumber="6" pagination="273 - 296" refId="ref29092" refString="Felsche C. 1907. Coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift: 273 - 296." type="book chapter" year="1907">Felsche, 1907</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B73FE9DFB1AFECC" box="[1138,1251,300,326]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="5" pageNumber="6">Figs 10A</figureCitation>
, 
<figureCitation id="45E32A41FFF389039BEEFE9DFABDFECC" box="[1263,1348,300,326]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. E–H. O. feae Felsche, 1907, AIMB/Co/Sc1000188. I–L. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742881" httpUri="https://zenodo.org/record/13742881/files/figure.png" pageId="5" pageNumber="6">11E–H</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFE9DFA94FECC" box="[1359,1389,300,326]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="5" pageNumber="6">18</figureCitation>
)
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</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFEE1FA8FFEE0" blockId="5.[189,1398,265,362]" box="[189,1398,335,362]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFEE1FA8FFEE0" box="[189,1398,335,362]" pageId="5" pageNumber="6">– Clypeus truncate or excised; head tuberculate; elytral intervals convex......................................... 15</keyLead>
</paragraph>
<keyStep id="662C2E61FFF389039FBCFE20FA8FFE7B" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFE20FA8FFE44" blockId="5.[189,1398,401,498]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFE20FA8FFE44" pageId="5" pageNumber="6">
15. Clypeal margin deeply excised with an obtuse angle; frontal carina with a small gap ....................... .......................................................................................... 
<taxonomicName id="1AD84D47FFF389039C86FE05FB2DFE45" ID-CoL="74MW8" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[903,1236,435,463]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFF389039C86FE05FBF3FE44" bold="true" box="[903,1034,435,462]" italics="true" pageId="5" pageNumber="6">O. brahma</emphasis>
<bibRefCitation id="B9494B35FFF389039B11FE05FB2DFE45" author="Lansberge G." box="[1040,1236,436,463]" pageId="5" pageNumber="6" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039BE2FE05FABBFE45" box="[1251,1346,436,463]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="5" pageNumber="6">Figs 5B</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFE05FA94FE44" box="[1359,1389,436,462]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="5" pageNumber="6">17</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFE69FA8FFE7B" blockId="5.[189,1398,401,498]" box="[189,1398,471,498]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFE69FA8FFE7B" box="[189,1398,471,498]" pageId="5" pageNumber="6">– Clypeal margin not deeply excised; frontal carina entire ................................................................ 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF389039FBCFDA8FA8FFD16" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFDA8FA8FFDDC" blockId="5.[189,1398,537,669]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFDA8FA8FFDDC" pageId="5" pageNumber="6">
16. Hair on the pygidium short; elytral interval angularly elevated along middle with fine close punctures................................................................................... 
<taxonomicName id="1AD84D47FFF389039CBEFD8DFB2BFDDD" ID-CoL="49KFD" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[959,1234,572,599]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFF389039CBEFD8DFBBDFDDC" bold="true" box="[959,1092,572,598]" italics="true" pageId="5" pageNumber="6">O. crassus</emphasis>
Sharp, 1875
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039BE0FD8DFABAFDDD" box="[1249,1347,572,599]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="5" pageNumber="6">Figs 5D</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFD8DFA94FDDC" box="[1359,1389,572,598]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="5" pageNumber="6">18</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFDD1FA8FFD16" blockId="5.[189,1398,537,669]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFDD1FA8FFD16" pageId="5" pageNumber="6">
– Hair on the pygidium long; elytral interval flat with feeble punctures................................................ .................................................................... 
<taxonomicName id="1AD84D47FFF389039DE5FD33FB89FD16" ID-CoL="6SN86" authority="Redtenbacher, 1848" authorityName="Redtenbacher" authorityYear="1848" box="[740,1136,642,669]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFF389039DE5FD33FC86FD17" bold="true" box="[740,895,642,669]" italics="true" pageId="5" pageNumber="6">O. castaneus</emphasis>
Redtenbacher, 1848
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B7EFD33FB26FD17" box="[1151,1247,642,669]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="5" pageNumber="6">Figs 5C</figureCitation>
, 
<figureCitation id="45E32A41FFF389039BEAFD33FABAFD16" box="[1259,1347,642,668]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. E–H. O. feae Felsche, 1907, AIMB/Co/Sc1000188. I–L. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742881" httpUri="https://zenodo.org/record/13742881/files/figure.png" pageId="5" pageNumber="6">11A–D</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFD33FA94FD16" box="[1359,1389,642,668]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="5" pageNumber="6">17</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF389039FBCFD75FA8FFC8B" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFD75FA8FFD54" blockId="5.[189,1398,708,769]" box="[189,1398,708,734]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFD75FA8FFD54" box="[189,1398,708,734]" pageId="5" pageNumber="6">17. Clypeal carina present ..................................................................................................................... 18</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFD56FA8FFC8B" blockId="5.[189,1398,708,769]" box="[189,1398,743,769]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFD56FA8FFC8B" box="[189,1398,743,769]" pageId="5" pageNumber="6">– Clypeal carina absent....................................................................................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF389039FBCFC99FA8FFC45" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFC99FA8FFCEC" blockId="5.[189,1398,808,976]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFC99FA8FFCEC" pageId="5" pageNumber="6">
18. First, third and fifth inter-striae of the elytra not much convex; elytra dark, metasternal shield smooth behind, with a median groove (
<emphasis id="EFACEAD6FFF389039D58FCFAFD24FCEC" bold="true" box="[601,733,843,870]" pageId="5" pageNumber="6">GROUP V</emphasis>
)............ 
<taxonomicName id="1AD84D47FFF389039C44FCFAFB89FCEC" ID-CoL="74N7T" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[837,1136,843,870]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFF389039C44FCFAFC5CFCEC" bold="true" box="[837,933,843,870]" italics="true" pageId="5" pageNumber="6">O. lama</emphasis>
<bibRefCitation id="B9494B35FFF389039CADFCFAFB89FCEC" author="Lansberge G." box="[940,1136,843,870]" pageId="5" pageNumber="6" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B7EFCFAFB17FCEC" box="[1151,1262,843,870]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="5" pageNumber="6">Figs 10C</figureCitation>
, 
<figureCitation id="45E32A41FFF389039BFBFCFDFABAFCEC" box="[1274,1347,844,870]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. E–H. O. feae Felsche, 1907, AIMB/Co/Sc1000188. I–L. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742881" httpUri="https://zenodo.org/record/13742881/files/figure.png" pageId="5" pageNumber="6">11I–L</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFCFDFA94FCEC" box="[1359,1389,844,870]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="5" pageNumber="6">19</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFCDEFA8FFC45" blockId="5.[189,1398,808,976]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFCDEFA8FFC45" pageId="5" pageNumber="6">
– First, third, and fifth inter-striae distinctly convex; elytra yellow with some intervals green; metasternal shield punctured behind, without a median groove (
<emphasis id="EFACEAD6FFF389039CD6FC23FB85FC26" bold="true" box="[983,1148,914,940]" pageId="5" pageNumber="6">GROUP XIV</emphasis>
) ............................... ....................................................................................... 
<taxonomicName id="1AD84D47FFF389039C70FC04FB3CFC45" ID-CoL="49KGS" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[881,1221,949,976]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFF389039C70FC04FBE2FC5A" bold="true" box="[881,1051,949,976]" italics="true" pageId="5" pageNumber="6">O. humerosus</emphasis>
(
<bibRefCitation id="B9494B35FFF389039B2BFC04FB45FC45" author="Pallas P. S." box="[1066,1212,949,975]" pageId="5" pageNumber="6" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039BD5FC04FABBFC45" box="[1236,1346,949,975]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="5" pageNumber="6">Figs 10B</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFC04FA94FC45" box="[1359,1389,949,975]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="5" pageNumber="6">18</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF389039FBCFC46FA8FFBF0" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFC46FA8FFBBE" blockId="5.[189,1398,1014,1147]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFC46FA8FFBBE" pageId="5" pageNumber="6">
19. Pronotum covered with close granules (
<emphasis id="EFACEAD6FFF389039DBCFC47FCB7FB9B" bold="true" box="[701,846,1014,1041]" pageId="5" pageNumber="6">GROUP IX</emphasis>
)......................................................................... ............................................................................ 
<taxonomicName id="1AD84D47FFF389039C1EFBABFB91FBBE" ID-CoL="74MVW" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[799,1128,1049,1076]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFF389039C1EFBABFC58FBBE" bold="true" box="[799,929,1049,1076]" italics="true" pageId="5" pageNumber="6">O. bordati</emphasis>
<bibRefCitation id="B9494B35FFF389039CA0FBABFB91FBBE" author="Cambefort Y." box="[929,1128,1050,1076]" pageId="5" pageNumber="6" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B76FBABFB3AFBBE" box="[1143,1219,1050,1076]" captionStart="Fig" captionStartId="11.[189,232,1953,1979]" captionTargetBox="[280,1294,921,1858]" captionTargetId="figure-246@11.[189,1398,900,1917]" captionTargetPageId="11" captionText="Fig. 3. Onitis bordati Cambefort, 1988, ♀, AIMB/Co/Sc1000018. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742859" httpUri="https://zenodo.org/record/13742859/files/figure.png" pageId="5" pageNumber="6">Figs 3</figureCitation>
, 
<figureCitation id="45E32A41FFF389039BD1FBABFAE8FBBE" box="[1232,1297,1050,1076]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="5" pageNumber="6">4G–J</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A1CFBABFABAFBBF" box="[1309,1347,1050,1077]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="5" pageNumber="6">5A</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFBABFA94FBBE" box="[1359,1389,1050,1076]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="5" pageNumber="6">17</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFB8FFA8FFBF0" blockId="5.[189,1398,1014,1147]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFB8FFA8FFBF0" pageId="5" pageNumber="6">
– Pronotum covered with strong punctures (
<emphasis id="EFACEAD6FFF389039DD8FB8CFC6CFBDD" bold="true" box="[729,917,1085,1111]" pageId="5" pageNumber="6">GROUP XVIII</emphasis>
)............................................................... .......................................................................................... 
<taxonomicName id="1AD84D47FFF389039C88FBD1FB96FBF0" ID-CoL="49KL8" authority="Gillet, 1911" authorityName="Gillet" authorityYear="1911" box="[905,1135,1120,1146]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFF389039C88FBD1FC25FBF0" bold="true" box="[905,988,1120,1146]" italics="true" pageId="5" pageNumber="6">O. siva</emphasis>
<bibRefCitation id="B9494B35FFF389039CE2FBD1FB96FBF0" author="Gillet J. J. E." box="[995,1135,1120,1146]" pageId="5" pageNumber="6" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B7FFBD1FAFDFBF0" box="[1150,1284,1120,1146]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="5" pageNumber="6">Figs 4K–N</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A11FBD1FABBFBF0" box="[1296,1346,1120,1146]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="5" pageNumber="6">12B</figureCitation>
, 
<figureCitation id="45E32A41FFF389039A4EFBD1FA97FBF0" box="[1359,1390,1120,1146]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="5" pageNumber="6">20</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
</key>
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</treatment>
</document>