186 lines
19 KiB
XML
186 lines
19 KiB
XML
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<document id="6804E9AEA289EB7C3EFEE2A6DA4CC54E" ENCODING="UTF8" ID-GBIF-Dataset="5c456f28-83a8-4a50-9dee-80551a76f456" ModsDocAuthor="Bayartogtokh, B. & Schatz, H." ModsDocDate="2008" ModsDocID="BAYARTOGTOKH2008" ModsDocOrigin="http://www.mapress.com/zootaxa/2008/2/zt01948p035.pdf" ModsDocTitle="Trichoribates and Jugatala (Acari: Oribatida: Ceratozetidae) from the Central and Southern Alps, with notes on their distribution" checkinTime="1317117753293" checkinUser="thomas" docAuthor="Bayartogtokh, B. & Schatz, H." docDate="2008" docId="BD0AC06413B089D323F935084DFE6DA5" docLanguage="en" docName="Bayartogtokh_gg2.xml" docOrigin="Zootaxa 1948" docSource="http://www.mapress.com/zootaxa/2008/2/zt01948p035.pdf" docTitle="Trichoribates novus" docType="treatment" docVersion="10" lastPageNumber="16" masterDocId="BA7F56815C1778EA6535B52EA2F791A2" masterDocTitle="Trichoribates and Jugatala (Acari: Oribatida: Ceratozetidae) from the Central and Southern Alps, with notes on their distribution" masterLastPageNumber="35" masterPageNumber="1" pageNumber="13" updateTime="1701312712183" updateUser="plazi">
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<mods:title id="7545B02218DDD6AFFCEE9DB0E9F4EAAD">Trichoribates and Jugatala (Acari: Oribatida: Ceratozetidae) from the Central and Southern Alps, with notes on their distribution</mods:title>
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<mods:namePart id="EA141CA864BC04448A43D60F6B5164DD">Bayartogtokh, B.</mods:namePart>
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<mods:roleTerm id="0763E53394998859C9CF0077D727A184">Author</mods:roleTerm>
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<mods:namePart id="AFA834D8E0E2434C1F45FED710C194E6">Schatz, H.</mods:namePart>
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<mods:typeOfResource id="761CB0A6F153C824C760638D6633C366">text</mods:typeOfResource>
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<mods:title id="41890CB820100E56EC9C825EB20E5047">Zootaxa</mods:title>
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<mods:part id="29D83F03EE4E0279D4ED20A47CB45B1A">
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<mods:date id="85B3F187D422E6EDC5E71D54E79E9324">2008</mods:date>
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<mods:detail id="97E8A55F04F005834A65B9A9C70028EC" type="volume">
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<mods:number id="1CEAE3C19F529ACEDC9E88038BEAF086">1948</mods:number>
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<mods:start id="16208A899778A8D0956EBCF5C730DCE7">1</mods:start>
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<mods:end id="10972A171929176B2EDCE4CB34D2ED3E">35</mods:end>
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<mods:url id="64584EF30A76B5C1D87C934191E192E1">http://www.mapress.com/zootaxa/2008/2/zt01948p035.pdf</mods:url>
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<mods:classification id="7A4E3467CB1C3F3B78287626DF6BCD12">journal article</mods:classification>
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<mods:identifier id="F38EE4072D907BC49F0FB13CA55D67B9" type="Plazi-Custom">BAYARTOGTOKH2008</mods:identifier>
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</mods:mods>
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<treatment id="BD0AC06413B089D323F935084DFE6DA5" ID-DOI="http://doi.org/10.5281/zenodo.6229531" ID-GBIF-Taxon="100113797" ID-Zenodo-Dep="6229531" LSID="urn:lsid:plazi:treatment:BD0AC06413B089D323F935084DFE6DA5" httpUri="http://treatment.plazi.org/id/BD0AC06413B089D323F935084DFE6DA5" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
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<subSubSection id="B5EE9D3E08B66D1C10E2030300354916" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="A1E2E1E3A66CA8047D58B6CF9647CD64" pageId="12" pageNumber="13">
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<taxonomicName id="CB1A6177433F49B67E3E8A90CC646A5B" ID-CoL="8T838" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="novus">Trichoribates novus</taxonomicName>
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(Sellnick, 1928)
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</paragraph>
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</subSubSection>
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<subSubSection id="410C8CA3CE3DDF30759BAF4C4B3D41D7" pageId="12" pageNumber="13" type="description">
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<paragraph id="EBFADF0746B8E90076E06DEF7A8BB827" pageId="12" pageNumber="13">(Figs. 7, 8)</paragraph>
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</subSubSection>
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<subSubSection id="D846D414A3EF6A94977BD83E401B2E98" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="327C044E35BA4443C4ACCA9BD457B92A" pageId="12" pageNumber="13">
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<taxonomicName id="AD314BE5C73848E6F79C3FC4155AE061" class="Arachnida" family="Ceratozetidae" genus="Murcia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nova">Murcia nova</taxonomicName>
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Sellnick 1928, p. 11.
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</paragraph>
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<paragraph id="49961BBF0C814025E0BB375F0185C03D" pageId="12" pageNumber="13">
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<taxonomicName id="BBDE55E22461465C6360B07D429A1557" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="novus">Trichoribates novus</taxonomicName>
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: Willmann 1931, p. 170, fig. 275; van der Hammen 1952, p. 100; Schweizer 1956, p. 326, fig. 285; Shaldybina 1975, p. 292, fig. 708;
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<normalizedToken id="E45A563D37123B2C7AAF78F7F2570920" originalValue="Pérez-Iñigo" pageId="12" pageNumber="13">Perez-Inigo</normalizedToken>
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1993, p. 180; Pavlitshenko 1994, p. 31, fig. 22; Bayartogtokh et al. 2002, p. 13, figs. 34
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<normalizedToken id="166308DD7AFD2FAC60BDD2AC019BCE46" originalValue="—" pageId="12" pageNumber="13">-</normalizedToken>
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41; Weigmann 2006, p. 390, fig. 208a-d.
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</paragraph>
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<paragraph id="26062E7E2879053F11032FB2EB075E3C" pageId="12" pageNumber="13">
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<taxonomicName id="756F3089259AAE0F7F0A86579A2BE8E6" class="Arachnida" family="Ceratozetidae" genus="Murcia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nova">Murcia nova</taxonomicName>
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:
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<normalizedToken id="7B1B256D72F4C0281A60FE17B033C768" originalValue="Subías" pageId="12" pageNumber="13">Subias</normalizedToken>
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2004, p. 177; 2008, p. 332.
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</paragraph>
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</subSubSection>
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<subSubSection id="79D5E661A2724C4078D3F66B8F19031A" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="6369578BE12629173FEBA883D458C1A5" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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Diagnosis. Rostrum rounded, with a pair of minute lateral dens; rostral, lamellar and interlamellar setae densely barbed; lamella wide, with broad translamella; lamellar cusp with streng lateral dens, medial dens minute or indistinct; sensillus short, with minutely barbed, club-shaped head; tutorium broad, distally pointed;
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<pageBreakToken id="1F2643875D202484A6ACE4DF2C282737" pageId="14" pageNumber="15" start="start">11</pageBreakToken>
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pairs of notogastral setae medium long, barbed, seta dp present; porose areas oval; epimeral setae lc and adanal setae ad1, ad2 barbed, other ventral setae smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="AD236B55104BCB23289AADCDA63C7FE8" pageId="14" pageNumber="15" type="description">
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<paragraph id="DF40ABA8394DB7A64302AFD2F501C13F" pageId="14" pageNumber="15">
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Measurements. (n = 3) Body length 616
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<normalizedToken id="A0F358A00B689ACD5584D8E84066B4D6" originalValue="–" pageId="14" pageNumber="15">-</normalizedToken>
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669 (641)
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<normalizedToken id="06DB30DD5E666CFCFEEA8FB175235D2A" originalValue="µm" pageId="14" pageNumber="15">ym</normalizedToken>
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; length of notogaster 555
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<normalizedToken id="9F0B8901677E7A808F364B83398A6588" originalValue="–" pageId="14" pageNumber="15">-</normalizedToken>
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608 (583)
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<normalizedToken id="BB25908DC44C3B6867FC8E5AF0D6AE23" originalValue="µm" pageId="14" pageNumber="15">ym</normalizedToken>
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; width of notogaster 456
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<normalizedToken id="AD35D12A60BC2815640081AF7602454C" originalValue="–" pageId="14" pageNumber="15">-</normalizedToken>
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509 (484)
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<normalizedToken id="304C494F0108FB5C9F18B737771F5ECD" originalValue="µm" pageId="14" pageNumber="15">ym</normalizedToken>
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.
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</paragraph>
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<paragraph id="75E6C35078B01357374DCE7D9A381A6A" pageId="14" pageNumber="15">Integument. Body colour dark brown to almost black. With thick cerotegument, roughened by minute granules. Faintly microtuberculate on cuticle of prodorsum, notogaster, ventral plate, leg segments and subcapitular mentum.</paragraph>
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<paragraph id="911149CCD77CE1D8E0450069D4AE2A84" pageId="14" pageNumber="15">
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Prodorsum (Fig. 7
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<normalizedToken id="3853DA88157B9D7E61F1D1A39E20ABE3" originalValue="A–C" pageId="14" pageNumber="15">A-C</normalizedToken>
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, E, F). Rostrum rounded, with a pair of minute lateral dens, but without nose-like protuberance. Rostral and lamellar setae similar in length, densely barbed. Interlamellar seta about 1.5 x longer than setae ro and le, barbed, almost reaching tip of rostrum. Lamella wide, with longitudinal striations along outer margin; translamella long and wide; lamellar cusp with strong lateral dens, medial dens usually absent, but sometimes minute dens present (Fig. 7F). Sensillus of medium length, with minutely barbed club-shaped head (Fig. 7C). Bothridium small, with pointed bothridial scale svm as seen in dorsal aspect (Fig. 7A). Tutorium broad, with comparatively long, distally pointed cusp, its proximal part with longitudinal striations (Fig. 7 B, E).
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</paragraph>
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<paragraph id="175EB8A20712A46F5B7A6CE2E7D5BF9F" pageId="14" pageNumber="15">Notogaster (Figs. 7A, 8A, B). Oval, about 1.1 x longer than wide. Anterior margin of notogaster arched anteriad, lenticulus poorly developed. Pteromorph large, curved ventrally, anterior portion protruding markedly anteriad. Eleven pairs of notogastral setae, dp present, all setae long, barbed. Porose areas oval to round, Aa largest, A2 smallest. Lyrifissure im small, but well visible in dorsal or lateral view, ia, ih, ip and ips well visible in lateral or posterior view.</paragraph>
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<paragraph id="297FFF30A20141B8E37B661B01A5B833" pageId="14" pageNumber="15">
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Gnathosoma (Figs. 7D). Subcapitular mentum conspicuously wider than long, without noticeable microtubercles. Hypostomal setae a, h and m of medium length, smooth, attenuated. Structure and setation of palp and chelicera typical for family, as shown in the description of
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<taxonomicName id="D5D55544DC74C64ADE73E78ED573EAB0" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" genus.bestMatchDistance="7745" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="scilierensis">T. scilierensis</taxonomicName>
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<taxonomicNameLabel id="C64EE44BAF1B26A8248989C7221C7514" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="B262F1DD033A8E28CF8457709EF08AC9" pageId="14" pageNumber="15">Epimeral region (Fig. 7D). Most epimeral setae thin, smooth; setae lb and lc conspicuously longer and thicker than other setae, lc barbed, setal formula 3-1-3-3. Custodium reaching anterior margin of pedotectum II; discidium conspicuously projecting laterally.</paragraph>
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<paragraph id="9698BDE595CB98E84A18BE1430B51842" pageId="14" pageNumber="15">Ano-genital region (Figs. 7D, 8A). Anal aperture distinctly larger than genital one, anal and genital plates smooth. All ano-genital setae of medium length, thin, smooth, except adanal setae ad1 and ad2 barbed; setal formula same as in the previous species. Distance between bases of aggenital setae nearly equal to that between adanal setae ad3-ad3. Adanal lyrifissure iad short, situated at same level as anal setae an2, adjacent and parallel to anterolateral margin of anal aperture. Postanal porose area very long, narrow, far longer than distance between bases of adanal setae ad2 (Fig. 8A).</paragraph>
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<paragraph id="11CC8634189B255EF96A6CC49924CEBF" pageId="14" pageNumber="15">
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Legs. Most leg setae distinctly barbed, except (p) on tarsi
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<normalizedToken id="E13BD0761A6DEE8C99DA619949F1E938" originalValue="I–IV" pageId="14" pageNumber="15">I-IV</normalizedToken>
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. Setae l
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<normalizedToken id="B44D5B8B7F225FDEA81BA72544523B66" originalValue="”" pageId="14" pageNumber="15">"</normalizedToken>
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on tibiae and genua
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<normalizedToken id="D12C594390E753A36F8182BD2E0F5BF6" originalValue="I–IV" pageId="14" pageNumber="15">I-IV</normalizedToken>
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heavily barbed, much thicker than other setae. Formula of leg setation (including famulus): I (1-5-3-4-20); II (1-5-3-4-15); III (2-2-1-3-15); IV (1-2-2-3-12); formula of solenidia: I (1-2-2); II (1-1-2); III (1-1-0); IV (0-1-0). Illustration of legs is given by Bayartogtokh et al. (2002).
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</paragraph>
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</subSubSection>
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<subSubSection id="431BD181F360AD35969E14AD0514FADB" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8B8B4FBF01DFFFBA4318AB531A82B4C4" pageId="14" pageNumber="15">
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Material examined.
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<materialsCitation id="0F003BF7619AB2902DD44A143E50F4BD" ID-GBIF-Occurrence="923893748" collectingDate="1991-06-13" collectorName="H. Schatz" country="Italy" latitude="46.0206" location="Monte Bondone, near Viote" longitude="11.0633" stateProvince="Provincia di Trento">
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<collectingCountry id="B6B07FF0D0E9545C405125424460B4EE">Italy</collectingCountry>
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,
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<collectingRegion id="7FEB59F07C800DBA945413D84C292F7A">Provincia di Trento</collectingRegion>
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,
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<location id="F2447DCE63BB6496EED6382573D6D08C" LSID="urn:lsid:plazi:treatment:BD0AC06413B089D323F935084DFE6DA5:F2447DCE63BB6496EED6382573D6D08C" country="Italy" latitude="46.0206" longitude="11.0633" name="Monte Bondone, near Viote" stateProvince="Provincia di Trento">Monte Bondone, near Viote</location>
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(1650 m a.s.l.,
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<date id="B582E57194DDC1230F346058DFF3FA85" value="1991-06-13">
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<collectingDate id="C04FF1A64A60657A6BA1DFEC40986D44" value="1991-06-13">13 June 1991</collectingDate>
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</date>
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, two females, one male, leg.
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<collectorName id="D8B2BC673246C8DD33E2E2FD3D2061A1">H. Schatz</collectorName>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="5F48BD7D6868879CB3477C7E6F56AA05" pageId="14" pageNumber="15" type="discussion">
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<paragraph id="9E432288ECD258A16AF1F056D7386920" pageId="14" pageNumber="15">Remarks. The main character states of the examined specimens agree with those of the specimens studied by Shaldybina (1975), Pavlitshenko (1994), Bayartogtokh et al. (2002) and Weigmann (2006). However, none of those descriptions and drawings show the lateral dens on rostrum, but the specimens examined by us have these lateral dens clearly developed. Moreover, Pavlitshenko (1994) shows distinct medial dens of the lamellar cusps, but our specimens have no or very minute medial dens as illustrated by Shaldybina (1975) and Weigmann (2006).</paragraph>
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</subSubSection>
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<subSubSection id="152C7B6F9F1BB99B7D4FFA5DF02C2FD1" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="distribution">
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<paragraph id="7A6A9C3107C742D343ED133FD0C79324" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
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Distribution. This species is commonly distributed in the Holarctic region. In North America it is known only from Virginia, USA (Marshall et al. 1987), but is common in Europe (Schatz 1983; Karppinen et al. 1987, 1992; Bernini et al. 1995; Luxton 1996;
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<normalizedToken id="6B8DD51778A9D39E287FF445FE138B7D" originalValue="Niedbała" pageId="14" pageNumber="15">Niedbala</normalizedToken>
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& Olszanowski 1997;
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<normalizedToken id="055BE24F4D4D087D1A2FDAFEDB522DE3" originalValue="Subías" pageId="14" pageNumber="15">Subias</normalizedToken>
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&
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<normalizedToken id="DCA433EA5461ABD638344DD0DF54906F" originalValue="Gil-Martín" pageId="14" pageNumber="15">Gil-Martin</normalizedToken>
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1997). In
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<pageBreakToken id="10D86F3BAB2412588AAB626AF844EAE5" pageId="16" pageNumber="17" start="start">Asia</pageBreakToken>
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it was recorded in Turkey, Caucasus, Siberia, Far East of Russia, Japan, Kazakhstan and Mongolia (Golosova et al. 1983; Karppinen et al. 1986; Fujikawa et al. 1993; Rahimbaeva 1995; Ryabinin & Pankov 1997; Shtanchaeva 2001; Bayartogtokh 2007; Erman et al. 2007).
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</paragraph>
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</subSubSection>
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<subSubSection id="71644A9D93DC9402E2B1C71AC6FD4465" pageId="15" pageNumber="16" type="multiple">
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<caption id="EECA249D104F92CC3AC510099AE63F4D" pageId="15" pageNumber="16">
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<paragraph id="D08BF7854937BFF3306F095D574ED434" pageId="15" pageNumber="16">
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<pageBreakToken id="68CA8A510BCB90F070220F36AB4E0D13" pageId="15" pageNumber="16" start="start">FIGURE</pageBreakToken>
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7.
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<taxonomicName id="3EB5FFFEE63D15082C72AC99452E118A" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="novus">Trichoribates novus</taxonomicName>
|
|||
|
(Sellnick, 1928).
|
|||
|
<normalizedToken id="E8B92647AC429C8DF97A023DEA4F154D" originalValue="A—Dorsal" pageId="15" pageNumber="16">A-Dorsal</normalizedToken>
|
|||
|
view;
|
|||
|
<normalizedToken id="0F6E24C46C9FE2B6A59AB6349C9BF9B3" originalValue="B—Lateral" pageId="15" pageNumber="16">B-Lateral</normalizedToken>
|
|||
|
view of proterosoma;
|
|||
|
<normalizedToken id="52526D6520B8AB23E8696D833671B36C" originalValue="C—Sensillus" pageId="15" pageNumber="16">C-Sensillus</normalizedToken>
|
|||
|
and bothridium (lateral view);
|
|||
|
<normalizedToken id="07420F5D9F23040400F51099261FC557" originalValue="D—Ventral" pageId="15" pageNumber="16">D-Ventral</normalizedToken>
|
|||
|
view;
|
|||
|
<normalizedToken id="0DB9CD689A419DE49C435BA631DB6181" originalValue="E—Tutorium" pageId="15" pageNumber="16">E-Tutorium</normalizedToken>
|
|||
|
and rostral seta (right, lateral view);
|
|||
|
<normalizedToken id="2ED29879B3D7EA149D127BDF6D400140" originalValue="F—Lamellae" pageId="15" pageNumber="16">F-Lamellae</normalizedToken>
|
|||
|
with lamellar setae. Scale bar is same for A and D; for other figures, respectively.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<caption id="C30F4FC9EEB1F0BF9DE5BA9EC94E857D" pageId="15" pageNumber="16">
|
|||
|
<paragraph id="96E4E75107420FA046507769E7F23017" pageId="15" pageNumber="16">
|
|||
|
FIGURE 8.
|
|||
|
<taxonomicName id="4704559C7C4F125BCC87F1DF1C0E0774" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="novus">Trichoribates novus</taxonomicName>
|
|||
|
(Sellnick, 1928).
|
|||
|
<normalizedToken id="8D75EA0C5098557D3C34A485BF08752C" originalValue="A—Posterior" pageId="15" pageNumber="16">A-Posterior</normalizedToken>
|
|||
|
view of opisthosoma;
|
|||
|
<normalizedToken id="3866EC73CE88E51C5B8E317C44DA30E0" originalValue="B—Lateral" pageId="15" pageNumber="16">B-Lateral</normalizedToken>
|
|||
|
view of notogaster.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|