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174 lines
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<document ID-DOI="10.1643/i2019299" ID-ISSN="2766-1520" ID-Zenodo-Dep="7846669" IM.illustrations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1681927084425" checkinUser="felipe" docAuthor="Armbruster, Jonathan W., Lujan, Nathan K. & Bloom, Devin D." docDate="2021" docId="0B02895DD579FFF9FF93F9113A9708CE" docLanguage="en" docName="IchthyolHerpetol.109.1.102-122.pdf" docOrigin="Ichthyology & Herpetology 109 (1)" docSource="http://dx.doi.org/10.1643/i2019299" docStyle="DocumentStyle:25DEBD253B07840CEE0E8BE8BE2C6752.1:IchthyolHerpetol.2021-.journal_article" docStyleId="25DEBD253B07840CEE0E8BE8BE2C6752" docStyleName="IchthyolHerpetol.2021-.journal_article" docStyleVersion="1" docTitle="Characidium Armbruster & Lujan & Bloom 2021" docType="key" docVersion="3" lastPageNumber="120" masterDocId="F73BF125D56BFFEBFFF4FFFF3F450C62" masterDocTitle="Redescription of the Guiana Shield Darter Species Characidium crandellii and C. declivirostre (Crenuchidae) with Descriptions of Two New Species" masterLastPageNumber="122" masterPageNumber="102" pageNumber="120" updateTime="1682004287626" updateUser="tatiana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Redescription of the Guiana Shield Darter Species Characidium crandellii and C. declivirostre (Crenuchidae) with Descriptions of Two New Species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Armbruster, Jonathan W.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lujan, Nathan K.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bloom, Devin D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Ichthyology & Herpetology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-03-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>109</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>102</mods:start>
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<mods:end>122</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1643/i2019299</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1643/i2019299</mods:identifier>
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<mods:identifier type="ISSN">2766-1520</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7846669</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:0B02895DD579FFF9FF93F9113A9708CE" httpUri="http://treatment.plazi.org/id/0B02895DD579FFF9FF93F9113A9708CE" lastPageNumber="120" pageId="18" pageNumber="120">
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<subSubSection pageId="18" pageNumber="120" type="nomenclature">
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<paragraph blockId="18.[103,773,1774,1826]" pageId="18" pageNumber="120">
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<heading allCaps="true" bold="true" centered="true" fontSize="9" level="1" pageId="18" pageNumber="120" reason="1">
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<emphasis bold="true" pageId="18" pageNumber="120">
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KEY TO SPECIES OF THE FAST-WATER
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<taxonomicName ID-CoL="3M73" ID-ENA="42560" authorityName="Armbruster & Lujan & Bloom" authorityYear="2021" box="[527,684,1774,1797]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="120" phylum="Chordata" rank="genus">
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<emphasis bold="true" box="[527,684,1774,1797]" italics="true" pageId="18" pageNumber="120">CHARACIDIUM</emphasis>
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</taxonomicName>
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OF THE GUIANA SHIELD
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="120" type="key">
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<keyStep pageId="18" pageNumber="120">
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<keyLead pageId="18" pageNumber="120">
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<paragraph blockId="18.[103,736,1855,2024]" pageId="18" pageNumber="120">
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1a. Twelve circumpeduncular scales. Dorsal fin falcate with a medial dark band and terminal white band (
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<figureCitation box="[172,244,1914,1936]" captionStart="FIG" captionStartId="3.[142,170,1639,1659]" captionTargetBox="[142,1528,178,1610]" captionTargetId="figure-171@3.[142,1529,178,1610]" captionTargetPageId="3" captionText="FIG. 4. Dorsal, lateral, and ventral views of Characidium crandellii in life, potential nuptial male, AUM 67142, 106.9 mm SL; Ireng River population. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846677" httpUri="https://zenodo.org/record/7846677/files/figure.png" pageId="18" pageNumber="120">Figs. 4</figureCitation>
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,
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<figureCitation box="[256,271,1914,1937]" captionStart="FIG" captionStartId="4.[103,131,1217,1237]" captionTargetBox="[103,1491,178,1188]" captionTargetId="figure-201@4.[103,1491,178,1188]" captionTargetPageId="4" captionText="FIG. 5. Dorsal and lateral views of Characidium crandellii in life, common coloration, AUM 62874, 57.0 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846679" httpUri="https://zenodo.org/record/7846679/files/figure.png" pageId="18" pageNumber="120">5</figureCitation>
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). Origin of adipose fin centered at vertical above insertion of posteriormost anal-fin ray (
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<figureCitation captionStart="FIG" captionStartId="3.[142,170,1639,1659]" captionTargetBox="[142,1528,178,1610]" captionTargetId="figure-171@3.[142,1529,178,1610]" captionTargetPageId="3" captionText="FIG. 4. Dorsal, lateral, and ventral views of Characidium crandellii in life, potential nuptial male, AUM 67142, 106.9 mm SL; Ireng River population. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846677" httpUri="https://zenodo.org/record/7846677/files/figure.png" pageId="18" pageNumber="120">Figs. 4</figureCitation>
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,
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<figureCitation box="[198,213,1972,1995]" captionStart="FIG" captionStartId="4.[103,131,1217,1237]" captionTargetBox="[103,1491,178,1188]" captionTargetId="figure-201@4.[103,1491,178,1188]" captionTargetPageId="4" captionText="FIG. 5. Dorsal and lateral views of Characidium crandellii in life, common coloration, AUM 62874, 57.0 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846679" httpUri="https://zenodo.org/record/7846679/files/figure.png" pageId="18" pageNumber="120">5</figureCitation>
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). Pectoral fin uniformly dark or peripherally dark with lighter center. Proximal dentary and
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</paragraph>
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<paragraph blockId="18.[882,1490,179,1196]" pageId="18" pageNumber="120">
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premaxillary teeth wide and tri- to pentacuspid (central cusp longest)
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<subScript attach="left" box="[1121,1167,220,228]" fontSize="3" pageId="18" pageNumber="120">_________</subScript>
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<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1191,1490,209,231]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="120" phylum="Chordata" rank="species" species="crandellii">
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<emphasis bold="true" box="[1191,1490,209,231]" italics="true" pageId="18" pageNumber="120">Characidium crandellii</emphasis>
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</taxonomicName>
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</paragraph>
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</keyLead>
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<paragraph blockId="18.[820,854,240,262]" lastBlockId="18.[882,1490,179,1196]" pageId="18" pageNumber="120">
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<keyLead pageId="18" pageNumber="120">
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1b. Ten circumpeduncular scales. Dorsal fin only slightly concave along distal margin and with two or more thin dark bands (
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<figureCitation box="[1144,1217,300,322]" captionStart="FIG" captionStartId="7.[142,170,1643,1663]" captionTargetBox="[142,1530,178,1614]" captionTargetId="figure-132@7.[142,1530,178,1614]" captionTargetPageId="7" captionText="FIG. 7. Dorsal, lateral, and ventral views of Characidium declivirostre in life, AUM 56680, 55.5 mm SL; Raudales Danto, Río Cuao, Amazonas, Venezuela. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846683" httpUri="https://zenodo.org/record/7846683/files/figure.png" pageId="18" pageNumber="120">Figs. 7</figureCitation>
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,
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<figureCitation box="[1231,1246,300,321]" captionStart="FIG" captionStartId="8.[103,131,1539,1559]" captionTargetBox="[103,1492,178,1510]" captionTargetId="figure-257@8.[103,1492,178,1510]" captionTargetPageId="8" captionText="FIG. 8. Dorsal, lateral, and ventral views of a specimen of Characidium declivirostre from the same stream as the type locality, AUM 36625, 50.1 mm SL. Scale ¼ 1 cm. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846685" httpUri="https://zenodo.org/record/7846685/files/figure.png" pageId="18" pageNumber="120">8</figureCitation>
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,
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<figureCitation box="[1260,1330,300,323]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="12.[103,131,1898,1918]" captionStartId-1="15.[142,170,1319,1339]" captionStartId-2="16.[103,131,1331,1351]" captionTargetBox-0="[156,1436,178,1868]" captionTargetBox-1="[142,1530,178,1290]" captionTargetBox-2="[103,1491,178,1301]" captionTargetId-0="figure-1@12.[156,1436,178,1868]" captionTargetId-1="figure-339@15.[142,1530,178,1290]" captionTargetId-2="figure-376@16.[103,1491,178,1301]" captionTargetPageId-0="12" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIG. 10. Lateral view (top) of Characidium duplicatum, new species, in life and dorsal, lateral, and ventral views after preservation, CSBD F-3614, holotype, 39.4 mm SL. Photos by J. W. Armbruster." captionText-1="FIG. 11. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, in life, CSBD F-3615, holotype, 72.6 mm SL. Photos by N. K. Lujan." captionText-2="FIG. 12. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi-0="http://doi.org/10.5281/zenodo.7846689" figureDoi-1="http://doi.org/10.5281/zenodo.7846691" figureDoi-2="http://doi.org/10.5281/zenodo.7846693" httpUri-0="https://zenodo.org/record/7846689/files/figure.png" httpUri-1="https://zenodo.org/record/7846691/files/figure.png" httpUri-2="https://zenodo.org/record/7846693/files/figure.png" pageId="18" pageNumber="120">10–12</figureCitation>
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). Origin of adipose fin at vertical over center of anal-fin base or more anterior. Pectoral fin light with at least one (usually two or three) dark bands made of separate elongate spots (
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<figureCitation box="[1057,1130,420,442]" captionStart="FIG" captionStartId="7.[142,170,1643,1663]" captionTargetBox="[142,1530,178,1614]" captionTargetId="figure-132@7.[142,1530,178,1614]" captionTargetPageId="7" captionText="FIG. 7. Dorsal, lateral, and ventral views of Characidium declivirostre in life, AUM 56680, 55.5 mm SL; Raudales Danto, Río Cuao, Amazonas, Venezuela. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846683" httpUri="https://zenodo.org/record/7846683/files/figure.png" pageId="18" pageNumber="120">Figs. 7</figureCitation>
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,
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<figureCitation box="[1143,1158,420,441]" captionStart="FIG" captionStartId="8.[103,131,1539,1559]" captionTargetBox="[103,1492,178,1510]" captionTargetId="figure-257@8.[103,1492,178,1510]" captionTargetPageId="8" captionText="FIG. 8. Dorsal, lateral, and ventral views of a specimen of Characidium declivirostre from the same stream as the type locality, AUM 36625, 50.1 mm SL. Scale ¼ 1 cm. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846685" httpUri="https://zenodo.org/record/7846685/files/figure.png" pageId="18" pageNumber="120">8</figureCitation>
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,
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<figureCitation box="[1171,1241,419,442]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="12.[103,131,1898,1918]" captionStartId-1="15.[142,170,1319,1339]" captionStartId-2="16.[103,131,1331,1351]" captionTargetBox-0="[156,1436,178,1868]" captionTargetBox-1="[142,1530,178,1290]" captionTargetBox-2="[103,1491,178,1301]" captionTargetId-0="figure-1@12.[156,1436,178,1868]" captionTargetId-1="figure-339@15.[142,1530,178,1290]" captionTargetId-2="figure-376@16.[103,1491,178,1301]" captionTargetPageId-0="12" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIG. 10. Lateral view (top) of Characidium duplicatum, new species, in life and dorsal, lateral, and ventral views after preservation, CSBD F-3614, holotype, 39.4 mm SL. Photos by J. W. Armbruster." captionText-1="FIG. 11. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, in life, CSBD F-3615, holotype, 72.6 mm SL. Photos by N. K. Lujan." captionText-2="FIG. 12. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi-0="http://doi.org/10.5281/zenodo.7846689" figureDoi-1="http://doi.org/10.5281/zenodo.7846691" figureDoi-2="http://doi.org/10.5281/zenodo.7846693" httpUri-0="https://zenodo.org/record/7846689/files/figure.png" httpUri-1="https://zenodo.org/record/7846691/files/figure.png" httpUri-2="https://zenodo.org/record/7846693/files/figure.png" pageId="18" pageNumber="120">10–12</figureCitation>
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). Proximal dentary and premaxillary teeth narrow, peg-like, tricuspid, but only central cusp conspicuous
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<subScript attach="left" box="[1269,1453,491,499]" fontSize="3" pageId="18" pageNumber="120">___________________________________</subScript>
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2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="18" pageNumber="120">
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<paragraph blockId="18.[820,853,511,532]" lastBlockId="18.[882,1490,179,1196]" pageId="18" pageNumber="120">
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<keyLead pageId="18" pageNumber="120">
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2a. Pelvic fin with two unbranched rays. Pectoral fin with four unbranched rays (
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<figureCitation box="[1195,1271,540,563]" captionStart="FIG" captionStartId="1.[142,170,670,690]" captionTargetBox="[142,1528,178,641]" captionTargetId="figure-704@1.[142,1528,178,641]" captionTargetPageId="1" captionText="FIG. 1. Thickened, keratinized pads on the undersurface of the anterior pectoral-fin rays of (A) Characidium wangyapoik, new species, CSBD F-3615, holotype, and (B) C. crandellii, AUM 67142. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846671" httpUri="https://zenodo.org/record/7846671/files/figure.png" pageId="18" pageNumber="120">Fig. 1B</figureCitation>
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)
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<subScript attach="left" fontSize="3" pageId="18" pageNumber="120">_________________________________ _____________________________________________</subScript>
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<emphasis bold="true" box="[1144,1490,571,593]" pageId="18" pageNumber="120">
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<taxonomicName authority="Armbruster & Lujan & Bloom, 2021" authorityName="Armbruster & Lujan & Bloom" authorityYear="2021" box="[1144,1325,571,593]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="120" phylum="Chordata" rank="species" species="duplicatum" status="new species">
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<emphasis bold="true" box="[1144,1325,571,593]" italics="true" pageId="18" pageNumber="120">C. duplicatum</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1340,1490,571,593]" pageId="18" pageNumber="120" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph blockId="18.[820,855,601,623]" lastBlockId="18.[882,1490,179,1196]" pageId="18" pageNumber="120">
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<keyLead pageId="18" pageNumber="120">
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2b. Pelvic fin with one unbranched ray. Pectoral fin with three unbranched rays (
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<figureCitation box="[1208,1285,631,654]" captionStart="FIG" captionStartId="1.[142,170,670,690]" captionTargetBox="[142,1528,178,641]" captionTargetId="figure-704@1.[142,1528,178,641]" captionTargetPageId="1" captionText="FIG. 1. Thickened, keratinized pads on the undersurface of the anterior pectoral-fin rays of (A) Characidium wangyapoik, new species, CSBD F-3615, holotype, and (B) C. crandellii, AUM 67142. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846671" httpUri="https://zenodo.org/record/7846671/files/figure.png" pageId="18" pageNumber="120">Fig. 1A</figureCitation>
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)
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<subScript attach="left" box="[1296,1453,643,651]" fontSize="3" pageId="18" pageNumber="120">______________________________</subScript>
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3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="18" pageNumber="120">
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<paragraph blockId="18.[820,853,663,684]" lastBlockId="18.[882,1490,179,1196]" pageId="18" pageNumber="120">
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<keyLead pageId="18" pageNumber="120">
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3a. Dorsal coloration consisting of prominent saddles that extend to a faint midlateral stripe that may appear as a series of blotches (stripe most prominent in preserved specimens). No accessory bars between saddles. Ventral portion of sides with few, rectangular to square blotches that do not extend dorsal to midlateral stripe (
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<figureCitation box="[1077,1150,842,864]" captionStart="FIG" captionStartId="7.[142,170,1643,1663]" captionTargetBox="[142,1530,178,1614]" captionTargetId="figure-132@7.[142,1530,178,1614]" captionTargetPageId="7" captionText="FIG. 7. Dorsal, lateral, and ventral views of Characidium declivirostre in life, AUM 56680, 55.5 mm SL; Raudales Danto, Río Cuao, Amazonas, Venezuela. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846683" httpUri="https://zenodo.org/record/7846683/files/figure.png" pageId="18" pageNumber="120">Figs. 7</figureCitation>
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,
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<figureCitation box="[1163,1176,842,865]" captionStart="FIG" captionStartId="8.[103,131,1539,1559]" captionTargetBox="[103,1492,178,1510]" captionTargetId="figure-257@8.[103,1492,178,1510]" captionTargetPageId="8" captionText="FIG. 8. Dorsal, lateral, and ventral views of a specimen of Characidium declivirostre from the same stream as the type locality, AUM 36625, 50.1 mm SL. Scale ¼ 1 cm. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846685" httpUri="https://zenodo.org/record/7846685/files/figure.png" pageId="18" pageNumber="120">8</figureCitation>
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)
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<subScript attach="left" box="[1190,1278,854,862]" fontSize="3" pageId="18" pageNumber="120">_________________</subScript>
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<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1303,1490,843,865]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="120" phylum="Chordata" rank="species" species="declivirostre">
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<emphasis bold="true" box="[1303,1490,843,865]" italics="true" pageId="18" pageNumber="120">C. declivirostre</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="18.[820,855,873,895]" lastBlockId="18.[882,1490,179,1196]" pageId="18" pageNumber="120">
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<keyLead pageId="18" pageNumber="120">
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3b. Dorsal saddles not prominent in adults, and usually confluent with thin bands from below midlateral stripe, which is usually demarcated by a faint stripe made of zig-zag lines with the stripe most prominent in smaller individuals. Thin bars also present in lighter interspaces between saddles, some of which extend below midlateral stripe (
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<figureCitation box="[1312,1398,1053,1075]" captionStart="FIG" captionStartId="15.[142,170,1319,1339]" captionTargetBox="[142,1530,178,1290]" captionTargetId="figure-339@15.[142,1530,178,1290]" captionTargetPageId="15" captionText="FIG. 11. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, in life, CSBD F-3615, holotype, 72.6 mm SL. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846691" httpUri="https://zenodo.org/record/7846691/files/figure.png" pageId="18" pageNumber="120">Figs. 11</figureCitation>
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,
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<figureCitation box="[1410,1440,1053,1076]" captionStart="FIG" captionStartId="16.[103,131,1331,1351]" captionTargetBox="[103,1491,178,1301]" captionTargetId="figure-376@16.[103,1491,178,1301]" captionTargetPageId="16" captionText="FIG. 12. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846693" httpUri="https://zenodo.org/record/7846693/files/figure.png" pageId="18" pageNumber="120">12</figureCitation>
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). Ventral portion of sides with numerous narrow bars that extend above midlateral stripe (in juveniles, only some bars may continue dorsal to midlateral stripe;
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<figureCitation box="[959,1045,1173,1195]" captionStart="FIG" captionStartId="15.[142,170,1319,1339]" captionTargetBox="[142,1530,178,1290]" captionTargetId="figure-339@15.[142,1530,178,1290]" captionTargetPageId="15" captionText="FIG. 11. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, in life, CSBD F-3615, holotype, 72.6 mm SL. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846691" httpUri="https://zenodo.org/record/7846691/files/figure.png" pageId="18" pageNumber="120">Figs. 11</figureCitation>
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,
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<figureCitation box="[1058,1085,1173,1196]" captionStart="FIG" captionStartId="16.[103,131,1331,1351]" captionTargetBox="[103,1491,178,1301]" captionTargetId="figure-376@16.[103,1491,178,1301]" captionTargetPageId="16" captionText="FIG. 12. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846693" httpUri="https://zenodo.org/record/7846693/files/figure.png" pageId="18" pageNumber="120">12</figureCitation>
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)
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<subScript attach="left" box="[1095,1115,1185,1193]" fontSize="3" pageId="18" pageNumber="120">____</subScript>
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<emphasis bold="true" box="[1142,1490,1174,1196]" pageId="18" pageNumber="120">
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<emphasis bold="true" box="[1142,1332,1174,1196]" italics="true" pageId="18" pageNumber="120">
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<taxonomicName authority="Armbruster & Lujan & Bloom, 2021" authorityName="Armbruster & Lujan & Bloom" authorityYear="2021" box="[1142,1326,1174,1196]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="120" phylum="Chordata" rank="species" species="wangyapoik" status="new species">C. wangyapoik</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel box="[1340,1490,1174,1196]" pageId="18" pageNumber="120" rank="species">new species</taxonomicNameLabel>
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</emphasis>
|
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</keyLead>
|
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document>
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