<documentid="BBA949D6A08DE60560042F3871C194FC"ID-CLB-Dataset="48704"ID-DOI="10.5281/zenodo.184552"ID-GBIF-Dataset="8eb92ccd-80d7-4381-bee3-7616ef59d4b1"ID-ISSN="1175-5326"ID-Zenodo-Dep="184552"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"checkinTime="1459987789263"checkinUser="plazi"docAuthor="Mclaughlin, Patsy A. & Lemaitre, Rafael"docDate="2008"docId="03D487909930FFFDFF2FF938FB5BFD29"docLanguage="en"docName="zt01911p068.pdf"docOrigin="Zootaxa 1911"docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article"docStyleId="6581673A57F01A3145754A1E615EDFF0"docStyleName="Zootaxa.2007-2008.journal_article"docStyleVersion="4"docTitle="Trizocheles Forest 1987"docType="treatment"docVersion="11"lastPageNumber="56"masterDocId="FFEDFFE89931FFF9FFB8FFFFFF83FFBB"masterDocTitle="Larvae of two species of Trizocheles (Decapoda: Anomura: Paguroidea: Pylochelidae: Trizochelinae), description of the adult of one, and preliminary implications of development on pylochelid phylogeny"masterLastPageNumber="68"masterPageNumber="52"pageNumber="53"updateTime="1732718119159"updateUser="ExternalLinkService">
<mods:titleid="0C5B5B8E5BF5F088F87F7DDFD0E4B3D9">Larvae of two species of Trizocheles (Decapoda: Anomura: Paguroidea: Pylochelidae: Trizochelinae), description of the adult of one, and preliminary implications of development on pylochelid phylogeny</mods:title>
<bibRefCitationid="EFEC4B779930FFF8FE5BF8F7FD1FF8A4"author="Henderson"box="[483,668,1800,1823]"pageId="1"pageNumber="67"refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger during the years 1873 - 76, (Zoology) volume 27, i - xi, 1 - 221."type="journal article"year="1888">Henderson, 1888</bibRefCitation>
<bibRefCitationid="EFEC4B779930FFF8FECCF8D7FDAEF884"author="Henderson"box="[372,557,1832,1855]"pageId="1"pageNumber="67"refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger during the years 1873 - 76, (Zoology) volume 27, i - xi, 1 - 221."type="journal article"year="1888">Henderson, 1888</bibRefCitation>
<dateid="FFC310469930FFF8FBA9F82AFB11F854"box="[1041,1170,2005,2031]"pageId="1"pageNumber="53"value="1999-06-05">5 Jun 1999</date>
(
<collectionCodeid="ED6CAE439930FFF8FB18F82AFB6CF854"box="[1184,1263,2005,2031]"httpUri="http://grbio.org/cool/nyq3-3g6s"name="National Institute of Water and Atmospheric Research"pageId="1"pageNumber="53">NIWA</collectionCode>
. a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b–i).
<figureCitationid="13462A039933FFFBFEF1FA8AFE0FFA34"box="[329,396,1397,1423]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="2"pageNumber="54">Fig. 1</figureCitation>
a). Globoid, lacking spines and setae, posterior margins rounded; rostrum approximately 0.2 of carapace length, moderately slender, frequently directed somewhat downward.
<figureCitationid="13462A039933FFFBFEF5FA12FE0CF9BC"box="[333,399,1517,1543]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="2"pageNumber="54">Fig. 1</figureCitation>
b). Biramous, with protopod elongate, unsegmented; endopod varying from nearly equal in length to approximately twice length of exopod, naked or with 1 terminal seta; exopod varying from naked to bearing 1 short terminal seta and 3 or 4 aesthetascs, 1 aesthetasc on inner margin.
<figureCitationid="13462A039933FFFBFE82F99AFEFCF9C4"box="[314,383,1637,1663]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="2"pageNumber="54">Fig. 1</figureCitation>
c). Biramous; protopod unsegmented, with small spine at endopodal junction and larger spine at scaphocerite junction; base of protopod sometimes enlarged, with internally directed lobe of future antennal gland orifice apparent; endopod varying from approximately half length of scaphocerite to nearly equal to tip of distolateral spine. Scaphocerite with 11 or 12 setae, often only partially evaginated, on inner margin, upper margin with or without small spine; distolateral spine subdistal in origin and very prominently bifid.
<figureCitationid="13462A039933FFFBFEF2F8AAFE13F8D4"box="[330,400,1877,1903]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="2"pageNumber="54">Fig. 1</figureCitation>
d). Varying from indistinguishably fused to epistome to subquadrate in structure with unsegmented, naked palp.
<figureCitationid="13462A039933FFFBFEF2F85AFE13F804"box="[330,400,1957,1983]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="2"pageNumber="54">Fig. 1</figureCitation>
e). Often indiscernible, occasionally with buds of endites and endopod developed, all lacking setae.
<figureCitationid="13462A039932FFFAFE8AFF67FEF5FF09"box="[306,374,152,178]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="3"pageNumber="55">Fig. 1</figureCitation>
f). Varying from bilobed bud to moderately well developed appendage with bilobed coxal and basial endites, and scaphognathite, endopod apparent or not; all lacking setae.
<figureCitationid="13462A039932FFFAFE2EFF17FE59FEB9"box="[406,474,232,258]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="3"pageNumber="55">
<figureCitationid="13462A039932FFFAFE01FEC7FD82FEE9"box="[441,513,312,338]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="3"pageNumber="55">Fig. 1</figureCitation>
h). Protopod unsegmented; exopod elongate, varying from unsegmented to incompletely 2-segmented; endopod varying from unsegmented and subequal in length to exopod to appreciably shorter than exopod and faintly 4-segmented, setae lacking.
<figureCitationid="13462A039932FFFAFE22FE4FFE5DFE71"box="[410,478,432,458]"captionStart="FIGURE 1"captionStartId="2.[151,254,1253,1277]"captionTargetBox="[272,1318,245,1221]"captionTargetId="figure@2.[264,1323,237,1227]"captionTargetPageId="2"captionText="FIGURE 1. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, carapace (lateral view); b, antennule; c, antenna; d, mandible; e, maxillule; f, maxilla; g, first maxilliped; h, second maxilliped; i, third maxilliped. Scales: 1 mm (a), and 0.25 mm (b – i)."httpUri="https://zenodo.org/record/184553/files/figure.png"pageId="3"pageNumber="55">
i). Protopod unsegmented; exopod elongate, varying from faintly to incompletely 2-segmented; endopod unsegmented, varying from rudimentary to moderately well developed bud.
<figureCitationid="13462A039935FFFDFEF3FF67FE11FF09"box="[331,402,152,178]"captionStart="FIGURE 2"captionStartId="3.[151,254,1924,1948]"captionTargetBox="[285,1307,549,1892]"captionTargetId="figure@3.[274,1315,540,1902]"captionTargetPageId="3"captionText="FIGURE 2. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, cheliped; b, second pereopod; c, third pereopod; d, fourth pereopod; e, fifth pereopod; f, pleon and telson. Scales: 0.25 mm (a – e), and 1 mm (f)."httpUri="https://zenodo.org/record/184554/files/figure.png"pageId="4"pageNumber="56">Fig. 2</figureCitation>
a). Chela with dactyl distinguishable, articulation with palm usually not apparent; carpus and merus delineated, unarmed; setae entirely lacking.
<emphasisid="B909EA949935FFFDFF7EFF17FE24FEBA"box="[198,423,232,257]"italics="true"pageId="4"pageNumber="56">Pereopods 2 and 3</emphasis>
(
<figureCitationid="13462A039935FFFDFE00FF17FD83FEB9"box="[440,512,232,258]"captionStart="FIGURE 2"captionStartId="3.[151,254,1924,1948]"captionTargetBox="[285,1307,549,1892]"captionTargetId="figure@3.[274,1315,540,1902]"captionTargetPageId="3"captionText="FIGURE 2. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, cheliped; b, second pereopod; c, third pereopod; d, fourth pereopod; e, fifth pereopod; f, pleon and telson. Scales: 0.25 mm (a – e), and 1 mm (f)."httpUri="https://zenodo.org/record/184554/files/figure.png"pageId="4"pageNumber="56">Fig. 2</figureCitation>
b, c). Dactyls, propodi, carpi and meri varying from partially to clearly delineated, no terminal claws apparent; setae entirely lacking.
<figureCitationid="13462A039935FFFDFED2FEC7FE3BFEE9"box="[362,440,312,338]"captionStart="FIGURE 2"captionStartId="3.[151,254,1924,1948]"captionTargetBox="[285,1307,549,1892]"captionTargetId="figure@3.[274,1315,540,1902]"captionTargetPageId="3"captionText="FIGURE 2. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, cheliped; b, second pereopod; c, third pereopod; d, fourth pereopod; e, fifth pereopod; f, pleon and telson. Scales: 0.25 mm (a – e), and 1 mm (f)."httpUri="https://zenodo.org/record/184554/files/figure.png"pageId="4"pageNumber="56">Fig. 2</figureCitation>
<figureCitationid="13462A039935FFFDFEE3FE77FE20FE19"box="[347,419,392,418]"captionStart="FIGURE 2"captionStartId="3.[151,254,1924,1948]"captionTargetBox="[285,1307,549,1892]"captionTargetId="figure@3.[274,1315,540,1902]"captionTargetPageId="3"captionText="FIGURE 2. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, cheliped; b, second pereopod; c, third pereopod; d, fourth pereopod; e, fifth pereopod; f, pleon and telson. Scales: 0.25 mm (a – e), and 1 mm (f)."httpUri="https://zenodo.org/record/184554/files/figure.png"pageId="4"pageNumber="56">Fig. 2</figureCitation>
e). Dactyl partially to clearly delineated, simple, terminal; remaining segments partially to clearly delineated; setae entirely lacking.
<figureCitationid="13462A039935FFFDFEA4FE27FEE0FE49"box="[284,355,472,498]"captionStart="FIGURE 2"captionStartId="3.[151,254,1924,1948]"captionTargetBox="[285,1307,549,1892]"captionTargetId="figure@3.[274,1315,540,1902]"captionTargetPageId="3"captionText="FIGURE 2. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, cheliped; b, second pereopod; c, third pereopod; d, fourth pereopod; e, fifth pereopod; f, pleon and telson. Scales: 0.25 mm (a – e), and 1 mm (f)."httpUri="https://zenodo.org/record/184554/files/figure.png"pageId="4"pageNumber="56">Fig. 2</figureCitation>
f). Six-segmented, with telson clearly separated; pleomeres 1–4 and 6 unarmed, pleomere 5 with pair of moderately prominent posterolateral spines; uniramous or weakly biramous pleopod buds present on pleomeres 2–5; pleomere 6 with uni- or biramous uropod buds.
<figureCitationid="13462A039935FFFDFE9AFDAFFEEBFDD1"box="[290,360,592,618]"captionStart="FIGURE 2"captionStartId="3.[151,254,1924,1948]"captionTargetBox="[285,1307,549,1892]"captionTargetId="figure@3.[274,1315,540,1902]"captionTargetPageId="3"captionText="FIGURE 2. Metazoea of Trizocheles spinosus spinosus (Henderson, 1888). a, cheliped; b, second pereopod; c, third pereopod; d, fourth pereopod; e, fifth pereopod; f, pleon and telson. Scales: 0.25 mm (a – e), and 1 mm (f)."httpUri="https://zenodo.org/record/184554/files/figure.png"pageId="4"pageNumber="56">Fig. 2</figureCitation>
f, 4h). Short and moderately stout; terminal margin with broad V-shaped median cleft and 8 + 8 processes, all articulated, 3–8 each with long, fine marginal setae, fourth process longest.
