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<document id="AC53E42DB45CDE76452359FEE7700188" ID-DOI="10.5281/zenodo.184177" ID-GBIF-Dataset="0ff63046-5a50-49eb-b64d-1899ddd56b3d" ID-ISSN="1175-5326" ID-Zenodo-Dep="184177" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459951158418" checkinUser="plazi" docAuthor="Zhang, Kuiyan, Yang, Ding &amp; Ren, Dong" docDate="2008" docId="03B587ED4C32FFF3DDF7F19BF57E89CD" docLanguage="en" docName="zt01879p064.pdf" docOrigin="Zootaxa 1879" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Protobrachyceron sinensis Zhang, Yang &amp; Ren, 2008, sp. nov." docType="treatment" docVersion="8" lastPageNumber="64" masterDocId="FF8CFF954C30FFF0DD60F41CF4428B79" masterDocTitle="The first Middle Jurassic Protobrachyceron Handlirsch fly (Diptera: Brachycera: Protobrachyceridae) from Inner Mongolia (China)" masterLastPageNumber="64" masterPageNumber="61" pageNumber="63" updateTime="1698230833419" updateUser="plazi">
<mods:mods id="B0A21F6A477FFE02306742E9C4E643FC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E714C8AB9F68ABFE6EF4F7E6136E524D">
<mods:title id="EB0F14712D189CB9A83167B615F426F7">The first Middle Jurassic Protobrachyceron Handlirsch fly (Diptera: Brachycera: Protobrachyceridae) from Inner Mongolia (China)</mods:title>
</mods:titleInfo>
<mods:name id="CA8EABA072137C67A5CCB0CB20AB63D1" type="personal">
<mods:role id="812E27847F75DC964E2C3D3D252CF2F6">
<mods:roleTerm id="6DB1BDFC9AA514FD0D1357C191F04969">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00C5BC9453066FD1BCF9B389DD4AC38F">Zhang, Kuiyan</mods:namePart>
</mods:name>
<mods:name id="253DB9A1975E1BF438CDED4E4B129A1C" type="personal">
<mods:role id="0AF872B3C4CE89AD3B18DFF05B947C31">
<mods:roleTerm id="0F3027EB3C9008C314C4C35DD997F19B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75F8E411C8E0B92112F770B0AC724693">Yang, Ding</mods:namePart>
</mods:name>
<mods:name id="8A01896EFCB7216420C2D2BF6A8DE864" type="personal">
<mods:role id="810FABC7C54B08EF4E9A7E8827C4C8C8">
<mods:roleTerm id="34AD3804977885766EDDAB2922909403">Author</mods:roleTerm>
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<mods:namePart id="57F7AA5EFE778613E3CE4562C1E243EF">Ren, Dong</mods:namePart>
</mods:name>
<mods:typeOfResource id="54A21C16D49D884810D7A6C0CD0FE081">text</mods:typeOfResource>
<mods:relatedItem id="628BC21F19F98DA09DF68CF9E6560C93" type="host">
<mods:titleInfo id="BE748E303004DABDE19B4A1B17B853E6">
<mods:title id="1DC684283CD882FD526FCA84B1490DA7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9E6BFDCF9B7D4620D7BEF9D7CF230CF3">
<mods:date id="7B21873C8A4FCF832C7C44629CA34726">2008</mods:date>
<mods:detail id="BDE33682BB9FF8D84E226EA2D1A70DED" type="volume">
<mods:number id="0336616607F44C851DD7C0D6F5DE795E">1879</mods:number>
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<mods:start id="38C47D512C6278BAC0C488B5B18F6DD3">61</mods:start>
<mods:end id="243FF8105BE1F8CE32F2A4981EB70068">64</mods:end>
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<mods:classification id="F8FE446C9D4D1A0438C7C72971A38ED6">journal article</mods:classification>
<mods:identifier id="5741ADE023F51D1C9342EF61E345478D" type="DOI">10.5281/zenodo.184177</mods:identifier>
<mods:identifier id="D63756D3E20C2CA9DEE58F6CD34FBEAD" type="GBIF-Dataset">0ff63046-5a50-49eb-b64d-1899ddd56b3d</mods:identifier>
<mods:identifier id="E69074BDD6F75D83F36440359811BC8F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A9D6C9F80F9C40E3C67E119059B59F72" type="Zenodo-Dep">184177</mods:identifier>
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<treatment id="03B587ED4C32FFF3DDF7F19BF57E89CD" ID-DOI="http://doi.org/10.5281/zenodo.6236014" ID-GBIF-Taxon="119376527" ID-Zenodo-Dep="6236014" LSID="urn:lsid:plazi:treatment:03B587ED4C32FFF3DDF7F19BF57E89CD" httpUri="http://treatment.plazi.org/id/03B587ED4C32FFF3DDF7F19BF57E89CD" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
<subSubSection id="C30665704C32FFF2DDF7F19BF5498EB8" pageId="2" pageNumber="63" type="nomenclature">
<paragraph id="8BA336FB4C32FFF2DDF7F19BF6558EE6" blockId="2.[151,535,1415,1473]" box="[151,535,1415,1439]" pageId="2" pageNumber="63">
<heading id="D0EB81974C32FFF2DDF7F19BF6558EE6" bold="true" box="[151,535,1415,1439]" fontSize="10" level="2" pageId="2" pageNumber="63" reason="2">
<emphasis id="B968EAE94C32FFF2DDF7F19BF6558EE6" bold="true" box="[151,535,1415,1439]" pageId="2" pageNumber="63">
<taxonomicName id="4C1C4D784C32FFF2DDF7F19BF5FE8EE6" box="[151,444,1415,1439]" class="Insecta" family="Protobrachycerontidae" genus="Protobrachyceron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="sinensis" status="sp. nov.">
<emphasis id="B968EAE94C32FFF2DDF7F19BF5FE8EE6" bold="true" box="[151,444,1415,1439]" italics="true" pageId="2" pageNumber="63">Protobrachyceron sinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57924C32FFF2DCA2F19BF6558EE6" box="[450,535,1415,1439]" pageId="2" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336FB4C32FFF2DDF7F1B6F5498EB8" blockId="2.[151,535,1415,1473]" box="[151,267,1450,1473]" pageId="2" pageNumber="63">
(
<figureCitation id="13272A7E4C32FFF2DDFFF1B6F5468EB8" box="[159,260,1450,1473]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,255,1431,1455]" captionStartId-1="2.[151,255,1311,1335]" captionTargetBox-0="[150,1436,194,1405]" captionTargetBox-1="[231,1359,202,1281]" captionTargetId-0="figure@1.[147,1440,194,1407]" captionTargetId-1="figure@2.[226,1371,194,1286]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Protobrachyceron sinensis sp. nov. Body with wings, photograph in dorsal view." captionText-1="FIGURE 2. Protobrachyceron sinensis sp. nov. camera lucida drawing, based on the original photograph, in dorsal view." httpUri-0="https://zenodo.org/record/184178/files/figure.png" httpUri-1="https://zenodo.org/record/184179/files/figure.png" pageId="2" pageNumber="63">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30665704C32FFF2DDF7F1F3F6968D0F" pageId="2" pageNumber="63" type="diagnosis">
<paragraph id="8BA336FB4C32FFF2DDF7F1F3F6968D0F" blockId="2.[151,1436,1519,2013]" pageId="2" pageNumber="63">
<emphasis id="B968EAE94C32FFF2DDF7F1F3F54E8D7E" bold="true" box="[151,268,1519,1543]" pageId="2" pageNumber="63">Diagnosis.</emphasis>
Body small, head elliptic. Vein Sc slightly longer than cell br (basal radial); veins R1 and R2+3 close to each other at wing margin; vein Rs slightly longer than distance between beginning of vein R2+3 and crossvein r-m; veins M2, M3 much shorter than vein M1; vein CuA1 arising from cell d, vein CuP well visible; mouth of cell r1 narrow; relative widths of mouths of cells m1, m2 and m3 about 2: 2: 1.
</paragraph>
</subSubSection>
<subSubSection id="C30665704C32FFF2DDA6F298F72A8DE2" box="[198,872,1668,1692]" pageId="2" pageNumber="63" type="etymology">
<paragraph id="8BA336FB4C32FFF2DDA6F298F72A8DE2" blockId="2.[151,1436,1519,2013]" box="[198,872,1668,1692]" pageId="2" pageNumber="63">
<emphasis id="B968EAE94C32FFF2DDA6F298F5058DE5" bold="true" box="[198,327,1668,1692]" pageId="2" pageNumber="63">Etymology.</emphasis>
The specific name refers to the Latin
<emphasis id="B968EAE94C32FFF2DFBCF298F74A8DE2" box="[732,776,1668,1691]" italics="true" pageId="2" pageNumber="63">sina</emphasis>
(
<collectingCountry id="F30B766B4C32FFF2DE77F298F71E8DE2" box="[791,860,1668,1691]" name="China" pageId="2" pageNumber="63">China</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C30665704C32FFF2DDA6F2BBF7488C7A" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="8BA336FB4C32FFF2DDA6F2BBF09A8DC7" blockId="2.[151,1436,1519,2013]" box="[198,1240,1703,1727]" pageId="2" pageNumber="63">
<emphasis id="B968EAE94C32FFF2DDA6F2BBF5778DC6" bold="true" box="[198,309,1703,1727]" pageId="2" pageNumber="63">
<typeStatus id="54A788594C32FFF2DDA6F2BBF56C8DC6" box="[198,302,1703,1727]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
.
</emphasis>
CNU-DB-NN2008001, an almost complete adult body with both wings in dorsal view.
</paragraph>
<paragraph id="8BA336FB4C32FFF2DDA6F2D5F7488C7A" blockId="2.[151,1436,1519,2013]" pageId="2" pageNumber="63">
<emphasis id="B968EAE94C32FFF2DDA6F2D5F5AD8D98" bold="true" box="[198,495,1737,1761]" pageId="2" pageNumber="63">
<typeStatus id="54A788594C32FFF2DDA6F2D5F4BC8D98" box="[198,254,1737,1761]" pageId="2" pageNumber="63">Type</typeStatus>
locality and horizon.
</emphasis>
Daohugou Village, Shantou Township, Ningcheng County, Inner
<collectingCountry id="F30B766B4C32FFF2D9D1F2D6F1598D98" box="[1201,1307,1738,1761]" name="Mongolia" pageId="2" pageNumber="63">Mongolia</collectingCountry>
,
<collectingCountry id="F30B766B4C32FFF2D846F2D6F12B8D98" box="[1318,1385,1738,1761]" name="China" pageId="2" pageNumber="63">China</collectingCountry>
; Jiulongshan Formation, Middle Jurassic (Aalenian-Bajocian).
</paragraph>
</subSubSection>
<subSubSection id="C30665704C32FFF3DDA6F313F6468A3D" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63" type="description">
<paragraph id="8BA336FB4C32FFF2DDA6F313F0CA8C5F" blockId="2.[151,1436,1519,2013]" box="[198,1160,1807,1831]" pageId="2" pageNumber="63">
<emphasis id="B968EAE94C32FFF2DDA6F313F5138C5E" bold="true" box="[198,337,1807,1831]" pageId="2" pageNumber="63">Description.</emphasis>
Body length
<quantity id="4CE49B1E4C32FFF2DC81F313F6718C5F" box="[481,563,1807,1830]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="63" unit="mm" value="6.2">6.2 mm</quantity>
as preserved; wing length
<quantity id="4CE49B1E4C32FFF2DE31F313F7E78C5F" box="[849,933,1807,1830]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="63" unit="mm" value="5.8">5.8 mm</quantity>
, wing width
<quantity id="4CE49B1E4C32FFF2D94FF313F0C68C5F" box="[1071,1156,1807,1830]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="63" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<paragraph id="8BA336FB4C32FFF3DDA6F32EF6468A3D" blockId="2.[151,1436,1519,2013]" lastBlockId="3.[151,1436,152,692]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
Head elliptic in dorsal view, width about
<quantity id="4CE49B1E4C32FFF2DFF0F32EF6A58C30" box="[656,743,1842,1865]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="63" unit="mm" value="1.5">1.5 mm</quantity>
. Eyes large, bare, ommatidia clearly visible. Antennae not preserved. Relative widths of head and thorax about 0.7: 1. Thorax black. Legs slender. Wing elongate, hyaline: Vein C continued around wing as an ambient vein; vein Sc ending at midwing, slightly longer than cell br. Veins R1 and R2+3 close to each other at wing margin; vein Rs slightly longer than distance between beginning of vein R2+3 and crossvein r-m; vein R5 ending at apex of wing, slightly shorter than distance between crossvein r-m and beginning of vein R4; both vein R4 and cell d distinctly shorter than vein M
<quantity id="4CE49B1E4C33FFF3DF24F4BAF6258BD6" box="[580,615,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="64" unit="in" value="1.0">1 in</quantity>
length. Crossvein r-m at basal 1/4 of cell d; veins M2, M3 much shorter than vein M1. Vein CuA1 arising from cell d, vein CuP well visible. Mouth of cell sc slightly wider than that of cell r2+3; mouth of cell r1 narrow, about half as wide as that of cell r2+3. Cell d wide; five posterior cells widely open; relative widths of mouths of cells m1, m2 and m3 about 2: 2: 1. Abdomen brown, cylindrical; 6 segments visible. Segments II-IV distinctly long and broad. Gender unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30665704C33FFF3DDA6F553F57E89CD" pageId="3" pageNumber="64" type="discussion">
<paragraph id="8BA336FB4C33FFF3DDA6F553F57E89CD" blockId="3.[151,1436,152,692]" pageId="3" pageNumber="64">
<emphasis id="B968EAE94C33FFF3DDA6F553F5778A1E" bold="true" box="[198,309,335,359]" pageId="3" pageNumber="64">Remarks.</emphasis>
<taxonomicName id="4C1C4D784C33FFF3DC20F54CF6228A1E" box="[320,608,336,359]" class="Insecta" family="Protobrachycerontidae" genus="Protobrachyceron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="sinensis" status="sp. nov.">
<emphasis id="B968EAE94C33FFF3DC20F54CF6228A1E" box="[320,608,336,359]" italics="true" pageId="3" pageNumber="64">Protobrachyceron sinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57924C33FFF3DF0CF54CF6868A1E" box="[620,708,336,359]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C1C4D784C33FFF3DEFBF54CF09F8A1E" authority="Handlirsch, 1920" authorityName="Handlirsch" authorityYear="1920" box="[923,1245,336,359]" class="Insecta" family="Protobrachycerontidae" genus="Protobrachyceron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="liasinum">
<emphasis id="B968EAE94C33FFF3DEFBF54CF0548A1E" box="[923,1046,336,359]" italics="true" pageId="3" pageNumber="64">P. liasinum</emphasis>
<bibRefCitation id="EF8D4B0A4C33FFF3D941F54CF09F8A1E" author="Handlirsch" box="[1057,1245,336,359]" pageId="3" pageNumber="64" refString="Handlirsch, A. (1920) Palaeontologie (part), pp. 117 - 208 [= Lieferung 5]. In: Schroeder, C. W. M. (Ed.), Handbuch der Entomologie, Band III. G. Fischer, Jena. 1, 201 pp." type="book" year="1920">
Handlirsch,
<quantity id="4CE49B1E4C33FFF3D9C8F54CF09F8A1E" box="[1192,1245,336,359]" metricMagnitude="1" metricUnit="m" metricValue="4.8768" pageId="3" pageNumber="64" unit="in" value="1920.0">1920</quantity>
</bibRefCitation>
</taxonomicName>
in the following aspects: Veins R1 and R2+3 close to each other at wing margin; both vein R4 and cell d distinctly shorter than vein M
<quantity id="4CE49B1E4C33FFF3D818F59DF1DE8AF0" box="[1400,1436,370,398]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="64" unit="in" value="1.0">1 in</quantity>
length; mouth of cell sc distinctly wider than that of cell r1; five posterior cells widely open. However, we can easily separate it from
<taxonomicName id="4C1C4D784C33FFF3DC43F5A1F5DB8AAD" box="[291,409,445,468]" class="Insecta" family="Protobrachycerontidae" genus="Protobrachyceron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="liasinum">
<emphasis id="B968EAE94C33FFF3DC43F5A1F5DB8AAD" box="[291,409,445,468]" italics="true" pageId="3" pageNumber="64">P. liasinum</emphasis>
</taxonomicName>
by vein Rs slightly longer than distance between beginning of vein R2+3 and crossvein r-m; distance between crossvein r-m and beginning of vein R4 about 1.3 times as long as vein R4; vein CuA1 arising from cell d; mouth of cell r2+3 about twice as wide as that of cell r1; relative widths of mouths of cells m1, m2 and m3 about 2: 2: 1. In
<taxonomicName id="4C1C4D784C33FFF3DDF7F631F54C893D" box="[151,270,557,580]" class="Insecta" family="Protobrachycerontidae" genus="Protobrachyceron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="liasinum">
<emphasis id="B968EAE94C33FFF3DDF7F631F54C893D" box="[151,270,557,580]" italics="true" pageId="3" pageNumber="64">P. liasinum</emphasis>
</taxonomicName>
, vein Rs is more than twice as long as distance between beginning of vein R2+3 and crossvein r-m; distance between crossvein r-m and beginning of vein R4 is about twice as long as vein R4; vein CuA1 is arising from the infall of cell d and cell bm; mouth of cell r2+3 is slightly wider than that of cell r1; relative widths of mouths of cells m1, m2 and m3 is about 6: 9: 1.
</paragraph>
</subSubSection>
</treatment>
</document>
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