treatments-xml/data/03/C1/87/03C187CFFFBAFFAEFF38FB3D9F8D56D5.xml

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<document ID-DOI="10.11646/zootaxa.5239.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7635110" ID-ZooBank="8C81FAA8-9157-4476-84BB-54C6636BCFB1" checkinTime="1676283117990" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Mumladze, Levan, Grego, Jozef &amp; Giokas, Sinos" docDate="2023" docId="03C187CFFFBAFFAEFF38FB3D9F8D56D5" docLanguage="en" docName="zootaxa.5239.4.1.pdf" docOrigin="Zootaxa 5239 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acicula parcelineata" docType="treatment" docVersion="4" lastPageNumber="463" masterDocId="FFF8FFB7FFB0FFA2FFAFFFF39D54504C" masterDocTitle="The genus Acicula W. Hartmann, 1821 in the Caucasus (Gastropoda: Caenogastropoda: Aciculidae)" masterLastPageNumber="476" masterPageNumber="451" pageNumber="461" updateTime="1676406524359" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Acicula W. Hartmann, 1821 in the Caucasus (Gastropoda: Caenogastropoda: Aciculidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Páll-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eötvös Loránd Research Network, H- 1022 Budapest, Herman Ottó út 15, Hungary. pallgergely 2 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6167 - 7221</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mumladze, Levan</mods:namePart>
<mods:affiliation>Institute of Zoology, Ilia State University, Chavchavadze ave. 32, Tbilisi, Georgia.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-4977-0415</mods:nameIdentifier>
<mods:affiliation>Horná Mičiná 219, SK- 97401 Banská Bystrica, Slovakia. jozef. grego @ gmail. com; https: // orcid. org / 0000 - 0002 - 4977 - 0415 Department of Biology, University of Patras, Patras, 26500, Greece.</mods:affiliation>
<mods:nameIdentifier type="email">jozef.grego@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giokas, Sinos</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5239</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>451</mods:start>
<mods:end>476</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5239.4.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5239.4.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7635110</mods:identifier>
<mods:identifier type="ZooBank">8C81FAA8-9157-4476-84BB-54C6636BCFB1</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7635221" ID-Zenodo-Dep="7635221" LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5" httpUri="http://treatment.plazi.org/id/03C187CFFFBAFFAEFF38FB3D9F8D56D5" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461">
<subSubSection box="[151,589,1230,1257]" pageId="10" pageNumber="461" type="nomenclature">
<paragraph blockId="10.[151,589,1230,1293]" box="[151,589,1230,1257]" pageId="10" pageNumber="461">
<heading bold="true" box="[151,589,1230,1257]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
<taxonomicName ID-CoL="64HRD" authority="(Clessin, 1911)" baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[151,589,1230,1257]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="461" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis bold="true" box="[151,589,1230,1257]" pageId="10" pageNumber="461">
<emphasis bold="true" box="[151,401,1230,1257]" italics="true" pageId="10" pageNumber="461">Acicula parcelineata</emphasis>
(
<bibRefCitation author="Clessin, S." box="[417,582,1230,1256]" pageId="10" pageNumber="461" pagination="165 - 167" refId="ref10530" refString="Clessin, S. (1911) Neue Acme - Arten. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 43 (4), 165 - 167." type="journal article" year="1911">Clessin, 1911</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,289,1267,1293]" pageId="10" pageNumber="461" type="description">
<paragraph blockId="10.[151,589,1230,1293]" box="[151,289,1267,1293]" pageId="10" pageNumber="461">
(
<figureCitation box="[159,238,1267,1293]" captionStart="FIGURE 8" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1920]" captionTargetId="figure-18@11.[152,1436,181,1920]" captionTargetPageId="11" captionText="FIGURE 8. Acicula parcelineata (Clessin, 1911). A: 2018/19, specimen1; B: 2018/19, specimen2; C: 2018/17, specimen 1; D: ex LN 10681, specimen2; E: 2021/28x; F and G: shells collected near Tatariv village, Ukraine, photos from Skvortsova &amp; Balashov (2019); H: ex LN 10680, specimen1; I: ex LN 10680, specimen2. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7635130" httpUri="https://zenodo.org/record/7635130/files/figure.png" pageId="10" pageNumber="461">Figs. 8</figureCitation>
,
<figureCitation box="[251,280,1267,1293]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[296,1292,181,1920]" captionTargetId="figure-28@14.[296,1292,181,1920]" captionTargetPageId="14" captionText="FIGURE 10. Synoptic view of Caucasian Acicula. Upper left shell: Acicula limbata Reuss, 1861; other shells in the upper two rows: Acicula moussoni O. Boettger, 1879, third row: Acicula telum Páll-Gergely, n. sp., fourth row: Acicula parcelineata (Clessin, 1911)." figureDoi="http://doi.org/10.5281/zenodo.7635136" httpUri="https://zenodo.org/record/7635136/files/figure.png" pageId="10" pageNumber="461">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="461" type="reference_group">
<paragraph blockId="10.[151,742,1335,1421]" box="[151,542,1335,1359]" pageId="10" pageNumber="461">
<treatmentCitationGroup box="[151,542,1335,1359]" pageId="10" pageNumber="461">
<treatmentCitation author="Clessin, S." box="[151,539,1335,1359]" page="165" pageId="10" pageNumber="461" year="1911">
<taxonomicName ID-CoL="7PT8W" authority="Clessin, 1911: 165" authorityName="Clessin" authorityPageNumber="165" authorityYear="1911" box="[151,539,1335,1359]" class="Gastropoda" family="Aciculidae" genus="Acme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="461" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[151,341,1335,1359]" italics="true" pageId="10" pageNumber="461">Acme parcelineata</emphasis>
<bibRefCitation author="Clessin, S." box="[347,539,1335,1359]" pageId="10" pageNumber="461" pagination="165 - 167" refId="ref10530" refString="Clessin, S. (1911) Neue Acme - Arten. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 43 (4), 165 - 167." type="journal article" year="1911">Clessin, 1911: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="10.[151,742,1335,1421]" box="[151,681,1366,1390]" pageId="10" pageNumber="461">
<treatmentCitationGroup box="[151,681,1366,1390]" pageId="10" pageNumber="461">
<treatmentCitation author="Steklov, A. A." box="[151,541,1366,1390]" page="122" pageId="10" pageNumber="461" year="1966">
<taxonomicName ID-CoL="7PSVS" authority="Steklov, 1966: 122" authorityName="Steklov" authorityPageNumber="122" authorityYear="1966" box="[151,541,1366,1390]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="461" phylum="Mollusca" rank="species" species="bakanense">
<emphasis box="[151,342,1366,1390]" italics="true" pageId="10" pageNumber="461">Acicula bakanense</emphasis>
<bibRefCitation author="Steklov, A. A." box="[348,541,1366,1390]" pageId="10" pageNumber="461" pagination="279" refId="ref12757" refString="Steklov, A. A. (1966) Nazemnyye mollyuski neogena Predkavkaz'ya i ikh stratigraficheskoye znacheniye [Terrestrial molluscs of the Neogene Ciscaucasia and their stratigraphic significance]. Academy of Sciences of the USSR Transactions 163. Akademia Nauka S. S. S. R., Moskva, 279." type="journal article" year="1966">Steklov, 1966: 122</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 1, fig. 11.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="10.[151,742,1335,1421]" box="[151,742,1396,1421]" pageId="10" pageNumber="461">
<treatmentCitationGroup box="[151,742,1396,1421]" pageId="10" pageNumber="461">
<emphasis box="[151,385,1396,1420]" italics="true" pageId="10" pageNumber="461">
<taxonomicName ID-CoL="64HRD" baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[151,359,1396,1420]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="461" phylum="Mollusca" rank="species" species="parcelineata">Acicula parcelineata</taxonomicName>
</emphasis>
<treatmentCitation author="Boeters, H. D. &amp; Gittenberger, E. &amp; Subai, P." box="[385,617,1396,1421]" page="78" pageId="10" pageNumber="461" year="1989">
<bibRefCitation author="Boeters, H. D. &amp; Gittenberger, E. &amp; Subai, P." box="[385,617,1396,1421]" pageId="10" pageNumber="461" pagination="1 - 234" refId="ref10251" refString="Boeters, H. D., Gittenberger, E. &amp; Subai, P. (1989) Die Aciculidae (Mollusca: Gastropoda Prosobranchia). Zoologische Verhandelingen, 252, 1 - 234." type="journal article" year="1989">
Boeters
<emphasis box="[468,520,1396,1420]" italics="true" pageId="10" pageNumber="461">et al.</emphasis>
1989: 78
</bibRefCitation>
</treatmentCitation>
, figs 77, 78.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="461" type="diagnosis">
<paragraph blockId="10.[151,1437,1466,2033]" pageId="10" pageNumber="461">
<emphasis bold="true" box="[151,274,1466,1492]" pageId="10" pageNumber="461">Diagnosis.</emphasis>
A small (H:
<quantity box="[420,539,1467,1493]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="10" pageNumber="461" unit="mm" value="2.2" valueMax="2.4" valueMin="2.0">22.4 mm</quantity>
), mostly conical
<taxonomicName box="[731,906,1467,1493]" pageId="10" pageNumber="461">
<emphasis box="[731,817,1467,1493]" italics="true" pageId="10" pageNumber="461">Acicula</emphasis>
species
</taxonomicName>
with few and irregularly arranged longitudinal grooves and without neck bulge.
</paragraph>
<paragraph blockId="10.[151,1437,1466,2033]" box="[199,1073,1538,1565]" pageId="10" pageNumber="461">
<emphasis bold="true" box="[199,596,1538,1565]" pageId="10" pageNumber="461">Number of grooves on last whorl.</emphasis>
311, average: 5.56 (SD=+/-1.6) (n = 34).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="461" type="description">
<paragraph blockId="10.[151,1437,1466,2033]" pageId="10" pageNumber="461">
<emphasis bold="true" box="[199,492,1574,1601]" pageId="10" pageNumber="461">Measurements (in mm).</emphasis>
H: 1.622.68, average: 2.04 (SD=+/-0.32); D: 0.640.91, average: 0.76 (SD=+/- 0.09); D/H: 0.320.41, average: 0.38 (SD=+/-0.02) (n = 34).
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461" type="materials_examined">
<paragraph blockId="10.[151,1437,1466,2033]" lastBlockId="12.[151,1437,150,1689]" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461">
<emphasis bold="true" box="[199,435,1646,1672]" pageId="10" pageNumber="461">Material examined.</emphasis>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="10" pageNumber="461" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
GEORGIA, Samegrelo Reg.,
<collectingCounty box="[784,882,1647,1673]" pageId="10" pageNumber="461">Mukhuri</collectingCounty>
vicinity, road to
<collectingMunicipality box="[1081,1162,1647,1673]" pageId="10" pageNumber="461">Lugela</collectingMunicipality>
, small karst spring well just at left side of road from
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FE48F9609F1D56E1" box="[487,585,1683,1709]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Mukhuri" pageId="10" pageNumber="461" stateProvince="Imereti">Mukhuri</location>
to
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FDDDF9609F9756E1" box="[626,707,1683,1709]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Lugela" pageId="10" pageNumber="461" stateProvince="Imereti">Lugela</location>
, behind
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FC8AF9609EFC56E1" box="[805,936,1683,1709]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Shurubumu" pageId="10" pageNumber="461" stateProvince="Imereti">Shurubumu</location>
springs,
<geoCoordinate box="[1042,1185,1684,1711]" degrees="42" direction="north" minutes="38.985" orientation="latitude" pageId="10" pageNumber="461" precision="1" value="42.64975">42°38.985N</geoCoordinate>
<geoCoordinate box="[1194,1334,1684,1711]" degrees="42" direction="east" minutes="12.300" orientation="longitude" pageId="10" pageNumber="461" precision="1" value="42.205">42°12.300E</geoCoordinate>
(locality code:
<date box="[223,332,1719,1745]" pageId="10" pageNumber="461" value="2021-10-28">
<collectingDate box="[223,332,1719,1745]" pageId="10" pageNumber="461" value="2021-10-28">2021/28x</collectingDate>
</date>
), leg.
<collectorName box="[409,508,1719,1745]" pageId="10" pageNumber="461">J. Grego</collectorName>
&amp;
<collectorName box="[552,697,1719,1745]" pageId="10" pageNumber="461">M. Szekeres</collectorName>
, 17
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FD40F9449E1F569D" box="[751,843,1719,1745]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="October" pageId="10" pageNumber="461" stateProvince="Imereti">October</location>
, 2021, 1 figured shell (H:
<quantity box="[1162,1268,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="2.39" pageId="10" pageNumber="461" unit="mm" value="2.39">2.39 mm</quantity>
, D:
<quantity box="[1323,1428,1719,1745]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="10" pageNumber="461" unit="mm" value="0.88">0.88 mm</quantity>
) (coll. JG);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="10" pageNumber="461" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
<collectingRegion box="[272,356,1755,1781]" country="Georgia" name="Imereti" pageId="10" pageNumber="461">Imereti</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FEDFF9289CBF56B9" box="[368,491,1755,1781]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Nakhriduri" pageId="10" pageNumber="461" stateProvince="Imereti">Nakhriduri</location>
5 spring cave of
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FD01F9289E8A56B9" box="[686,990,1755,1781]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Turchu Gamosadivari river" pageId="10" pageNumber="461" stateProvince="Imereti">Turchu Gamosadivari river</location>
at the end of the canyon (locality code: 2018/17),
<geoCoordinate box="[268,414,1792,1819]" degrees="42" direction="north" minutes="29.216" orientation="latitude" pageId="10" pageNumber="461" precision="1" value="42.486935">42°29.216N</geoCoordinate>
,
<geoCoordinate box="[427,569,1792,1819]" degrees="42" direction="east" minutes="31.341" orientation="longitude" pageId="10" pageNumber="461" precision="1" value="42.52235">42°31.341E</geoCoordinate>
, leg.
<collectorName box="[633,730,1791,1817]" pageId="10" pageNumber="461">J. Grego</collectorName>
&amp;
<collectorName box="[770,927,1791,1817]" pageId="10" pageNumber="461">L. Mumladze</collectorName>
, 3
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FC6CF90C9EA35755" box="[963,1015,1791,1817]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="May" pageId="10" pageNumber="461" stateProvince="Imereti">May</location>
, 2018, 1 shell (H:
<quantity box="[1232,1336,1791,1817]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="10" pageNumber="461" unit="mm" value="2.03">2.03 mm</quantity>
, D:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="10" pageNumber="461" unit="mm" value="0.76">0.76 mm</quantity>
) (coll. JG);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="10" pageNumber="461" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
<collectingRegion box="[332,413,1827,1853]" country="Georgia" name="Imereti" pageId="10" pageNumber="461">Imereti</collectingRegion>
reg.,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FE72F8D09F665771" box="[477,562,1827,1853]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Tkibuli" pageId="10" pageNumber="461" stateProvince="Imereti">Tkibuli</location>
,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FD92F8D09E675771" box="[573,819,1827,1853]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Tkibuli-Nikortsminda" pageId="10" pageNumber="461" stateProvince="Imereti">Tkibuli-Nikortsminda</location>
road left side, spring with travertine waterfall (locality code: 2018/19),
<geoCoordinate box="[339,481,1864,1891]" degrees="42" direction="north" minutes="22.976" orientation="latitude" pageId="10" pageNumber="461" precision="1" value="42.382935">42°22.976N</geoCoordinate>
<geoCoordinate box="[492,620,1864,1891]" degrees="43" direction="east" minutes="0.730" orientation="longitude" pageId="10" pageNumber="461" precision="1" value="43.012165">43°0.730E</geoCoordinate>
, leg.
<collectorName box="[685,783,1863,1889]" pageId="10" pageNumber="461">J. Grego</collectorName>
&amp;
<collectorName box="[824,981,1863,1889]" pageId="10" pageNumber="461">L. Mumladze</collectorName>
,
<date box="[994,1166,1863,1889]" pageId="10" pageNumber="461" value="2018-05-04" valueMax="2018-05-08" valueMin="2018-05-04">
<collectingDate box="[994,1166,1863,1889]" pageId="10" pageNumber="461" value="2018-05-04" valueMax="2018-05-08" valueMin="2018-05-04">48 May 2018</collectingDate>
</date>
, 6 shells (H:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.285" metricValueMax="2.34" metricValueMin="2.23" pageId="10" pageNumber="461" unit="mm" value="2.285" valueMax="2.34" valueMin="2.23">2.232.34 mm</quantity>
, D:
<quantity box="[246,410,1898,1925]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="8.7" metricValueMin="8.5" pageId="10" pageNumber="461" unit="mm" value="0.86" valueMax="0.87" valueMin="0.85">0.850.87 mm</quantity>
) (coll. JG);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="10" pageNumber="461" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
<collectingRegion box="[549,630,1899,1925]" country="Georgia" name="Imereti" pageId="10" pageNumber="461">Imereti</collectingRegion>
reg.,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FD18F8989E1057C9" box="[695,836,1899,1925]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Tskhradjvari" pageId="10" pageNumber="461" stateProvince="Imereti">Tskhradjvari</location>
pass, limestone rocks and karst forest (locality code: C2458),
<geoCoordinate box="[249,394,1936,1963]" degrees="42" direction="north" minutes="23.405" orientation="latitude" pageId="10" pageNumber="461" precision="1" value="42.390083">42°23.405N</geoCoordinate>
,
<geoCoordinate box="[405,547,1936,1963]" degrees="42" direction="east" minutes="58.421" orientation="longitude" pageId="10" pageNumber="461" precision="1" value="42.973682">42°58.421E</geoCoordinate>
, leg.
<collectorName box="[606,704,1935,1961]" pageId="10" pageNumber="461">J. Grego</collectorName>
,
<date box="[714,852,1935,1961]" pageId="10" pageNumber="461" value="2018-05-04">
<collectingDate box="[714,852,1935,1961]" pageId="10" pageNumber="461" value="2018-05-04">4 May 2018</collectingDate>
</date>
, 2 shells (coll. JG);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="L. Mumladze" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="10" pageNumber="461" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
<collectingRegion box="[1073,1154,1935,1961]" country="Georgia" name="Imereti" pageId="10" pageNumber="461">Imereti</collectingRegion>
reg.,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FB6EF87C984457E5" box="[1217,1296,1935,1961]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Sairme" pageId="10" pageNumber="461" stateProvince="Imereti">Sairme</location>
gorge,
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.925" pageId="10" pageNumber="461" unit="m" value="1925.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.925" pageId="10" pageNumber="461" unit="m" value="1925.0">1925 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate box="[243,388,1972,1999]" degrees="41" direction="north" minutes="51.386" orientation="latitude" pageId="10" pageNumber="461" precision="1" value="41.856434">41°51.386N</geoCoordinate>
,
<geoCoordinate box="[398,538,1972,1999]" degrees="42" direction="east" minutes="47.406" orientation="longitude" pageId="10" pageNumber="461" precision="1" value="42.7901">42°47.406E</geoCoordinate>
(locality code: sp8s3), leg.
<collectorName box="[841,994,1971,1997]" pageId="10" pageNumber="461">L. Mumladze</collectorName>
,
<date box="[1003,1191,1971,1997]" pageId="10" pageNumber="461" value="2013-10-11">
<collectingDate box="[1003,1191,1971,1997]" pageId="10" pageNumber="461" value="2013-10-11">11 October 2013</collectingDate>
</date>
, 1 shell (
<collectionCode box="[1287,1337,1971,1997]" country="USA" httpUri="http://grbio.org/cool/jgjv-qdg3" name="Indiana State University" pageId="10" pageNumber="461">ISU</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Mukhuri" elevation="1925" lastPageId="12" lastPageNumber="463" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="10" pageNumber="461" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
<collectingRegion box="[1355,1436,1971,1997]" country="Georgia" name="Imereti" pageId="10" pageNumber="461">Imereti</collectingRegion>
reg.,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FF7EF8249C2357BD" box="[209,375,2007,2033]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Kinchkhaperdi" pageId="10" pageNumber="461" stateProvince="Imereti">Kinchkhaperdi</location>
vicinity,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FE4DF8249F6757BD" box="[482,563,2007,2033]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Okatse" pageId="10" pageNumber="461" stateProvince="Imereti">Okatse</location>
, spring at left bank of
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBAFFA8FC9AF8249EB957BD" box="[821,1005,2007,2033]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Satsiskvilo river" pageId="10" pageNumber="461" stateProvince="Imereti">Satsiskvilo river</location>
canyon, leakage among larger stones,
<geoCoordinate box="[151,297,152,179]" degrees="42" direction="north" minutes="31.102" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="42.518368">42°31.102N</geoCoordinate>
,
<geoCoordinate box="[311,450,152,179]" degrees="42" direction="east" minutes="33.437" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="42.55728">42°33.437E</geoCoordinate>
(locality code: F1940), leg.
<collectorName box="[784,882,151,177]" pageId="12" pageNumber="463">J. Grego</collectorName>
&amp;
<collectorName box="[924,1068,151,177]" pageId="12" pageNumber="463">M. Szekeres</collectorName>
, 15
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFBCEFF6499E950FD" box="[1121,1213,151,177]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="October" pageId="12" pageNumber="463" stateProvince="Imereti">October</location>
, 2021, 2 dry shells (coll. JG);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="L. Mumladze" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="12" pageNumber="463" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFEBEFF489CDD5099" box="[273,393,187,213]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Samegrelo" pageId="12" pageNumber="463" stateProvince="Imereti">Samegrelo</location>
reg.,
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFE65FF489FE75099" box="[458,691,187,213]" country="Georgia" county="Mukhuri" latitude="42.497017" longLatPrecision="1" longitude="42.547115" municipality="Lugela" name="Tekhuri River valley" pageId="12" pageNumber="463" stateProvince="Imereti">Tekhuri River valley</location>
,
<elevation box="[703,836,186,213]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="12" pageNumber="463" unit="m" value="465.0">
<quantity box="[703,775,186,213]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="12" pageNumber="463" unit="m" value="465.0">465 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate box="[849,994,188,215]" degrees="42" direction="north" minutes="36.035" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="42.600582">42°36.035N</geoCoordinate>
,
<geoCoordinate box="[1006,1145,188,215]" degrees="42" direction="east" minutes="20.797" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="42.346615">42°20.797E</geoCoordinate>
(locality code: leb3), leg.
<collectorName box="[151,306,223,249]" pageId="12" pageNumber="463">L. Mumladze</collectorName>
,
<date box="[317,498,222,249]" pageId="12" pageNumber="463" value="2015-08-21">
<collectingDate box="[317,498,222,249]" pageId="12" pageNumber="463" value="2015-08-21">21 August 2015</collectingDate>
</date>
,
<specimenCount box="[509,626,223,249]" pageId="12" pageNumber="463" type="subadult">2 subadult</specimenCount>
shells (
<collectionCode box="[712,762,223,249]" country="USA" httpUri="http://grbio.org/cool/jgjv-qdg3" name="Indiana State University" pageId="12" pageNumber="463">ISU</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="12" pageNumber="463" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
Samegrelo reg., Chkvaleri vicinity, Kvatsalara, small cave spring just left side of the road, to Intsira waterfall,
<geoCoordinate box="[743,889,260,287]" degrees="42" direction="north" minutes="43.275" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="42.72125">42°43.275N</geoCoordinate>
,
<geoCoordinate box="[901,1026,260,287]" degrees="42" direction="east" minutes="5.498" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="42.091633">42°5.498E</geoCoordinate>
(locality ode: F1968), leg.
<collectorName box="[1340,1436,259,285]" pageId="12" pageNumber="463">J. Grego</collectorName>
&amp;
<collectorName box="[180,321,295,321]" pageId="12" pageNumber="463">M. Szekeres</collectorName>
, 19 October, 2021, 1 broken shell (coll. JG);
</materialsCitation>
<materialsCitation collectingDate="2013-10-11" collectingDateMax="2021-10-28" collectingDateMin="2013-10-11" collectionCode="ISU" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Mukhuri" elevation="1925" latitude="42.497017" location="Mukhuri" longLatPrecision="1" longitude="42.547115" municipality="Lugela" pageId="12" pageNumber="463" specimenCount="2" specimenCount-subadult="2" stateProvince="Imereti">
Georgia,
<collectingRegion box="[931,1012,295,321]" country="Georgia" name="Imereti" pageId="12" pageNumber="463">Imereti</collectingRegion>
reg., Satsiskvilo vicinity, Turchismta, concrete wall and natural outlets among limestone beds at right side of path from Satsiskvilo to Okatse Spring (locality code: 2021/26),
<geoCoordinate box="[433,578,368,395]" degrees="42" direction="north" minutes="29.821" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="42.497017">42°29.821N</geoCoordinate>
,
<geoCoordinate box="[589,732,368,395]" degrees="42" direction="east" minutes="32.827" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="42.547115">42°32.827E</geoCoordinate>
, leg.
<collectorName box="[790,885,367,393]" pageId="12" pageNumber="463">J. Grego</collectorName>
&amp;
<collectorName box="[920,1060,367,393]" pageId="12" pageNumber="463">M. Szekeres</collectorName>
, 15 October, 2021, 1 broken shell (coll. JG)
</materialsCitation>
.
<materialsCitation collectingDate="1988-06-16" collectingDateMax="1988-06-20" collectingDateMin="1988-06-16" collectionCode="LN" collectorName="L. Nemeth" country="Russia" county="Western Caucasus" latitude="43.5476" location="Khosta river" longLatPrecision="1" longitude="39.881535" municipality="Sochi" pageId="12" pageNumber="463" specimenCode="LN 10680, LN 10524, LN 10563, LN 10720" specimenCount="42" specimenCount-adult="30" specimenCount-juvenile="12">
<collectingCountry box="[276,378,403,429]" name="Russia" pageId="12" pageNumber="463">RUSSIA</collectingCountry>
,
<collectingCounty box="[391,601,403,429]" pageId="12" pageNumber="463">Western Caucasus</collectingCounty>
, vicinity of
<collectingMunicipality box="[742,809,403,429]" pageId="12" pageNumber="463">Sochi</collectingMunicipality>
, valley of the
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFC7FFE60993451E1" box="[976,1120,403,429]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Khosta river" pageId="12" pageNumber="463">Khosta river</location>
, at
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFB24FE60986F51E1" box="[1163,1339,403,429]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Rassvet village" pageId="12" pageNumber="463">Rassvet village</location>
, flotsam of
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFF18FE449C5D519D" box="[183,265,439,465]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Khosta" pageId="12" pageNumber="463">Khosta</location>
,
<geoCoordinate box="[278,423,440,467]" degrees="43" direction="north" minutes="32.856" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="43.5476">43°32.856N</geoCoordinate>
,
<geoCoordinate box="[437,579,440,467]" degrees="39" direction="east" minutes="52.892" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="39.881535">39°52.892E</geoCoordinate>
, leg.
<collectorName box="[642,767,439,465]" pageId="12" pageNumber="463">L. Németh</collectorName>
,
<date box="[779,935,439,465]" pageId="12" pageNumber="463" value="1988-06-20">
<collectingDate box="[779,935,439,465]" pageId="12" pageNumber="463" value="1988-06-20">20 June 1988</collectingDate>
</date>
, 68 j/b shells +
<specimenCount box="[1120,1212,439,465]" pageId="12" pageNumber="463" type="adult">22 adult</specimenCount>
shells (
<specimenCode box="[1301,1424,439,465]" collectionCode="LN" pageId="12" pageNumber="463">LN 10680</specimenCode>
);
</materialsCitation>
<materialsCitation collectingDate="1988-06-16" collectingDateMax="1988-06-20" collectingDateMin="1988-06-16" collectionCode="LN" collectorName="L. Nemeth" country="Russia" county="Western Caucasus" latitude="43.5476" location="Khosta river" longLatPrecision="1" longitude="39.881535" municipality="Sochi" pageId="12" pageNumber="463" specimenCode="LN 10680, LN 10524, LN 10563, LN 10720" specimenCount="42" specimenCount-adult="30" specimenCount-juvenile="12">
Abkhazia, Novij Afon (Akhali Athoni), above the tunnel near the railway station,
<geoCoordinate box="[1068,1198,476,503]" degrees="43" direction="north" minutes="5.518" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="43.09197">43°5.518N</geoCoordinate>
,
<geoCoordinate box="[1210,1353,476,503]" degrees="40" direction="east" minutes="48.938" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="40.81563">40°48.938E</geoCoordinate>
, leg.
<collectorName pageId="12" pageNumber="463">L. Németh</collectorName>
,
<date box="[255,410,511,537]" pageId="12" pageNumber="463" value="1988-06-16">
<collectingDate box="[255,410,511,537]" pageId="12" pageNumber="463" value="1988-06-16">16 June 1988</collectingDate>
</date>
, 1 broken shell (
<specimenCode box="[598,720,510,537]" collectionCode="LN" pageId="12" pageNumber="463">LN 10524</specimenCode>
);
</materialsCitation>
<materialsCitation collectingDate="1988-06-16" collectingDateMax="1988-06-20" collectingDateMin="1988-06-16" collectionCode="LN" collectorName="L. Nemeth" country="Russia" county="Western Caucasus" latitude="43.5476" location="Khosta river" longLatPrecision="1" longitude="39.881535" municipality="Sochi" pageId="12" pageNumber="463" specimenCode="LN 10680, LN 10524, LN 10563, LN 10720" specimenCount="42" specimenCount-adult="30" specimenCount-juvenile="12">
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFD4BFE0C9EE05255" box="[740,948,511,537]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Western Caucasus" pageId="12" pageNumber="463">Western Caucasus</location>
, valley of
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFB83FE0C999D5255" box="[1068,1225,511,537]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="River Mzimta" pageId="12" pageNumber="463">River Mzimta</location>
towards Krasnaya Polyana, 26/33. km, in the &quot;gorge&quot; (approximate GPS coordinates:
<geoCoordinate box="[906,1051,548,575]" degrees="43" direction="north" minutes="31.267" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="43.52112">43°31.267N</geoCoordinate>
,
<geoCoordinate box="[1063,1206,548,575]" degrees="39" direction="east" minutes="59.903" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="39.998383">39°59.903E</geoCoordinate>
), leg.
<collectorName box="[1274,1397,547,573]" pageId="12" pageNumber="463">L. Németh</collectorName>
,
<date pageId="12" pageNumber="463" value="1988-06-18">
<collectingDate pageId="12" pageNumber="463" value="1988-06-18">18 June 1988</collectingDate>
</date>
, 3 shells (
<specimenCode box="[387,509,582,609]" collectionCode="LN" pageId="12" pageNumber="463">LN 10563</specimenCode>
);
</materialsCitation>
<materialsCitation collectingDate="1988-06-16" collectingDateMax="1988-06-20" collectingDateMin="1988-06-16" collectionCode="LN" collectorName="L. Nemeth" country="Russia" county="Western Caucasus" latitude="43.5476" location="Khosta river" longLatPrecision="1" longitude="39.881535" municipality="Sochi" pageId="12" pageNumber="463" specimenCode="LN 10680, LN 10524, LN 10563, LN 10720" specimenCount="42" specimenCount-adult="30" specimenCount-juvenile="12">
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFDBCFDB49FB0522D" box="[531,740,583,609]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Western Caucasus" pageId="12" pageNumber="463">Western Caucasus</location>
, valley of
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFCF1FDB49948522D" box="[862,1052,583,609]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="River Dagomysi" pageId="12" pageNumber="463">River Dagomysi</location>
, ca.
<quantity box="[1106,1182,583,609]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="463" unit="km" value="10.0">10 km</quantity>
, vicinity of “Nor Luis village” (=Altmets), approximate GPS coordinates:
<geoCoordinate box="[726,872,620,647]" degrees="43" direction="north" minutes="43.333" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="43.722218">43°43.333N</geoCoordinate>
,
<geoCoordinate box="[882,1024,620,647]" degrees="39" direction="east" minutes="40.500" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="39.675">39°40.500E</geoCoordinate>
, river debris, leg.
<collectorName box="[1220,1342,619,645]" pageId="12" pageNumber="463">L. Németh</collectorName>
,
<date pageId="12" pageNumber="463" value="1988-06-19">
<collectingDate pageId="12" pageNumber="463" value="1988-06-19">19 June 1988</collectingDate>
</date>
, 7 shell fragments (
<specimenCode box="[437,560,655,681]" collectionCode="LN" pageId="12" pageNumber="463">LN 10720</specimenCode>
);
</materialsCitation>
<materialsCitation collectingDate="1988-06-16" collectingDateMax="1988-06-20" collectingDateMin="1988-06-16" collectionCode="LN" collectorName="L. Nemeth" country="Russia" county="Western Caucasus" latitude="43.5476" location="Khosta river" longLatPrecision="1" longitude="39.881535" municipality="Sochi" pageId="12" pageNumber="463" specimenCode="LN 10680, LN 10524, LN 10563, LN 10720" specimenCount="42" specimenCount-adult="30" specimenCount-juvenile="12">
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFDEAFD7C9E4252E5" box="[581,790,655,681]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Western Caucasus" pageId="12" pageNumber="463">Western Caucasus</location>
, vicinity of Sochi, valley of the
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFB23FD7C984F52E5" box="[1164,1307,655,681]" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Khosta river" pageId="12" pageNumber="463">Khosta river</location>
, at
<location LSID="urn:lsid:plazi:treatment:03C187CFFFBAFFAEFF38FB3D9F8D56D5:8EB76002FFBCFFAEFAEAFD7C9DB25281" country="Russia" county="Western Caucasus" latitude="43.5476" longLatPrecision="1" longitude="39.881535" municipality="Sochi" name="Rassvet village" pageId="12" pageNumber="463">Rassvet village</location>
, flotsam of Khosta,
<geoCoordinate box="[458,603,692,719]" degrees="43" direction="north" minutes="32.856" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="43.5476">43°32.856N</geoCoordinate>
,
<geoCoordinate box="[615,757,692,719]" degrees="39" direction="east" minutes="52.892" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="39.881535">39°52.892E</geoCoordinate>
, leg.
<collectorName box="[817,941,691,717]" pageId="12" pageNumber="463">L. Németh</collectorName>
,
<date box="[952,1105,691,717]" pageId="12" pageNumber="463" value="1988-06-20">
<collectingDate box="[952,1105,691,717]" pageId="12" pageNumber="463" value="1988-06-20">20 June 1988</collectingDate>
</date>
,
<specimenCount box="[1117,1242,691,717]" pageId="12" pageNumber="463" type="juvenile">12 juvenile</specimenCount>
/broken +
<specimenCount box="[1359,1436,691,717]" pageId="12" pageNumber="463" type="adult">8 adult</specimenCount>
shells (ex LN 10681)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635130" ID-Zenodo-Dep="7635130" httpUri="https://zenodo.org/record/7635130/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1943,1967]" targetBox="[152,1436,181,1920]" targetPageId="11">
<paragraph blockId="11.[151,1436,1943,2040]" pageId="11" pageNumber="462">
<emphasis bold="true" box="[151,277,1943,1967]" pageId="11" pageNumber="462">FIGURE 8.</emphasis>
<taxonomicName authority="(Clessin, 1911)" baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[285,665,1944,1968]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="462" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[285,498,1944,1968]" italics="true" pageId="11" pageNumber="462">Acicula parcelineata</emphasis>
(
<bibRefCitation author="Clessin, S." box="[513,657,1944,1968]" pageId="11" pageNumber="462" pagination="165 - 167" refId="ref10530" refString="Clessin, S. (1911) Neue Acme - Arten. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 43 (4), 165 - 167." type="journal article" year="1911">Clessin, 1911</bibRefCitation>
)
</taxonomicName>
. A: 2018/19, specimen1; B: 2018/19, specimen2; C: 2018/17, specimen 1; D: ex LN 10681, specimen2; E: 2021/28x; F and G: shells collected near Tatariv village, Ukraine, photos from
<bibRefCitation author="Skvortsova, V. &amp; Balashov, I." pageId="11" pageNumber="462" pagination="94 - 102" refId="ref12693" refString="Skvortsova, V. &amp; Balashov, I. (2019) Land snail Acicula parcelineata (Architaenioglossa: Cyclophoroidea: Aciculidae) in Ukraine: distribution, variability, habitat preferences and conservation status. Ruthenica, 29 (2), 94 - 102. https: // doi. org / 10.35885 / ruthenica. 2019.29 (2). 3" type="journal article" year="2019">Skvortsova &amp; Balashov (2019)</bibRefCitation>
; H: ex LN 10680, specimen1; I: ex LN 10680, specimen2. All photos: B. Páll-Gergely.
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="463" type="diagnosis">
<paragraph blockId="12.[151,1437,150,1689]" pageId="12" pageNumber="463">
<emphasis bold="true" box="[199,463,762,788]" pageId="12" pageNumber="463">Differential diagnosis.</emphasis>
<taxonomicName authorityName="O. Boettger" authorityYear="1879" box="[470,670,763,789]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="moussoni">
<emphasis box="[470,670,763,789]" italics="true" pageId="12" pageNumber="463">Acicula moussoni</emphasis>
</taxonomicName>
was found syntopically with
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[1004,1177,763,789]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[1004,1177,763,789]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
at six sites (LN10561- LN10680-LN10681, sp8s3, F1968, 2021/26, F1940,
<date box="[758,867,799,825]" pageId="12" pageNumber="463" value="2021-10-28">2021/28x</date>
, see Appendix
<figureCitation box="[1046,1182,799,825]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1840,1864]" captionStartId-1="3.[151,250,1981,2005]" captionStartId-2="4.[151,250,1795,1819]" captionTargetBox-0="[151,1436,414,1814]" captionTargetBox-1="[239,1353,1367,1950]" captionTargetBox-2="[154,1427,185,1763]" captionTargetId-0="figure-129@2.[151,1436,414,1814]" captionTargetId-1="figure-408@3.[235,1353,1367,1958]" captionTargetId-2="figure-28@4.[151,1436,181,1771]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. A: An Acicula shell positioned with the axis of the shell on the y-axis and the aperture in the same plane as the objective, showing the location of the 18 landmarks used. B: counting of grooves on the last whorl. Images not to scale. Photos: B. Páll-Gergely." captionText-1="FIGURE 2. Means ± 95% Confidence Intervals of (A) Log centroid size, and (B) of the number of grooves, for the studied Acicula species, based on landmark data." captionText-2="FIGURE 3. (A) Principal Components Analysis and (B) Canonical Variates Analysis for the studied Acicula species, and the ordinations of specimens, with 90% confidence ellipses, based on landmark data." figureDoi-0="http://doi.org/10.5281/zenodo.7635112" figureDoi-1="http://doi.org/10.5281/zenodo.7635118" figureDoi-2="http://doi.org/10.5281/zenodo.7635120" httpUri-0="https://zenodo.org/record/7635112/files/figure.png" httpUri-1="https://zenodo.org/record/7635118/files/figure.png" httpUri-2="https://zenodo.org/record/7635120/files/figure.png" pageId="12" pageNumber="463">Figures 13</figureCitation>
) without intermediate forms, which indicates that in spite of the close similarity, they are distinct species. The shell differences are the following:
<taxonomicName authorityName="O. Boettger" authorityYear="1879" box="[273,409,871,897]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="moussoni">
<emphasis box="[273,409,871,897]" italics="true" pageId="12" pageNumber="463">A. moussoni</emphasis>
</taxonomicName>
is generally darker and larger (although the size ranges largely overlap, see Appendix
<figureCitation box="[1356,1421,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[154,1427,185,1763]" captionTargetId="figure-28@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 3. (A) Principal Components Analysis and (B) Canonical Variates Analysis for the studied Acicula species, and the ordinations of specimens, with 90% confidence ellipses, based on landmark data." figureDoi="http://doi.org/10.5281/zenodo.7635120" httpUri="https://zenodo.org/record/7635120/files/figure.png" pageId="12" pageNumber="463">Fig. 3</figureCitation>
), which is the most important trait that helps distinguish the two species in the case of mixed samples. Furthermore,
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[151,325,943,969]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[151,325,943,969]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
is rather conical or cylindrical, i.e., the sides of the shell are straight; the whorls grow regularly. In contrast,
<taxonomicName authorityName="O. Boettger" authorityYear="1879" box="[290,431,979,1005]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="moussoni">
<emphasis box="[290,431,979,1005]" italics="true" pageId="12" pageNumber="463">A. moussoni</emphasis>
</taxonomicName>
is rather ovoid-spindle-shaped. The whorls are flatter (less bulging) in
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[1262,1437,979,1005]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[1262,1437,979,1005]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
than in
<taxonomicName authorityName="O. Boettger" authorityYear="1879" box="[236,373,1015,1041]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="moussoni">
<emphasis box="[236,373,1015,1041]" italics="true" pageId="12" pageNumber="463">A. moussoni</emphasis>
</taxonomicName>
. Lastly,
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[470,642,1015,1041]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[470,642,1015,1041]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
has generally fewer grooves above the aperture, and they are arranged less regularly than those of
<taxonomicName authorityName="O. Boettger" authorityYear="1879" box="[463,601,1051,1077]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="moussoni">
<emphasis box="[463,601,1051,1077]" italics="true" pageId="12" pageNumber="463">A. moussoni</emphasis>
</taxonomicName>
(i.e., between two grooves, where a third one would be expected if it was regularly arranged, the intermediate one is absent).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="463" type="distribution">
<paragraph blockId="12.[151,1437,150,1689]" pageId="12" pageNumber="463">
<emphasis bold="true" box="[199,353,1122,1148]" pageId="12" pageNumber="463">Distribution.</emphasis>
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[361,596,1123,1149]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[361,596,1123,1149]" italics="true" pageId="12" pageNumber="463">Acicula parcelineata</emphasis>
</taxonomicName>
is known from the northern half of the Greater Caucasus Mountains, and one locality is known in the Lesser Caucasus Mountains (
<figureCitation box="[796,865,1159,1185]" captionStart="FIGURE 7" captionStartId="9.[151,250,1123,1147]" captionTargetBox="[151,1436,453,1100]" captionTargetId="figure-157@9.[151,1436,453,1100]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Acicula species in the Caucasus Mts. The symbols marked with numbers (1: Surami range, 2: Manglisi, 3: Martkopi, 4: North Ossetia) do not represent georeferenced localities for Acicula moussoni O. Boettger, 1879 and Acicula parcelineata (Clessin, 1911)." figureDoi="http://doi.org/10.5281/zenodo.7635128" httpUri="https://zenodo.org/record/7635128/files/figure.png" pageId="12" pageNumber="463">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="463" type="discussion">
<paragraph blockId="12.[151,1437,150,1689]" pageId="12" pageNumber="463">
<emphasis bold="true" box="[199,315,1195,1221]" pageId="12" pageNumber="463">Remarks.</emphasis>
This species was originally described from 'Dzingelau' (Dzięgielów in southern
<collectingCountry box="[1240,1322,1195,1221]" name="Poland" pageId="12" pageNumber="463">Poland</collectingCountry>
), and was subsequently reported from
<collectingCountry box="[473,656,1231,1257]" name="Czech Republic" pageId="12" pageNumber="463">Czech Republic</collectingCountry>
,
<collectingCountry box="[669,773,1231,1257]" name="Romania" pageId="12" pageNumber="463">Romania</collectingCountry>
,
<collectingCountry box="[787,885,1231,1257]" name="Slovakia" pageId="12" pageNumber="463">Slovakia</collectingCountry>
and the Carpathian part of
<collectingCountry box="[1205,1295,1231,1257]" name="Ukraine" pageId="12" pageNumber="463">Ukraine</collectingCountry>
(
<bibRefCitation author="Jackiewicz, M." pageId="12" pageNumber="463" pagination="445 - 454" refId="ref11273" refString="Jackiewicz, M. (1974) Krajowe gatunki rodziny Aciculidae (Mollusca, Prosobranchia). Fragmenta Faunistica, 19 (17), 445 - 454. https: // doi. org / 10.3161 / 00159301 FF 1974.19.17.445" type="journal article" year="1974">Jackiewicz 1974</bibRefCitation>
;
<bibRefCitation author="Boeters, H. D. &amp; Gittenberger, E. &amp; Subai, P." box="[222,436,1267,1293]" pageId="12" pageNumber="463" pagination="1 - 234" refId="ref10251" refString="Boeters, H. D., Gittenberger, E. &amp; Subai, P. (1989) Die Aciculidae (Mollusca: Gastropoda Prosobranchia). Zoologische Verhandelingen, 252, 1 - 234." type="journal article" year="1989">
Boeters
<emphasis box="[314,371,1267,1293]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
1989
</bibRefCitation>
;
<bibRefCitation author="Horsak, M. &amp; Jurickova, L. &amp; Picka, J." box="[448,660,1267,1293]" pageId="12" pageNumber="463" refId="ref11215" refString="Horsak, M., Jurickova, L. &amp; Picka, J. (2013) Molluscs of the Czech and Slovak Republics. Kabourek, Zlin, 264 pp. [in Czech and English]" type="book" year="2013">
Horsák
<emphasis box="[536,592,1267,1293]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
2013
</bibRefCitation>
).
<bibRefCitation author="Boeters, H. D. &amp; Gittenberger, E. &amp; Subai, P." box="[678,908,1267,1293]" pageId="12" pageNumber="463" pagination="1 - 234" refId="ref10251" refString="Boeters, H. D., Gittenberger, E. &amp; Subai, P. (1989) Die Aciculidae (Mollusca: Gastropoda Prosobranchia). Zoologische Verhandelingen, 252, 1 - 234." type="journal article" year="1989">
Boeters
<emphasis box="[771,827,1267,1293]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
(1989)
</bibRefCitation>
also reported it from flotsam of the river Rioni at Poti,
<collectingCountry box="[238,331,1303,1329]" name="Georgia" pageId="12" pageNumber="463">Georgia</collectingCountry>
, and mentioned that
<bibRefCitation author="Jackiewicz, M." box="[564,769,1303,1329]" pageId="12" pageNumber="463" pagination="445 - 454" refId="ref11273" refString="Jackiewicz, M. (1974) Krajowe gatunki rodziny Aciculidae (Mollusca, Prosobranchia). Fragmenta Faunistica, 19 (17), 445 - 454. https: // doi. org / 10.3161 / 00159301 FF 1974.19.17.445" type="journal article" year="1974">Jackiewicz (1974)</bibRefCitation>
reported it from North Ossetia in the Caucasus.
<bibRefCitation author="Jackiewicz, M." pageId="12" pageNumber="463" pagination="445 - 454" refId="ref11273" refString="Jackiewicz, M. (1974) Krajowe gatunki rodziny Aciculidae (Mollusca, Prosobranchia). Fragmenta Faunistica, 19 (17), 445 - 454. https: // doi. org / 10.3161 / 00159301 FF 1974.19.17.445" type="journal article" year="1974">Jackiewicz (1974)</bibRefCitation>
, in fact, referred to a paper by
<bibRefCitation author="Kalitina, Z. I. &amp; Botcharova, M. M." box="[584,946,1339,1365]" pageId="12" pageNumber="463" pagination="111 - 112" refId="ref11451" refString="Kalitina, Z. I. &amp; Botcharova, M. M. (1971) Molljuski vypasov i skotoprogonnyh putej kolhozov Severo-Osetinskoj ASSR [Molluscs of pastures and of the roads leading to pastures of collective farms on North Ossetian ASSR]. In: Likharev, I. M. (Ed.), Molluscs. Trends, Methods and some results of their investigation. These and Communications. Academy of Sciences USSR, Zoological Institute, Fourth meeting of the investigation of molluscs, Leningrad, pp. 111 - 112. [in Russian]" type="proceedings paper" year="1971">Kalitina and Botcharova (1971)</bibRefCitation>
, in which they collected molluscs from the mountain part of North Ossetia (= Republic of North-Ossetia-Alaniya). Those authors cited three localities where the land snails were collected (the Ardon and Kurtaty gorges, and the Military-Georgian [= Voenno-Gruzinskaya] Road near Lars settlement). Unfortunately, it is not specified where
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[935,1110,1447,1473]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[935,1110,1447,1473]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
was collected. Furthermore,
<taxonomicName authority="Steklov, 1966" authorityName="Steklov" authorityYear="1966" box="[151,527,1483,1509]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="bakanense">
<emphasis box="[151,362,1483,1509]" italics="true" pageId="12" pageNumber="463">Acicula bakanense</emphasis>
<bibRefCitation author="Steklov, A. A." box="[370,527,1483,1509]" pageId="12" pageNumber="463" pagination="279" refId="ref12757" refString="Steklov, A. A. (1966) Nazemnyye mollyuski neogena Predkavkaz'ya i ikh stratigraficheskoye znacheniye [Terrestrial molluscs of the Neogene Ciscaucasia and their stratigraphic significance]. Academy of Sciences of the USSR Transactions 163. Akademia Nauka S. S. S. R., Moskva, 279." type="journal article" year="1966">Steklov, 1966</bibRefCitation>
</taxonomicName>
, which was treated as a synonym of
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[941,1114,1483,1509]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[941,1114,1483,1509]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
by
<bibRefCitation author="Boeters, H. D. &amp; Gittenberger, E. &amp; Subai, P." box="[1157,1388,1483,1509]" pageId="12" pageNumber="463" pagination="1 - 234" refId="ref10251" refString="Boeters, H. D., Gittenberger, E. &amp; Subai, P. (1989) Die Aciculidae (Mollusca: Gastropoda Prosobranchia). Zoologische Verhandelingen, 252, 1 - 234." type="journal article" year="1989">
Boeters
<emphasis box="[1250,1306,1483,1509]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
(1989)
</bibRefCitation>
and
<bibRefCitation author="Egorov, R. V. &amp; Greie, C." box="[151,416,1519,1545]" pageId="12" pageNumber="463" refId="ref10771" refString="Egorov, R. V. &amp; Greie, C. (2003) Treasure of Russian Shells. Vol. 6. Terrestrial Pectinibranchia: Helicinidae, Cyclophoridae, Diplommatinidae, Cochlostomatidae, Aciculidae, Tateidae, Terrestribythinellidae, Pomatiidae. Colus-Doverie, Moscow, 43 pp." type="book" year="2003">Egorov &amp; Greíe (2003)</bibRefCitation>
, was described from the Mio-Pliocene at Upper Bakanskaya (Krasnodar, North Caucasus). Based on this information,
<bibRefCitation author="Boeters, H. D. &amp; Gittenberger, E. &amp; Subai, P." box="[454,684,1555,1581]" pageId="12" pageNumber="463" pagination="1 - 234" refId="ref10251" refString="Boeters, H. D., Gittenberger, E. &amp; Subai, P. (1989) Die Aciculidae (Mollusca: Gastropoda Prosobranchia). Zoologische Verhandelingen, 252, 1 - 234." type="journal article" year="1989">
Boeters
<emphasis box="[547,603,1555,1581]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
(1989)
</bibRefCitation>
speculated that
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[864,1036,1555,1581]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[864,1036,1555,1581]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
may have a wide distribution in the Caucasus Mountains. The data presented herein partly proves this claim. Namely,
<taxonomicName baseAuthorityName="Clessin" baseAuthorityYear="1911" box="[1081,1254,1591,1617]" class="Gastropoda" family="Aciculidae" genus="Acicula" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="463" phylum="Mollusca" rank="species" species="parcelineata">
<emphasis box="[1081,1254,1591,1617]" italics="true" pageId="12" pageNumber="463">A. parcelineata</emphasis>
</taxonomicName>
was only found in the northern half of the Caucasus Mountains, but the multiple newly reported sites show that this species is widespread and not rare in the area where it occurs.
</paragraph>
</subSubSection>
</treatment>
</document>