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<document id="17C40E790C0A7DFAAF8DECA266D68DC1" ID-CLB-Dataset="3305" ID-DOI="10.5852/ejt.2021.779.1579" ID-GBIF-Dataset="c66d7415-6743-496d-a518-e5fbd4b71a22" ID-ISSN="2118-9773" ID-Zenodo-Dep="5724311" ID-ZooBank="A60F479C-8055-4AC9-8F49-44311CA6EA0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637760838361" checkinUser="felipe" docAuthor="Salazar-Vallejo, Sergio I., León-González, Jesús Angel De &amp; Conde-Vela, Víctor M." docDate="2021" docId="03EF3A1F8A224C0BFE15282281267291" docLanguage="en" docName="ejt-779_1-70.pdf" docOrigin="European Journal of Taxonomy 779" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nereis ambigua Treadwell 1937" docType="treatment" docVersion="11" lastPageNumber="49" masterDocId="FFD642678A0C4C3BFC0F2C628041740A" masterDocTitle="Revision of the species confused with “ Nereis falsa ” de Quatrefages, 1866 (Annelida, Nereididae)" masterLastPageNumber="70" masterPageNumber="1" pageNumber="47" updateTime="1699469980804" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EA4C2237BA9EC34F922C0CDB67046573">Revision of the species confused with “ Nereis falsa ” de Quatrefages, 1866 (Annelida, Nereididae)</mods:title>
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<mods:name id="D16C345086B738D82EF01F48BE8B394D" type="personal">
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<mods:roleTerm id="E122F3769A31BC63D0606BD0C5D8DA4A">Author</mods:roleTerm>
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<mods:namePart id="8274846D0D6F4DA1516724714E25B8A7">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:affiliation id="B61D90E813D190676CA97C824FDEE108">El Colegio de la Frontera Sur, Departamento Sistemática y Ecología Acuática, Chetumal, Quintana Roo, Mexico.</mods:affiliation>
<mods:nameIdentifier id="B5F0ADF5E01A9711D60646CE8DBCE084" type="email">savs551216@hotmail.com,ssalazar@ecosur.mx</mods:nameIdentifier>
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<mods:namePart id="30F4F73B037800ADE06E4B7ED4C87242">León-González, Jesús Angel De</mods:namePart>
<mods:nameIdentifier id="DF6B2743F267487ECC3EDC00A5C28E6C" type="LSID">F347D71F-DDEA-4105-953F-4FE71A5D6809</mods:nameIdentifier>
<mods:affiliation id="0F3FBBF259221F6CB367AAB8FD37B9E8">Universidad Autónoma de Nuevo León, Facultad de Ciencias Biológicas, Laboratorio Biosistemática, San Nicolás de los Garza, N. L., Mexico.</mods:affiliation>
<mods:nameIdentifier id="0AC4DA2697A52C6EDFD88B33ECE0CA17" type="email">jesus.deleongn@uanl.edu.mx</mods:nameIdentifier>
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<mods:namePart id="5ED16A78C5824C43C164C2332C891FE4">Conde-Vela, Víctor M.</mods:namePart>
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<mods:affiliation id="BB2CD3AFC8AA495BA6DB53BAD3093DF5">El Colegio de la Frontera Sur, Departamento Sistemática y Ecología Acuática, Chetumal, Quintana Roo, Mexico. &amp; Universidad Autónoma de Nuevo León, Facultad de Ciencias Biológicas, Laboratorio Biosistemática, San Nicolás de los Garza, N. L., Mexico.</mods:affiliation>
<mods:nameIdentifier id="69C5F9D214FFDDCD8F26D6F05F560F9C" type="email">victorconde2323@gmail.com</mods:nameIdentifier>
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<mods:title id="F1280A4A2A95E66F737E478D773097F9">European Journal of Taxonomy</mods:title>
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<mods:date id="0453FECF060199E29F3E4B110A882D3A">2021</mods:date>
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<mods:number id="F393FC8A05EEE7AA52639DF016D8CB79">2021-11-24</mods:number>
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<mods:number id="A68C14E0BCE037C7DF44D999003BAC0A">779</mods:number>
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<mods:start id="3307466A84BCE6E52A90DD9C8557024C">1</mods:start>
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<mods:classification id="28B40C23112A8166B902786A479FDA14">journal article</mods:classification>
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<treatment id="03EF3A1F8A224C0BFE15282281267291" ID-DOI="http://doi.org/10.5281/zenodo.5727444" ID-GBIF-Taxon="190703783" ID-Zenodo-Dep="5727444" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291" httpUri="http://treatment.plazi.org/id/03EF3A1F8A224C0BFE15282281267291" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="C35CD8828A224C15FE15282284587050" box="[538,1049,1088,1115]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BF98B098A224C15FE15282284587050" blockId="46.[538,1049,1088,1115]" box="[538,1049,1088,1115]" pageId="46" pageNumber="47">
<heading id="D0B13C658A224C15FE15282284587050" box="[538,1049,1088,1115]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="4C46F08A8A224C15FE15282283E17050" ID-CoL="62dd322d-2b38-4f8c-998d-0c6b51070e42" authority="Treadwell, 1937" authorityName="Treadwell" authorityYear="1937" box="[538,928,1088,1115]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A224C15FE152822829B7051" bold="true" box="[538,730,1088,1115]" italics="true" pageId="46" pageNumber="47">Nereis ambigua</emphasis>
<bibRefCitation id="EFD7F6F88A224C15FEEE282283E17050" author="Treadwell A. L." box="[737,928,1088,1114]" pageId="46" pageNumber="47" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell, 1937</bibRefCitation>
</taxonomicName>
reinstated
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CD8828A224C15FCB2281983717132" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8BF98B098A224C15FCB2281983CA709E" blockId="46.[189,907,1146,1173]" box="[189,907,1146,1173]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB56AC278A224C15FCB2281983CA709E" box="[189,907,1146,1173]" pageId="46" pageNumber="47">
<treatmentCitation id="0AE7AD188A224C15FCB228198283709F" author="Treadwell A. L." box="[189,706,1146,1173]" page="150" pageId="46" pageNumber="47" year="1937">
<taxonomicName id="4C46F08A8A224C15FCB228198283709F" ID-CoL="474GT" authority="Treadwell, 1937: 149 - 150" authorityName="Treadwell" authorityPageNumber="149 - 150" authorityYear="1937" box="[189,706,1146,1173]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="ambiguus">
<emphasis id="B932571B8A224C15FCB2281981C3709E" box="[189,386,1146,1172]" italics="true" pageId="46" pageNumber="47">Nereis ambiguus</emphasis>
<bibRefCitation id="EFD7F6F88A224C15FD8628188283709F" author="Treadwell A. L." box="[393,706,1146,1173]" pageId="46" pageNumber="47" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell, 1937: 149–150</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 2 figs 19–24.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF98B098A224C15FCB228D783717132" blockId="46.[189,1398,1204,1338]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB56AC278A224C15FCB228D783717132" pageId="46" pageNumber="47">
<taxonomicName id="4C46F08A8A224C15FCB228D7810F70C4" ID-CoL="474TJ" authority="Monro 1933" box="[189,334,1205,1230]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="riisei">
<emphasis id="B932571B8A224C15FCB228D7810F70C4" box="[189,334,1205,1230]" italics="true" pageId="46" pageNumber="47">Nereis riisei</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE7AD188A224C15FD7F28D7822170C4" author="Monro C. C. A." box="[368,608,1204,1231]" page="43" pageId="46" pageNumber="47" year="1933">
<bibRefCitation id="EFD7F6F88A224C15FD7F28D7822170C4" author="Monro C. C. A." box="[368,608,1204,1231]" pageId="46" pageNumber="47" pagination="1 - 96" refId="ref44267" refString="Monro C. C. A. 1933. The Polychaeta Errantia collected by Dr. Crossland at Colon, in the Panama Region, and the Galapagos Islands during the expedition of the S. Y. &quot; St. George. &quot; Proceedings of the Zoological Society of London 103 (1): 1 - 96. https: // doi. org / 10.1111 / j. 1096 - 3642.1933. tb 01578. x" type="journal article" year="1933">Monro 1933: 43–44</bibRefCitation>
</treatmentCitation>
(living pigmentation). —
<treatmentCitation id="0AE7AD188A224C15FF8428D784F070C4" author="Hartman O." box="[907,1201,1204,1231]" page="221" pageId="46" pageNumber="47" year="1940">
<bibRefCitation id="EFD7F6F88A224C15FF8428D784F070C4" author="Hartman O." box="[907,1201,1204,1231]" pageId="46" pageNumber="47" pagination="173 - 287" refId="ref43181" refString="Hartman O. 1940. Polychaetous annelids, 2. Chrysopetalidae to Goniadidae. Allan Hancock Pacific Expeditions 7: 173 - 287. Available from https: // www. biodiversitylibrary. org / page / 27822405 # page / 191 / mode / 1 up [accessed 16 Sep. 2021]." type="journal article" year="1940">Hartman 1940: 221–222</bibRefCitation>
</treatmentCitation>
, pl. 33 fig. 37;
<treatmentCitation id="0AE7AD188A224C15FCE528B5812370FB" author="Hartman" box="[234,354,1239,1266]" page="279" pageId="46" pageNumber="47" year="1956">1956: 279</treatmentCitation>
(synonymy). —
<treatmentCitation id="0AE7AD188A224C15FE1228B5829470F8" author="Reish D. J." box="[541,725,1239,1266]" page="77" pageId="46" pageNumber="47" year="1968">
<bibRefCitation id="EFD7F6F88A224C15FE1228B5829470F8" author="Reish D. J." box="[541,725,1239,1266]" pageId="46" pageNumber="47" pagination="67 - 106" refId="ref45158" refString="Reish D. J. 1968. A biological survey of Bahia de Los Angeles, Gulf of California, Mexico, 2. Benthic polychaetous annelids. Transactions of the San Diego Society of Natural History 15 (7): 67 - 106. https: // doi. org / 10.5962 / bhl. part. 12054" type="journal article" year="1968">Reish 1968: 77</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AE7AD188A224C15FEFB28B583A370FB" author="Fauchald K." box="[756,994,1239,1265]" page="31" pageId="46" pageNumber="47" year="1977">
<bibRefCitation id="EFD7F6F88A224C15FEFB28B583A370FB" author="Fauchald K." box="[756,994,1239,1265]" pageId="46" pageNumber="47" pagination="1 - 81" refId="ref41743" refString="Fauchald K. 1977 b. Polychaetes from intertidal areas in Panama, with a review of previous shallow-water records. Smithsonian Contributions to Zoology 221: 1 - 81. https: // doi. org / 10.5479 / si. 00810282.221" type="journal article" year="1977">Fauchald 1977b: 31</bibRefCitation>
</treatmentCitation>
, fig. 8c–e (partim, only Pacific
<collectingCountry id="F351CB998A224C15FCE528998106711F" box="[234,327,1275,1301]" name="Panama" pageId="46" pageNumber="47">Panama</collectingCountry>
specimens). —
<treatmentCitation id="0AE7AD188A224C15FDF52898840B711F" author="De Leon-Gonzalez J. A. &amp; Solis-Weiss V." box="[506,1098,1274,1301]" page="560" pageId="46" pageNumber="47" year="2000">
<bibRefCitation id="EFD7F6F88A224C15FDF52898840B711F" author="De Leon-Gonzalez J. A. &amp; Solis-Weiss V." box="[506,1098,1274,1301]" pageId="46" pageNumber="47" pagination="549 - 569" refId="ref41266" refString="De Leon-Gonzalez J. A. &amp; Solis-Weiss V. 2000. A review of the polychaete family Nereididae from Western Mexico. Bulletin of Marine Science 67: 549 - 569." type="journal article" year="2000">de León-González &amp; Solís-Weiss 2000: 560–561</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AE7AD188A224C15F8642899850D711F" author="Dean H. K." box="[1131,1356,1274,1301]" page="57" pageId="46" pageNumber="47" year="2001">
<bibRefCitation id="EFD7F6F88A224C15F8642899850D711F" author="Dean H. K." box="[1131,1356,1274,1301]" pageId="46" pageNumber="47" pagination="37 - 67" refId="ref41350" refString="Dean H. K. 2001. Some Nereididae (Annelida: Polychaeta) from the Pacific coast of Costa Rica. Revista de Biologia Tropical 49 (Suppl. 2): 37 - 67." type="journal article" year="2001">Dean 2001: 57–58</bibRefCitation>
</treatmentCitation>
, figs 44–48 (non Grube &amp; Örsted in
<bibRefCitation id="EFD7F6F88A224C15FE83297C83627132" author="Grube A. E." box="[652,803,1309,1336]" pageId="46" pageNumber="47" pagination="158 - 186" refId="ref42608" refString="Grube A. E. 1857. Annulata orstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845 - 1848 suscepto legit cl. A. S. Orsted, adjectis speciebus nonnullis a cl. H. Kroyero in itinere ad Americam meridionalem collectis, (Fortsaettelse [continued]). 2. Familia Euniceae - F. Syllidea. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening for 1857: 158 - 186. Available from https: // www. biodiversitylibrary. org / item / 110674 # page / 159 / mode / 1 up [accessed 16 Sep. 2021]." type="journal article" year="1857">Grube, 1857</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35CD8828A224C15FCB22935821C71EC" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BF98B098A224C15FCB229358177717B" blockId="46.[189,310,1367,1393]" box="[189,310,1367,1393]" pageId="46" pageNumber="47">
<heading id="D0B13C658A224C15FCB229358177717B" bold="true" box="[189,310,1367,1393]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B932571B8A224C15FCB229358177717B" bold="true" box="[189,310,1367,1393]" pageId="46" pageNumber="47">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF98B098A224C15FCB229E5821C71EC" blockId="46.[189,1398,1414,1510]" pageId="46" pageNumber="47">
<taxonomicName id="4C46F08A8A224C15FCB229E5814871AA" authorityName="Linnaeus" authorityYear="1758" box="[189,265,1415,1440]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="46" pageNumber="47" phylum="Annelida" rank="genus">
<emphasis id="B932571B8A224C15FCB229E5814871AA" box="[189,265,1415,1440]" italics="true" pageId="46" pageNumber="47">Nereis</emphasis>
</taxonomicName>
with posterior eyes completely exposed; tentacular belt 2 × as long as first chaetiger; pharyngeal areas VII–VIII with a single row of large denticles; median and posterior parapodia with ventral ligules reaching tip of neuracicular ligules.
</paragraph>
</subSubSection>
<subSubSection id="C35CD8828A224C15FCB22A6483CF73E8" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BF98B098A224C15FCB22A6481EB722A" blockId="46.[189,426,1542,1568]" box="[189,426,1542,1568]" pageId="46" pageNumber="47">
<heading id="D0B13C658A224C15FCB22A6481EB722A" bold="true" box="[189,426,1542,1568]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B932571B8A224C15FCB22A6481EB722A" bold="true" box="[189,426,1542,1568]" pageId="46" pageNumber="47">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF98B098A224C15FCE52A56841572B3" blockId="46.[189,1399,1588,1721]" pageId="46" pageNumber="47">
<materialsCitation id="3B2E81548A224C15FCE52A568525729C" ID-GBIF-Occurrence="3411820380" collectingDate="1936-05-11" collectionCode="AMNH" country="Mexico" county="Templeton Crocker Expedition" elevation="0" latitude="18.344444" location="Revillagigedo Islands" longLatPrecision="21" longitude="-114.738884" municipality="Clarion Island" pageId="46" pageNumber="47" specimenCode="AMNH 36595" specimenCount="1" typeStatus="syntype">
<emphasis id="B932571B8A224C15FCE52A56811B7244" bold="true" box="[234,346,1588,1614]" pageId="46" pageNumber="47">
<typeStatus id="54FD35AB8A224C15FCE52A56811B7244" box="[234,346,1588,1614]" pageId="46" pageNumber="47" type="syntype">Syntypes</typeStatus>
</emphasis>
<collectingCountry id="F351CB998A224C15FCB22A3A81707278" box="[189,305,1624,1650]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
•
<specimenCount id="9D4040808A224C15FD5F2A3A81D97278" box="[336,408,1624,1650]" pageId="46" pageNumber="47" type="generic">1 juv.</specimenCount>
;
<location id="8E99DDD28A224C15FDA92A3A82EA7278" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FDA92A3A82EA7278" box="[422,683,1624,1650]" country="Mexico" county="Templeton Crocker Expedition" latitude="18.344444" longLatPrecision="21" longitude="-114.738884" municipality="Clarion Island" name="Revillagigedo Islands" pageId="46" pageNumber="47">Revillagigedo Islands</location>
,
<location id="8E99DDD28A224C15FEB52A3A83297278" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FEB52A3A83297278" box="[698,872,1624,1650]" country="Mexico" county="Templeton Crocker Expedition" latitude="18.344444" longLatPrecision="21" longitude="-114.738884" municipality="Clarion Island" name="Clarion Island" pageId="46" pageNumber="47">Clarion Island</location>
,
<location id="8E99DDD28A224C15FF792A3A844C7279" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FF792A3A844C7279" box="[886,1037,1624,1651]" country="Mexico" county="Templeton Crocker Expedition" latitude="18.344444" longLatPrecision="21" longitude="-114.738884" municipality="Clarion Island" name="Sulphur Bay" pageId="46" pageNumber="47">Sulphur Bay</location>
,
<collectingMunicipality id="6B9D11738A224C15F8132A3A848B7278" box="[1052,1226,1624,1650]" pageId="46" pageNumber="47">Clarion Island</collectingMunicipality>
;
<geoCoordinate id="EE72EDCE8A224C15F8D62A3485337279" box="[1241,1394,1622,1651]" degrees="18" direction="north" minutes="20" orientation="latitude" pageId="46" pageNumber="47" precision="15" seconds="40" value="18.344444">18°20′40″ N</geoCoordinate>
,
<geoCoordinate id="EE72EDCE8A224C15FCB22A188127729C" box="[189,358,1658,1686]" degrees="114" direction="west" minutes="44" orientation="longitude" pageId="46" pageNumber="47" precision="15" seconds="20" value="-114.738884">114°44′20″ W</geoCoordinate>
;
<collectingCounty id="6298F3858A224C15FD7D2A1982A1729F" box="[370,736,1659,1685]" pageId="46" pageNumber="47">Templeton Crocker Expedition</collectingCounty>
, stn 163 D1;
<elevation id="006B6C3A8A224C15FF732A198445729C" box="[892,1028,1659,1686]" pageId="46" pageNumber="47">
depth
<quantity id="4CBE26EC8A224C15FFCA2A198445729C" box="[965,1028,1659,1686]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="46" pageNumber="47" unit="m" value="36.0">36 m</quantity>
</elevation>
;
<date id="FFF8ADC98A224C15F81F2A1984EE729F" box="[1040,1199,1659,1686]" pageId="46" pageNumber="47" value="1936-05-11">
<collectingDate id="EFBC54218A224C15F81F2A1984EE729F" box="[1040,1199,1659,1686]" pageId="46" pageNumber="47" value="1936-05-11">11 May 1936</collectingDate>
</date>
;
<specimenCode id="DBE023728A224C15F8B52A198525729C" box="[1210,1380,1659,1686]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="46" pageNumber="47">AMNH 36595</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B2E81548A224C15FCB22AFD841172B3" ID-GBIF-Occurrence="3411820349" box="[189,1104,1694,1721]" collectingDate="1936-05-11" collectionCode="AMNH" country="Mexico" county="Templeton Crocker Expedition" elevation="36" latitude="18.344444" location="Revillagigedo Islands" longLatPrecision="21" longitude="-114.738884" municipality="Clarion Island" pageId="46" pageNumber="47" specimenCode="AMNH 36883" specimenCount="1" typeStatus="syntype">
<specimenCount id="9D4040808A224C15FCB22AFD808D72B3" box="[189,204,1695,1721]" pageId="46" pageNumber="47">1</specimenCount>
heteroneris; same collection data as for preceding;
<specimenCode id="DBE023728A224C15FF292AFD839172B2" box="[806,976,1694,1721]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="46" pageNumber="47">AMNH 36883</specimenCode>
(not seen)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF98B098A224C15FCE52AB281E772E0" blockId="46.[189,1400,1744,2018]" box="[234,422,1744,1770]" pageId="46" pageNumber="47">
<emphasis id="B932571B8A224C15FCE52AB281E772E0" bold="true" box="[234,422,1744,1770]" pageId="46" pageNumber="47">Other material</emphasis>
</paragraph>
<paragraph id="8BF98B098A224C15FCB22A9683CF73E8" blockId="46.[189,1400,1744,2018]" pageId="46" pageNumber="47">
<materialsCitation id="3B2E81548A224C15FCB22A968402733B" ID-GBIF-Occurrence="3411820327" collectingDate="2010-04-18" collectionCode="UANL" collectorName="J. A. de Leon" country="Mexico" latitude="28.947166" location="Gulf of California" longLatPrecision="1" longitude="-113.55717" pageId="46" pageNumber="47" specimenCode="UANL 8146" specimenCount="3" stateProvince="Baja California">
<collectingCountry id="F351CB998A224C15FCB22A9681707304" box="[189,305,1780,1806]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
–
<emphasis id="B932571B8A224C15FD562A9682057304" bold="true" box="[345,580,1780,1806]" pageId="46" pageNumber="47">
<location id="8E99DDD28A224C15FD562A9682057304" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FD562A9682057304" box="[345,580,1780,1806]" country="Mexico" latitude="28.947166" longLatPrecision="1" longitude="-113.55717" name="Gulf of California" pageId="46" pageNumber="47" stateProvince="Baja California">Gulf of California</location>
</emphasis>
•
<specimenCount id="9D4040808A224C15FE682A9682877304" box="[615,710,1780,1806]" pageId="46" pageNumber="47">3 specs</specimenCount>
;
<collectingRegion id="498245EB8A224C15FED82A9683D27304" box="[727,915,1780,1806]" country="Mexico" name="Baja California" pageId="46" pageNumber="47">Baja California</collectingRegion>
,
<location id="8E99DDD28A224C15FFAB2A9684F67304" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FFAB2A9684F67304" box="[932,1207,1780,1806]" country="Mexico" latitude="28.947166" longLatPrecision="1" longitude="-113.55717" name="Bahia de Los Angeles" pageId="46" pageNumber="47" stateProvince="Baja California">Bahía de Los Angeles</location>
, fiscal harbour;
<geoCoordinate id="EE72EDCE8A224C15FCB22B7481297338" box="[189,360,1814,1842]" degrees="28" direction="north" minutes="56" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="49.8" value="28.947166">28°56′49.8″ N</geoCoordinate>
,
<geoCoordinate id="EE72EDCE8A224C15FD7C2B74826F7338" box="[371,558,1814,1842]" degrees="113" direction="west" minutes="33" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="25.8" value="-113.55717">113°33′25.8″ W</geoCoordinate>
,
<date id="FFF8ADC98A224C15FE362B758292733B" box="[569,723,1815,1841]" pageId="46" pageNumber="47" value="2010-04-18">
<collectingDate id="EFBC54218A224C15FE362B758292733B" box="[569,723,1815,1841]" pageId="46" pageNumber="47" value="2010-04-18">18 Apr. 2010</collectingDate>
</date>
;
<collectorName id="26B3EEDF8A224C15FED12B758330733B" box="[734,881,1815,1841]" pageId="46" pageNumber="47">J.A. de León</collectorName>
leg.;
<specimenCode id="DBE023728A224C15FFBD2B758402733B" box="[946,1091,1815,1841]" pageId="46" pageNumber="47">UANL 8146</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B2E81548A224C15F86F2B7580B97372" ID-GBIF-Occurrence="3411820362" collectingDate="2011-04-02" collectionCode="UANL" collectorName="J. A. de Leon" country="Mexico" latitude="25.814444" location="Puerto Escondido Marina" longLatPrecision="15" longitude="-111.31142" pageId="46" pageNumber="47" specimenCode="UANL 8147" specimenCount="3" stateProvince="Baja California Sur">
<specimenCount id="9D4040808A224C15F86F2B7584F97338" box="[1120,1208,1815,1842]" pageId="46" pageNumber="47">3 specs</specimenCount>
;
<collectingRegion id="498245EB8A224C15F8CC2B7580A6735E" country="Mexico" name="Baja California Sur" pageId="46" pageNumber="47">Baja California Sur</collectingRegion>
,
<location id="8E99DDD28A224C15FCFA2B598269735E" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FCFA2B598269735E" box="[245,552,1850,1877]" country="Mexico" latitude="25.814444" longLatPrecision="15" longitude="-111.31142" name="Puerto Escondido Marina" pageId="46" pageNumber="47" stateProvince="Baja California Sur">Puerto Escondido Marina</location>
,
<geoCoordinate id="EE72EDCE8A224C15FE392B5B828C735F" box="[566,717,1849,1877]" degrees="25" direction="north" minutes="48" orientation="latitude" pageId="46" pageNumber="47" precision="15" seconds="52" value="25.814444">25°48′52″ N</geoCoordinate>
,
<geoCoordinate id="EE72EDCE8A224C15FED42B5B83DA735F" box="[731,923,1849,1877]" degrees="111" direction="west" minutes="18" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="41.1" value="-111.31142">111°18′41.1″ W</geoCoordinate>
;
<date id="FFF8ADC98A224C15FFA62B588479735E" box="[937,1080,1850,1877]" pageId="46" pageNumber="47" value="2011-04-02">
<collectingDate id="EFBC54218A224C15FFA62B588479735E" box="[937,1080,1850,1877]" pageId="46" pageNumber="47" value="2011-04-02">2 Apr. 2011</collectingDate>
</date>
,
<collectorName id="26B3EEDF8A224C15F8492B5984A1735E" box="[1094,1248,1850,1877]" pageId="46" pageNumber="47">J.A. de León</collectorName>
leg.;
<specimenCode id="DBE023728A224C15F9292B5980B97372" pageId="46" pageNumber="47">UANL 8147</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B2E81548A224C15FD1C2B3C84BD7372" ID-GBIF-Occurrence="3411820319" box="[275,1276,1885,1912]" collectingDate="2011-09-02" collectionCode="UANL" collectorName="J. A. de Leon" country="Mexico" latitude="25.814444" location="Puerto Escondido Marina" longLatPrecision="15" longitude="-111.31142" pageId="46" pageNumber="47" specimenCode="UANL 8148" specimenCount="3" stateProvince="Gulf of California">
<specimenCount id="9D4040808A224C15FD1C2B3C812B7372" box="[275,362,1886,1912]" pageId="46" pageNumber="47" type="generic">3 spec.</specimenCount>
; same locality as for preceding;
<date id="FFF8ADC98A224C15FEE22B3C833C7372" box="[749,893,1886,1912]" pageId="46" pageNumber="47" value="2011-09-02">
<collectingDate id="EFBC54218A224C15FEE22B3C833C7372" box="[749,893,1886,1912]" pageId="46" pageNumber="47" value="2011-09-02">2 Sep. 2011</collectingDate>
</date>
,
<collectorName id="26B3EEDF8A224C15FF852B3C84627372" box="[906,1059,1885,1912]" pageId="46" pageNumber="47">J.A. de León</collectorName>
leg.;
<specimenCode id="DBE023728A224C15F8672B3C84BD7372" box="[1128,1276,1886,1912]" pageId="46" pageNumber="47">UANL 8148</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B2E81548A224C15F9182B3F84037391" ID-GBIF-Occurrence="3411820313" collectingDate="2013-11-05" collectionCode="UANL" collectorName="J. A. de Leon" country="Mexico" latitude="25.814444" location="Puerto Escondido Marina" longLatPrecision="15" longitude="-111.31142" pageId="46" pageNumber="47" specimenCode="UANL 8149" specimenCount="6" stateProvince="Gulf of California">
<specimenCount id="9D4040808A224C15F9182B3F85337372" box="[1303,1394,1885,1912]" pageId="46" pageNumber="47">6 specs</specimenCount>
; same locality as for preceding;
<date id="FFF8ADC98A224C15FE3D2BE382877391" box="[562,710,1921,1948]" pageId="46" pageNumber="47" value="2013-11-05">
<collectingDate id="EFBC54218A224C15FE3D2BE382877391" box="[562,710,1921,1948]" pageId="46" pageNumber="47" value="2013-11-05">5 Nov. 2013</collectingDate>
</date>
,
<collectorName id="26B3EEDF8A224C15FEDC2BE3832B7391" box="[723,874,1921,1947]" pageId="46" pageNumber="47">J.A. de León</collectorName>
leg.;
<specimenCode id="DBE023728A224C15FFA12BE384037391" box="[942,1090,1921,1947]" pageId="46" pageNumber="47">UANL 8149</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B2E81548A224C15F8532BE380B973E8" ID-GBIF-Occurrence="3411820307" collectingDate="2013-11-12" collectionCode="UANL" collectorName="J. A. de Leon" country="Mexico" latitude="27.337057" location="Santa Rosalia Marina" longLatPrecision="1" longitude="-112.26322" pageId="46" pageNumber="47" specimenCode="UANL 8150" specimenCount="3" stateProvince="Baja California Sur">
<specimenCount id="9D4040808A224C15F8532BE384F77391" box="[1116,1206,1921,1947]" pageId="46" pageNumber="47">3 specs</specimenCount>
;
<collectingRegion id="498245EB8A224C15F8CC2BE380A973B4" country="Mexico" name="Baja California Sur" pageId="46" pageNumber="47">Baja California Sur</collectingRegion>
,
<location id="8E99DDD28A224C15FCF92BC681BA73B4" LSID="urn:lsid:plazi:treatment:03EF3A1F8A224C0BFE15282281267291:8E99DDD28A224C15FCF92BC681BA73B4" box="[246,507,1956,1982]" country="Mexico" latitude="27.337057" longLatPrecision="1" longitude="-112.26322" name="Santa Rosalia Marina" pageId="46" pageNumber="47" stateProvince="Baja California Sur">Santa Rosalia Marina</location>
;
<geoCoordinate id="EE72EDCE8A224C15FE072BC082F773B4" box="[520,694,1954,1982]" degrees="27" direction="north" minutes="20" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="13.4" value="27.337057">27°20′13.4″ N</geoCoordinate>
,
<geoCoordinate id="EE72EDCE8A224C15FECB2BC083C473B4" box="[708,901,1954,1982]" degrees="112" direction="west" minutes="15" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="47.6" value="-112.26322">112°15′47.6″ W</geoCoordinate>
;
<date id="FFF8ADC98A224C15FF9C2BC6847973B4" box="[915,1080,1956,1982]" pageId="46" pageNumber="47" value="2013-11-12">
<collectingDate id="EFBC54218A224C15FF9C2BC6847973B4" box="[915,1080,1956,1982]" pageId="46" pageNumber="47" value="2013-11-12">12 Nov. 2013</collectingDate>
</date>
,
<collectorName id="26B3EEDF8A224C15F8482BC684A173B4" box="[1095,1248,1956,1982]" pageId="46" pageNumber="47">J.A. de León</collectorName>
leg.;
<specimenCode id="DBE023728A224C15F9282BC680B973E8" pageId="46" pageNumber="47">UANL 8150</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B2E81548A224C15FD1E2BA583CF73E8" ID-GBIF-Occurrence="3411820306" box="[273,910,1991,2018]" collectingDate="2017-06-24" collectionCode="UANL" collectorName="J. A. de Leon" pageId="46" pageNumber="47" specimenCode="UANL 8151" specimenCount="2">
<specimenCount id="9D4040808A224C15FD1E2BA5812B73E8" box="[273,362,1991,2018]" pageId="46" pageNumber="47">2 specs</specimenCount>
;
<date id="FFF8ADC98A224C15FD792BA5825173EB" box="[374,528,1991,2018]" pageId="46" pageNumber="47" value="2017-06-24">
<collectingDate id="EFBC54218A224C15FD792BA5825173EB" box="[374,528,1991,2018]" pageId="46" pageNumber="47" value="2017-06-24">24 Jun. 2017</collectingDate>
</date>
;
<collectorName id="26B3EEDF8A224C15FE132BA582F073EB" box="[540,689,1991,2017]" pageId="46" pageNumber="47">J.A. de León</collectorName>
leg.;
<specimenCode id="DBE023728A224C15FEFB2BA583C873E8" box="[756,905,1991,2018]" pageId="46" pageNumber="47">UANL 8151</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35CD8828A234C14FCB22D6B84477265" pageId="47" pageNumber="48" type="description">
<paragraph id="8BF98B098A234C14FCB22D6B828B7529" blockId="47.[189,714,265,291]" box="[189,714,265,291]" pageId="47" pageNumber="48">
<emphasis id="B932571B8A234C14FCB22D6B810F7529" bold="true" box="[189,334,265,291]" pageId="47" pageNumber="48">Description</emphasis>
(abridged from several sources)
</paragraph>
<paragraph id="8BF98B098A234C14FCB22D5A83C5757F" blockId="47.[187,1399,311,373]" pageId="47" pageNumber="48">BODY. Dark brownish pigmentation on dorsal surfaces of prostomium and tentacular belt, often continued as two transverse bars per segment along anterior chaetigers.</paragraph>
<paragraph id="8BF98B098A234C14FCB22DFE850275D3" blockId="47.[189,1399,412,474]" pageId="47" pageNumber="48">PROSTOMIUM. Pyriform, longer than wide; eyes round, of similar size, in a rectangular arrangement. Antennae slightly surpassing palp tips. Palpophores globose, as long as wide, palpostyles rounded.</paragraph>
<paragraph id="8BF98B098A234C14FCB22E6384DC7635" blockId="47.[189,1398,513,575]" pageId="47" pageNumber="48">TENTACULAR BELT. 2× as long as first chaetiger, anterior margin middorsally projected anteriorly, not covering posterior eyes. Tentacular cirri smooth, longest one reaching chaetiger 5–9.</paragraph>
<paragraph id="8BF98B098A234C14FCB22E04810F76CC" blockId="47.[189,1399,613,710]" pageId="47" pageNumber="48">
PHARYNX. Jaws brownish with 9–10 teeth. Paragnaths formula: I:
<quantity id="4CBE26EC8A234C14FFCF2E0483AC7675" box="[960,1005,613,640]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="47" pageNumber="48" unit="in" value="2.0">2 in</quantity>
tandem; II:
<quantity id="4CBE26EC8A234C14F8722E0484A47675" box="[1149,1253,613,640]" metricMagnitude="-1" metricUnit="m" metricValue="8.509" metricValueMax="8.636" metricValueMin="8.382" pageId="47" pageNumber="48" unit="in" value="33.5" valueMax="34.0" valueMin="33.0">33–34 in</quantity>
an arch; III:
<quantity id="4CBE26EC8A234C14FCB22EEA80BB76A9" box="[189,250,648,675]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="47" pageNumber="48" unit="in" value="36.0">36 in</quantity>
an oval; IV: 33, 35 of different sizes; V: 0; VI: 8, 11 in a cluster; VII–VIII: single row with 4–5 small cones.
</paragraph>
<paragraph id="8BF98B098A234C14FCB22E8F84777744" blockId="47.[189,1400,749,846]" pageId="47" pageNumber="48">PATTERN OF PARAPODIAL LIGULES. Anterior chaetigers with ligules blunt, slightly longer than wide, of similar length and width, becoming tapered, blunt, slightly longer than wide in median and posterior chaetigers; ventral ligule progressively smaller, thinner, 2× as long as wide.</paragraph>
<paragraph id="8BF98B098A234C14FCB22F17847D779A" blockId="47.[189,1084,885,912]" box="[189,1084,885,912]" pageId="47" pageNumber="48">CHAETIGERS 1–2. With neuraciculae only. Ligules without glandular masses.</paragraph>
<paragraph id="8BF98B098A234C14FCB22FD5813070CD" blockId="47.[188,1401,950,1223]" pageId="47" pageNumber="48">MEDIAN PARAPODIA. With dorsal cirrus longer than ventral one, both thin; dorsal cirrus almost 2 × as long as dorsal ligule; dorsal ligule slightly longer and wider than median ligule, both with dark inner compact glandular masses. Neuracicular ligule blunt, subconical, about as long as wide. Ventral ligule digitate, 2× as long as wide. Ventral cirrus not reaching tip of ventral ligule. Notochaetae: homogomph spinigers and falcigers from chaetiger 39, blades 3 × as long as wide, margin denticulate, distal tooth incurved. Neurochaetae: homogomph spinigers and heterogomph falcigers in supracicular and subacicular fascicles; falcigers with blades slightly more than 2× as long as wide, with margin denticulate, distal tooth incurved.</paragraph>
<paragraph id="8BF98B098A234C14FCB2288C85367102" blockId="47.[189,1399,1262,1289]" box="[189,1399,1262,1289]" pageId="47" pageNumber="48">POSTERIOR REGION.Tapered. Pygidium with anus terminal with two anal cirri, as long as last 11 chaetigers.</paragraph>
<paragraph id="8BF98B098A234C14FCB2294D81727143" blockId="47.[189,307,1327,1353]" box="[189,307,1327,1353]" pageId="47" pageNumber="48">
<heading id="D0B13C658A234C14FCB2294D81727143" bold="true" box="[189,307,1327,1353]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B932571B8A234C14FCB2294D81727143" bold="true" box="[189,307,1327,1353]" pageId="47" pageNumber="48">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BF98B098A234C14FCB2293C84477265" blockId="47.[188,1401,1374,1648]" pageId="47" pageNumber="48">
Gulf of
<collectingRegion id="498245EB8A234C14FD12293C81D57172" box="[285,404,1374,1400]" country="United States of America" name="California" pageId="47" pageNumber="48">California</collectingRegion>
specimens (n = 14) showed some variation in their paragnath formula which is not size-dependent. Pharyngeal area I: 0–3 cones (50% of specimens with 2; 25% with only one; 15% without cones; 10% with 3 teeth and when the third cone is present, it is minute). Area II: right side 7–23 cones, left side 10–26 cones. Area III: oval patch with 13–30 cones in three lines, proximal line with larger cones (20% of specimens with 21 cones). Area IV: right side 12–30 cones, left side 12–34 cones. Area V: none. Area VI: right side 3–10 cones, left side 3–9 cones (25% of specimens with 5–5 cones, respectively). Area VII–VIII:
<quantity id="4CBE26EC8A234C14FE2E2A50822C7241" box="[545,621,1585,1612]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.778" metricValueMin="0.508" pageId="47" pageNumber="48" unit="in" value="4.5" valueMax="7.0" valueMin="2.0">2–7 in</quantity>
a row (60% of specimens with 5 cones). Longest tentacular cirri reach chaetiger 2–6 (in almost 25% of specimens reaching chaetiger 4).
</paragraph>
</subSubSection>
<subSubSection id="C35CD8828A234C0BFCB22AF481E371EE" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8BF98B098A234C14FCB22AF4816E72BA" blockId="47.[189,303,1686,1712]" box="[189,303,1686,1712]" pageId="47" pageNumber="48">
<heading id="D0B13C658A234C14FCB22AF4816E72BA" bold="true" box="[189,303,1686,1712]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B932571B8A234C14FCB22AF4816E72BA" bold="true" box="[189,303,1686,1712]" pageId="47" pageNumber="48">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF98B098A234C14FCB22AA785377308" blockId="47.[189,1399,1733,1794]" pageId="47" pageNumber="48">
The specific epithet must be modified because the original one was masculine (‛
<taxonomicName id="4C46F08A8A234C14F8542AA7848C72D5" authorityName="Treadwell" authorityYear="1937" box="[1115,1229,1733,1759]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="ambiguus">
<emphasis id="B932571B8A234C14F8542AA7848C72D5" box="[1115,1229,1733,1759]" italics="true" pageId="47" pageNumber="48">ambiguus</emphasis>
</taxonomicName>
’, ‛uncertain’), but the gender of
<taxonomicName id="4C46F08A8A234C14FD8B2A8B81917308" authorityName="Linnaeus" authorityYear="1758" box="[388,464,1769,1794]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="B932571B8A234C14FD8B2A8B81917308" box="[388,464,1769,1794]" italics="true" pageId="47" pageNumber="48">Nereis</emphasis>
</taxonomicName>
is feminine. Therefore, the epithet must be used in the feminine form (
<taxonomicName id="4C46F08A8A234C14F8F02A8A85277308" authorityName="Treadwell" authorityYear="1937" box="[1279,1382,1768,1794]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A234C14F8F02A8A85277308" box="[1279,1382,1768,1794]" italics="true" pageId="47" pageNumber="48">ambigua</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF98B098A234C14FCB22B4B853773DB" blockId="47.[188,1399,1833,2001]" pageId="47" pageNumber="48">
There are some problems regarding the
<typeStatus id="54FD35AB8A234C14FE852B4882FC734E" box="[650,701,1834,1860]" pageId="47" pageNumber="48">type</typeStatus>
material of
<taxonomicName id="4C46F08A8A234C14FF482B48838E7349" authorityName="Treadwell" authorityYear="1937" box="[839,975,1833,1859]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A234C14FF482B48838E7349" box="[839,975,1833,1859]" italics="true" pageId="47" pageNumber="48">N. ambigua</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFD7F6F88A234C14FFD32B4B84A37349" author="Treadwell A. L." box="[988,1250,1833,1859]" pageId="47" pageNumber="48" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell (1937: 149)</bibRefCitation>
selected and described as “
<typeStatus id="54FD35AB8A234C14FD6F2B2F81D6736D" box="[352,407,1869,1895]" pageId="47" pageNumber="48">type</typeStatus>
” a juvenile collected in the Revillagigedo Islands. It was
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long,
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wide, without the posterior region, and he recorded another, smaller specimen. Surprisingly, he indicated “the
<typeStatus id="54FD35AB8A234C14FCB32BF180AE73A7" box="[188,239,1939,1965]" pageId="47" pageNumber="48">type</typeStatus>
retained only one anal cirrus” (
<bibRefCitation id="EFD7F6F88A234C14FE532BF1831273A4" author="Treadwell A. L." box="[604,851,1939,1966]" pageId="47" pageNumber="48" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell 1937: 150</bibRefCitation>
), and this might correspond to the heteronereis he referred to in the last line of his description.
<bibRefCitation id="EFD7F6F88A234C14FEEC2BD483EF73DA" author="Treadwell A. L." box="[739,942,1974,2000]" pageId="47" pageNumber="48" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell (1937)</bibRefCitation>
described the largest specimen,
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</paragraph>
<paragraph id="8BF98B098A3C4C0BFCB22D6B8105754C" blockId="48.[189,1398,265,326]" pageId="48" pageNumber="49">
long and
<quantity id="4CBE26EC8A3C4C0BFD242D6B812E7529" box="[299,367,265,291]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="1.0">1 mm</quantity>
wide, and its pharynx was not exposed, preventing him to clearly count the numbers of paragnaths.
</paragraph>
<paragraph id="8BF98B098A3C4C0BFCB22D0C829D775B" blockId="48.[187,1401,366,850]" pageId="48" pageNumber="49">
<bibRefCitation id="EFD7F6F88A3C4C0BFCB22D0C81C07582" author="Hartman O." box="[189,385,366,392]" pageId="48" pageNumber="49" pagination="173 - 287" refId="ref43181" refString="Hartman O. 1940. Polychaetous annelids, 2. Chrysopetalidae to Goniadidae. Allan Hancock Pacific Expeditions 7: 173 - 287. Available from https: // www. biodiversitylibrary. org / page / 27822405 # page / 191 / mode / 1 up [accessed 16 Sep. 2021]." type="journal article" year="1940">Hartman (1940)</bibRefCitation>
synonymized this species with
<taxonomicName id="4C46F08A8A3C4C0BFF122D0C83C2758D" box="[797,899,366,391]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="riisei">
<emphasis id="B932571B8A3C4C0BFF122D0C83C2758D" box="[797,899,366,391]" italics="true" pageId="48" pageNumber="49">N. riisei</emphasis>
</taxonomicName>
on the basis of parapodial ligules and homogomph falciger shape only, but no further comments were made. Later,
<bibRefCitation id="EFD7F6F88A3C4C0BF8542DF3855975A6" author="Hartman O." box="[1115,1304,401,428]" pageId="48" pageNumber="49" pagination="239 - 310" refId="ref43398" refString="Hartman O. 1956. Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology. Bulletin of the American Museum of Natural History 109: 239 - 310." type="journal article" year="1956">Hartman (1956)</bibRefCitation>
studied one of the
<typeStatus id="54FD35AB8A3C4C0BFD362DD7812D75C5" box="[313,364,437,463]" pageId="48" pageNumber="49">type</typeStatus>
specimens, but she failed to indicate which one it was or whether there was only a single specimen remaining in the jar, the largest one being destroyed after dissection. She dissected the pharynx and found a paragnath distribution different from the one originally indicated by
<bibRefCitation id="EFD7F6F88A3C4C0BF8912D988530761F" author="Treadwell A. L." box="[1182,1393,506,533]" pageId="48" pageNumber="49" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell (1937)</bibRefCitation>
. Treadwell indicated that areas I and II were smooth, whereas Hartman found 3 paragnaths in I and a patch of larger paragnaths in II; furthermore, larger differences relate to areas VI and VII–VIII, because Treadwell indicated 3–4 cones in VI, 7–8 cones in VII and a circular patch with about 20 paragnaths in VIII, but Hartman noted 12 cones in VI and a single continuous band with about 10 alternating larger and smaller cones in VII–VIII. On the other hand,
<bibRefCitation id="EFD7F6F88A3C4C0BFF252EC9846676CF" author="Hartman O." box="[810,1063,683,709]" pageId="48" pageNumber="49" pagination="173 - 287" refId="ref43181" refString="Hartman O. 1940. Polychaetous annelids, 2. Chrysopetalidae to Goniadidae. Allan Hancock Pacific Expeditions 7: 173 - 287. Available from https: // www. biodiversitylibrary. org / page / 27822405 # page / 191 / mode / 1 up [accessed 16 Sep. 2021]." type="journal article" year="1940">Hartman (1940: 222)</bibRefCitation>
found a different pattern in her Eastern Pacific specimens, with 4–5 cones in VI and a single band with
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VII–VIII. A similar pattern, matching Hartman’s account, was recorded by
<bibRefCitation id="EFD7F6F88A3C4C0BFF522E9383B27701" author="Dean H. K." box="[861,1011,753,779]" pageId="48" pageNumber="49" pagination="37 - 67" refId="ref41350" refString="Dean H. K. 2001. Some Nereididae (Annelida: Polychaeta) from the Pacific coast of Costa Rica. Revista de Biologia Tropical 49 (Suppl. 2): 37 - 67." type="journal article" year="2001">Dean (2001)</bibRefCitation>
for his
<collectingRegion id="498245EB8A3C4C0BF8572E9384DA7701" box="[1112,1179,753,779]" country="New Zealand" name="West Coast" pageId="48" pageNumber="49">Costa</collectingRegion>
Rican specimens.
<bibRefCitation id="EFD7F6F88A3C4C0BFCB22F7781387725" author="Hartman O." box="[189,377,788,815]" pageId="48" pageNumber="49" pagination="239 - 310" refId="ref43398" refString="Hartman O. 1956. Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology. Bulletin of the American Museum of Natural History 109: 239 - 310." type="journal article" year="1956">Hartman (1956)</bibRefCitation>
also detailed 9 teeth on the jaws, dark aciculae and homogomph falcigers (presumably notopodial ones) appearing from chaetiger 26.
</paragraph>
<paragraph id="8BF98B098A3C4C0BFCB22F1B83E8706C" blockId="48.[188,1399,889,1127]" pageId="48" pageNumber="49">
Because of this series of differences, which were more or less corroborated in the Gulf of
<collectingRegion id="498245EB8A3C4C0BF90F2F1B85367799" box="[1280,1399,889,915]" country="United States of America" name="California" pageId="48" pageNumber="49">California</collectingRegion>
material, and because
<bibRefCitation id="EFD7F6F88A3C4C0BFDCE2FFE82CC77BC" author="Treadwell A. L." box="[449,653,924,950]" pageId="48" pageNumber="49" pagination="139 - 160" refId="ref46120" refString="Treadwell A. L. 1937. The Templeton Crocker Expedition. VIII. Polychaetous annelids from the west coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society 22 (2): 139 - 160. https: // doi. org / 10.5962 / p. 184684" type="journal article" year="1937">Treadwell (1937)</bibRefCitation>
indicated he had
<specimenCount id="9D4040808A3C4C0BFF542FFE844D77BC" box="[859,1036,924,950]" pageId="48" pageNumber="49" type="generic">two specimens</specimenCount>
, the chances are that those
<specimenCount id="9D4040808A3C4C0BF9452FFE817677D3" pageId="48" pageNumber="49" type="generic">two specimens</specimenCount>
belong to different species. An alternative is that Treadwell confused the pharyngeal areas in the non-everted pharynx, as he indicated, and that would explain the specific epithet. The specimens that
<bibRefCitation id="EFD7F6F88A3C4C0BFCFF286481EA702A" author="Hartman O." box="[240,427,1030,1056]" pageId="48" pageNumber="49" pagination="173 - 287" refId="ref43181" refString="Hartman O. 1940. Polychaetous annelids, 2. Chrysopetalidae to Goniadidae. Allan Hancock Pacific Expeditions 7: 173 - 287. Available from https: // www. biodiversitylibrary. org / page / 27822405 # page / 191 / mode / 1 up [accessed 16 Sep. 2021]." type="journal article" year="1940">Hartman (1940)</bibRefCitation>
and
<bibRefCitation id="EFD7F6F88A3C4C0BFDEC28648237702A" author="Dean H. K." box="[483,630,1030,1056]" pageId="48" pageNumber="49" pagination="37 - 67" refId="ref41350" refString="Dean H. K. 2001. Some Nereididae (Annelida: Polychaeta) from the Pacific coast of Costa Rica. Revista de Biologia Tropical 49 (Suppl. 2): 37 - 67." type="journal article" year="2001">Dean (2001)</bibRefCitation>
studied were more similar to
<taxonomicName id="4C46F08A8A3C4C0BFFDE2864846E7015" box="[977,1071,1030,1055]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="riisei">
<emphasis id="B932571B8A3C4C0BFFDE2864846E7015" box="[977,1071,1030,1055]" italics="true" pageId="48" pageNumber="49">N. riisei</emphasis>
</taxonomicName>
than the specimen
<bibRefCitation id="EFD7F6F88A3C4C0BF91F2864814B7049" author="Hartman O." pageId="48" pageNumber="49" pagination="239 - 310" refId="ref43398" refString="Hartman O. 1956. Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology. Bulletin of the American Museum of Natural History 109: 239 - 310." type="journal article" year="1956">Hartman (1956)</bibRefCitation>
studied. Regretfully, subsequent collecting trips to the Revillagigedo Islands (
<bibRefCitation id="EFD7F6F88A3C4C0BF8CF284B85307049" author="Hartman O." box="[1216,1393,1065,1091]" pageId="48" pageNumber="49" pagination="1 - 22" refId="ref43122" refString="Hartman O. 1939. The polychaetous annelids collected on the Presidential Cruise of 1938. Smithsonian Miscellaneous Collections 98 (13): 1 - 22. Available from https: // www. biodiversitylibrary. org / item / 35649 # page / 371 / mode / 1 up [accessed 16 Sep. 2021]." type="journal article" year="1939">Hartman 1939</bibRefCitation>
;
<bibRefCitation id="EFD7F6F88A3C4C0BFCB2282E8101706D" author="Rioja E." box="[189,320,1100,1127]" pageId="48" pageNumber="49" pagination="243 - 259" refId="ref45218" refString="Rioja E. 1960. Estudios anelidologicos, 23. Contribucion al conocimiento de los anelidos poliquetos de las Islas de Revillagigedo. Anales del Instituto de Biologia, Mexico 30: 243 - 259." type="journal article" year="1960">
<collectingRegion id="498245EB8A3C4C0BFCB2282E80BC706C" box="[189,253,1100,1126]" country="Spain" name="La Rioja" pageId="48" pageNumber="49">Rioja</collectingRegion>
1960
</bibRefCitation>
) did not provide topotype specimens of
<taxonomicName id="4C46F08A8A3C4C0BFF16282F83E2706C" authorityName="Treadwell" authorityYear="1937" box="[793,931,1100,1126]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A3C4C0BFF16282F83E2706C" box="[793,931,1100,1126]" italics="true" pageId="48" pageNumber="49">N. ambigua</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF98B098A3C4C0BFCB228EF81E371EE" blockId="48.[188,1399,1165,1508]" pageId="48" pageNumber="49">
Consequently, the key below indicates that
<taxonomicName id="4C46F08A8A3C4C0BFEE028EC83C070AD" authorityName="Treadwell" authorityYear="1937" box="[751,897,1165,1191]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A3C4C0BFEE028EC83C070AD" box="[751,897,1165,1191]" italics="true" pageId="48" pageNumber="49">N. ambigua</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C46F08A8A3C4C0BF81928EC843C70AD" box="[1046,1149,1166,1191]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="riisei">
<emphasis id="B932571B8A3C4C0BF81928EC843C70AD" box="[1046,1149,1166,1191]" italics="true" pageId="48" pageNumber="49">N. riisei</emphasis>
</taxonomicName>
because both have posterior eyes completely exposed and a single row of paragnaths in areas VII–VIII, after the careful characterization by
<bibRefCitation id="EFD7F6F88A3C4C0BFDA528B6820270E4" author="Dean H. K." box="[426,579,1236,1262]" pageId="48" pageNumber="49" pagination="37 - 67" refId="ref41350" refString="Dean H. K. 2001. Some Nereididae (Annelida: Polychaeta) from the Pacific coast of Costa Rica. Revista de Biologia Tropical 49 (Suppl. 2): 37 - 67." type="journal article" year="2001">Dean (2001)</bibRefCitation>
. Based on the original description,
<taxonomicName id="4C46F08A8A3C4C0BFFE228B6843970E4" authorityName="Treadwell" authorityYear="1937" box="[1005,1144,1236,1262]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A3C4C0BFFE228B6843970E4" box="[1005,1144,1236,1262]" italics="true" pageId="48" pageNumber="49">N. ambigua</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C46F08A8A3C4C0BF91928B6853770E7" box="[1302,1398,1236,1261]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="riisei">
<emphasis id="B932571B8A3C4C0BF91928B6853770E7" box="[1302,1398,1236,1261]" italics="true" pageId="48" pageNumber="49">N. riisei</emphasis>
</taxonomicName>
mainly by having larger eyes, tentacular cirri reaching chaetiger 9 and dorsal cirri in posterior chaetigers shorter than dorsal ligules. These two species further differ because in
<taxonomicName id="4C46F08A8A3C4C0BFFF1297984C9713E" authorityName="Treadwell" authorityYear="1937" box="[1022,1160,1306,1332]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A3C4C0BFFF1297984C9713E" box="[1022,1160,1306,1332]" italics="true" pageId="48" pageNumber="49">N. ambigua</emphasis>
</taxonomicName>
the ventral ligule in median and posterior parapodia reach the tips of the neuracicular ligules, and in fresh specimens the tentacular belt is pale. On the contrary, in
<taxonomicName id="4C46F08A8A3C4C0BFEA8290383447170" box="[679,773,1377,1402]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="riisei">
<emphasis id="B932571B8A3C4C0BFEA8290383447170" box="[679,773,1377,1402]" italics="true" pageId="48" pageNumber="49">N. riisei</emphasis>
</taxonomicName>
the ventral ligules in median and posterior parapodia surpass the neuracicular ligules, and fresh specimens have a blackish tentacular belt. Nevertheless, these differences must be taken as preliminary pending the finding of topotype specimens and confirming their diagnostic features.
</paragraph>
</subSubSection>
<subSubSection id="C35CD8828A3C4C0BFCB22A6981267291" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BF98B098A3C4C0BFCB22A698117722F" blockId="48.[189,342,1547,1573]" box="[189,342,1547,1573]" pageId="48" pageNumber="49">
<heading id="D0B13C658A3C4C0BFCB22A698117722F" bold="true" box="[189,342,1547,1573]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B932571B8A3C4C0BFCB22A698117722F" bold="true" box="[189,342,1547,1573]" pageId="48" pageNumber="49">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF98B098A3C4C0BFCB22A5881267291" blockId="48.[189,1398,1594,1691]" pageId="48" pageNumber="49">
Originally described from the Revillagigedo Islands, the potential distribution of
<taxonomicName id="4C46F08A8A3C4C0BF87E2A59856A725E" authorityName="Treadwell" authorityYear="1937" box="[1137,1323,1594,1620]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="48" pageNumber="49" phylum="Annelida" rank="species" species="ambigua">
<emphasis id="B932571B8A3C4C0BF87E2A59856A725E" box="[1137,1323,1594,1620]" italics="true" pageId="48" pageNumber="49">Nereis ambigua</emphasis>
</taxonomicName>
might extend along the Eastern Pacific, from the Gulf of California to the
<collectingRegion id="498245EB8A3C4C0BFFD22A3F84F5727D" box="[989,1204,1629,1655]" country="Ecuador" name="Galapagos" pageId="48" pageNumber="49">Galápagos Islands</collectingRegion>
and
<collectingCountry id="F351CB998A3C4C0BF8E02A3F8512727D" box="[1263,1363,1629,1655]" name="Ecuador" pageId="48" pageNumber="49">Ecuador</collectingCountry>
, in shallow water.
</paragraph>
</subSubSection>
</treatment>
</document>
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