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<document id="DC4E7BD88A4B7181873E810DBE5D7E53" ID-CLB-Dataset="4228" ID-DOI="10.12782/sd.20.1.001" ID-GBIF-Dataset="c824e298-6eb3-43bf-8d6a-6dcf6baf691a" ID-ISSN="2189-7301" ID-Zenodo-Dep="4583130" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614892188657" checkinUser="felipe" docAuthor="Motomura, Hiroyuki, Ota, Ryohei, Meguro, Masatoshi &amp; Tashiro, Satokuni" docDate="2015" docId="9F56D34B4953FFA6FC1A3D02B60AFD2F" docLanguage="en" docName="SpeciesDiversity.20.1.1-12.pdf" docOrigin="Species Diversity 20 (1)" docSource="http://dx.doi.org/10.12782/sd.20.1.001" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Enneapterygius atriceps" docType="treatment" docVersion="6" lastPageNumber="11" masterDocId="636FAB33495AFFACFF80383BB224F533" masterDocTitle="Enneapterygius phoenicosoma, a New Species of Triplefin (Tripterygiidae) from the Western Pacific Ocean" masterLastPageNumber="12" masterPageNumber="1" pageNumber="10" updateTime="1698932605625" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="52467F41775AF9CE9471B8E923EBD3A0">Enneapterygius phoenicosoma, a New Species of Triplefin (Tripterygiidae) from the Western Pacific Ocean</mods:title>
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<mods:namePart id="86AA0F0C28D1F31C42C2B6EA40FCA599">Motomura, Hiroyuki</mods:namePart>
<mods:affiliation id="A4B9BD36B497C563C3BB063138F55FF3">The Kagoshima University Museum, 1 - 21 - 30 Korimoto, Kagoshima 890 - 0065, Japan E-mail: motomura @ kaum. kagoshima-u. ac. jp &amp; Corresponding author</mods:affiliation>
<mods:nameIdentifier id="B81C4C5D28509E3435C5D8A5A4C2D6AC" type="email">motomura@kaum.kagoshima-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="5A5AC5084EC4A858A9A418C73F9AD884">Ota, Ryohei</mods:namePart>
<mods:affiliation id="76DD7D258A1D1CC8EDDA80B710CFC40A">Numazu Deepsea Aquarium, 83 Senbonminatomachi, Numazu, Shizuoka 410 - 0845, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="A45AD7520443B20E6C6823F68F179662">Meguro, Masatoshi</mods:namePart>
<mods:affiliation id="75A4B8B92734D2583C08AE97783582BB">Kagoshima Environmental Research and Service, 1 - 1 - 10 Nanatsujima, Kagoshima 891 - 0132, Japan</mods:affiliation>
</mods:name>
<mods:name id="A4495639FB391D7F6A2A3E326479F2C6" type="personal">
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<mods:roleTerm id="2691300B122316AB4E25A983A4701638">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B216B7344BA55A24A19E1EE43456F27D">Tashiro, Satokuni</mods:namePart>
<mods:affiliation id="621D34ABD498029170D3588B9D4B2031">The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24 Korimoto, Kagoshima 890 - 0065, Japan</mods:affiliation>
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<mods:title id="BF799276A64BD37BAF8D5748A1893EF0">Species Diversity</mods:title>
</mods:titleInfo>
<mods:part id="D9CB471764405E44B0B74BC9DFDE5421">
<mods:date id="5EABACE95C361633AFA7D8C2606D4111">2015</mods:date>
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<mods:number id="234AD29CFE58EF01EE574383FDF69DA4">1</mods:number>
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<treatment id="9F56D34B4953FFA6FC1A3D02B60AFD2F" ID-DOI="http://doi.org/10.5281/zenodo.5526683" ID-GBIF-Taxon="188482711" ID-Zenodo-Dep="5526683" LSID="urn:lsid:zoobank.org:act:72378A18-95EE-4CA4-AE58-BA9220905B32" httpUri="http://treatment.plazi.org/id/9F56D34B4953FFA6FC1A3D02B60AFD2F" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="5FE531D64953FFA5FC1A3D02B716F062" box="[922,1330,1337,1361]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="1740625D4953FFA5FC1A3D02B716F062" blockId="9.[893,1359,1337,1426]" box="[922,1330,1337,1361]" pageId="9" pageNumber="10">
<taxonomicName id="D0FF19DE4953FFA5FC1A3D02B716F062" ID-CoL="39VYP" authority="(Jenkins, 1903)" baseAuthorityName="Jenkins" baseAuthorityYear="1903" box="[922,1330,1337,1361]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4953FFA5FC1A3D02B6B6F062" bold="true" box="[922,1170,1337,1361]" italics="true" pageId="9" pageNumber="10">Enneapterygius atriceps</emphasis>
(
<bibRefCitation id="736E1FAC4953FFA5FB1F3D02B70EF062" author="Jenkins, O. P." box="[1183,1322,1337,1361]" pageId="9" pageNumber="10" pagination="415 - 511" refId="ref9683" refString="Jenkins, O. P. 1903. Report on collections of fishes made in the Hawaiian Islands, with descriptions of new species. Bulletin of the U. S. Fish Commission 22 (1902): 415 - 511." type="journal article" year="1903">Jenkins, 1903</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5FE531D64953FFA5FCFD3D62B76BF042" box="[893,1359,1369,1393]" pageId="9" pageNumber="10" type="discussion">
<paragraph id="1740625D4953FFA5FCFD3D62B76BF042" blockId="9.[893,1359,1337,1426]" box="[893,1359,1369,1393]" pageId="9" pageNumber="10">[English name: Hawaiian Blackhead Triplefin]</paragraph>
</subSubSection>
<subSubSection id="5FE531D64953FFA5FC483D41B727F0A1" box="[968,1283,1402,1426]" pageId="9" pageNumber="10" type="discussion">
<paragraph id="1740625D4953FFA5FC483D41B727F0A1" blockId="9.[893,1359,1337,1426]" box="[968,1283,1402,1426]" pageId="9" pageNumber="10">
(
<figureCitation id="8FC47ED84953FFA5FC4F3D41B603F0A1" box="[975,1063,1402,1426]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[821,852,646,668]" captionStartId-1="8.[156,187,1519,1541]" captionTargetBox-0="[823,1428,206,605]" captionTargetBox-1="[177,739,206,1479]" captionTargetId-0="figure-452@4.[822,1430,205,607]" captionTargetId-1="figure-452@8.[176,746,205,1481]" captionTargetPageId-0="4" captionTargetPageId-1="8" captionText-0="Fig. 6. Distributional records of Enneapterygius phoenicosoma n. sp. and E. atriceps. Stars, localities of specimens of E. phoenicosoma; closed circles, localities of specimens of E. atriceps examined in this study; open circles, localities of specimens of E. atriceps examined by Fricke (1997)." captionText-1="Fig. 7. Cephalic sensory pore system of Enneapterygius atriceps. Dorsal (A), lateral (B), and ventral (C) views of head (ANSP 95035, male, 20.8mm SL, Waikiki, Oahu, Hawaiian Islands). Nostrils and tentacles not illustrated; scale bar 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.4583154" figureDoi-1="http://doi.org/10.5281/zenodo.4583166" httpUri-0="https://zenodo.org/record/4583154/files/figure.png" httpUri-1="https://zenodo.org/record/4583166/files/figure.png" pageId="9" pageNumber="10">Figs 6–7</figureCitation>
,
<figureCitation id="8FC47ED84953FFA5FBB13D41B618F0A1" box="[1073,1084,1402,1426]" captionStart="Fig" captionStartId="8.[821,852,893,915]" captionTargetBox="[843,1407,206,852]" captionTargetId="figure-404@8.[841,1411,205,854]" captionTargetPageId="8" captionText="Fig. 8. Tentacle on anterior nostril. A–B, holotype (KAUM–I. 21960, 35.1mm SL) of Enneapterygius phoenicosoma n. sp.; C–D, non-type (ANSP 28022, 18.6mm SL) of E. atriceps. Upper (A, C) and lower (B, D) represent lateral and posterior views respectively. Scale bars 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4583162" httpUri="https://zenodo.org/record/4583162/files/figure.png" pageId="9" pageNumber="10">8</figureCitation>
C–D, 9;
<tableCitation id="5A7D57E64953FFA5FB103D41B6FAF0A1" box="[1168,1246,1402,1426]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="9" pageNumber="10" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Tables 1</tableCitation>
–3)
</paragraph>
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<treatmentCitationGroup id="37EF45734953FFA5FCB53D87B6C5F0C7" pageId="9" pageNumber="10">
<treatmentCitation id="965E444C4953FFA5FCB53D87B6EBF0E0" author="Jenkins, O. P." box="[821,1231,1467,1491]" page="505" pageId="9" pageNumber="10" year="1903">
<taxonomicName id="D0FF19DE4953FFA5FCB53D87B6EBF0E0" authority="Jenkins, 1903: 505" authorityName="Jenkins" authorityPageNumber="505" authorityYear="1903" box="[821,1231,1467,1491]" class="Magnoliopsida" family="Celastraceae" genus="Tripterygium" kingdom="Animalia" order="Celastrales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="atriceps">
<emphasis id="258BBE4F4953FFA5FCB53D87B62FF0E0" box="[821,1035,1468,1491]" italics="true" pageId="9" pageNumber="10">Tripterygium atriceps</emphasis>
<bibRefCitation id="736E1FAC4953FFA5FB933D80B6EBF0E0" author="Jenkins, O. P." box="[1043,1231,1467,1491]" pageId="9" pageNumber="10" pagination="415 - 511" refId="ref9683" refString="Jenkins, O. P. 1903. Report on collections of fishes made in the Hawaiian Islands, with descriptions of new species. Bulletin of the U. S. Fish Commission 22 (1902): 415 - 511." type="journal article" year="1903">Jenkins, 1903: 505</bibRefCitation>
</taxonomicName>
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, fig. 46 (
<typeStatus id="C844DCFF4953FFA5FAA93D80B772F0E0" box="[1321,1366,1467,1491]" pageId="9" pageNumber="10">type</typeStatus>
locality: Honolulu, Oahu, Hawaiian Islands).
</treatmentCitationGroup>
</paragraph>
<paragraph id="1740625D4953FFA5FCB53DC6B77CF3E4" blockId="9.[821,1432,1467,1751]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="37EF45734953FFA5FCB53DC6B77CF3E4" pageId="9" pageNumber="10">
<treatmentCitation id="965E444C4953FFA5FCB53DC6B6C1F327" author="Fricke, R." box="[821,1253,1532,1556]" page="161" pageId="9" pageNumber="10" year="1997">
<taxonomicName id="D0FF19DE4953FFA5FCB53DC6B6C1F327" ID-CoL="39VYP" authority=": Fricke 1997: 161" authorityName=": Fricke" authorityPageNumber="161" authorityYear="1997" box="[821,1253,1532,1556]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4953FFA5FCB53DC6B600F327" box="[821,1060,1533,1556]" italics="true" pageId="9" pageNumber="10">Enneapterygius atriceps</emphasis>
:
<bibRefCitation id="736E1FAC4953FFA5FBB43DC7B6C1F327" author="Fricke, R." box="[1076,1253,1532,1556]" pageId="9" pageNumber="10" refId="ref9457" refString="Fricke, R. 1997. Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein, ix + 607 pp." type="book" year="1997">Fricke 1997: 161</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 566, figs 25, 26 (Hawaiian Archipelago, including French Frigate Shoals, Laysan Island, and
<collectingCountry id="6FE822CD4953FFA5FB9C3E06B68BF366" box="[1052,1199,1597,1621]" name="United States Pacific Island Wildlife Refuges" pageId="9" pageNumber="10">Midway Atoll</collectingCountry>
);
<treatmentCitation id="965E444C4953FFA5FB433E06B1E4F345" author="Williams, J. T. &amp; Fricke, R." page="3534" pageId="9" pageNumber="10" year="2001">
<bibRefCitation id="736E1FAC4953FFA5FB433E06B1E4F345" author="Williams, J. T. &amp; Fricke, R." pageId="9" pageNumber="10" refId="ref10047" refString="Williams, J. T. and Fricke, R. 2001. Tripterygiidae. Triplefins. Pp. 3532 - 3535. In: Carpenter, K. E. and Niem, V. E. (Eds) FAO Species Identification Guide for Fisheries Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 6. Bony Fishes Part 4 (Labridae to Latimeriidae), Estuarine Crocodiles, Sea Turtles, Sea Snakes and Marine Mammals. Food and Agricultural Organization, Rome." type="book" year="2001">Williams and Fricke 2001: 3534</bibRefCitation>
</treatmentCitation>
(name only);
<treatmentCitation id="965E444C4953FFA5FBDC3E65B6DAF345" author="Fricke, R." box="[1116,1278,1630,1654]" page="38" pageId="9" pageNumber="10" year="2009">
<bibRefCitation id="736E1FAC4953FFA5FBDC3E65B6DAF345" author="Fricke, R." box="[1116,1278,1630,1654]" pageId="9" pageNumber="10" refId="ref9486" refString="Fricke, R. 2009. Systematics of the Tripterygiidae (triplefins). Pp. 31 - 67. In: Patzner, R. A., Goncalves, E. J., Hastings, P. A., and Kapoor, B. G. (Eds) The Biology of Blennies. Science Publishers, Enfield." type="book" year="2009">Fricke 2009: 38</bibRefCitation>
</treatmentCitation>
(Hawaiian Islands, north to
<collectingCountry id="6FE822CD4953FFA5FC793E45B6A9F3A5" box="[1017,1165,1662,1686]" name="United States Pacific Island Wildlife Refuges" pageId="9" pageNumber="10">Midway Atoll</collectingCountry>
);
<treatmentCitation id="965E444C4953FFA5FB223E45B740F3A5" author="Randall, J. E." box="[1186,1380,1662,1686]" page="373" pageId="9" pageNumber="10" year="2007">
<bibRefCitation id="736E1FAC4953FFA5FB223E45B740F3A5" author="Randall, J. E." box="[1186,1380,1662,1686]" pageId="9" pageNumber="10" refId="ref9974" refString="Randall, J. E. 2007. Reef and Shore Fishes of the Hawaiian Islands. Sea Grant College Program, University of Hawai' i, Honolulu, xiv + 546 pp." type="book" year="2007">Randall 2007: 373</bibRefCitation>
</treatmentCitation>
, unnumbered figs (Hawaiian Islands); Meguro and Motomura
<emphasis id="258BBE4F4953FFA5FCE83EFBB158F3E4" box="[872,892,1728,1751]" italics="true" pageId="9" pageNumber="10">in</emphasis>
<treatmentCitation id="965E444C4953FFA5FC023EFBB6EBF3E4" author="Motomura, H. &amp; Matsuura, K." box="[898,1231,1727,1751]" page="6" pageId="9" pageNumber="10" year="2010">
<bibRefCitation id="736E1FAC4953FFA5FC023EFBB6EBF3E4" author="Motomura, H. &amp; Matsuura, K." box="[898,1231,1727,1751]" pageId="9" pageNumber="10" refId="ref9871" refString="Motomura, H. and Matsuura, K. (Eds). 2010. Fishes of Yaku-shima Island-A World Heritage Island in the Osumi Group, Kagoshima Prefecture, Southern Japan. National Museum of Nature and Science, Tokyo, viii + 264 pp." type="book" year="2010">Motomura and Matsuura 2010: 6</bibRefCitation>
</treatmentCitation>
(name only).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5FE531D64953FFA6FCCE3F3BB2E2F777" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="1740625D4953FFA5FCCE3F3BB6F9F20A" blockId="9.[821,1432,1792,2077]" pageId="9" pageNumber="10">
<materialsCitation id="A79768004953FFA5FCCE3F3BB6F9F20A" ID-GBIF-Occurrence="3357498357" collectionCode="USNM" collectorName="O. Jenkins." location="Hawaiian Islands" pageId="9" pageNumber="10" specimenCode="USNM 50719" specimenCount="1" typeStatus="holotype">
<emphasis id="258BBE4F4953FFA5FCCE3F3BB192F22B" bold="true" box="[846,950,1792,1816]" pageId="9" pageNumber="10">
<typeStatus id="C844DCFF4953FFA5FCCE3F3BB196F22B" box="[846,946,1792,1816]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="4759CA264953FFA5FC503F3BB67BF22B" box="[976,1119,1792,1816]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM 50719</specimenCode>
, male,
<quantity id="D007CFB84953FFA5FB2A3F3BB724F22B" box="[1194,1280,1792,1816]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="9" pageNumber="10" unit="mm" value="22.1">22.1 mm</quantity>
SL, Honolulu, Oahu,
<location id="122034864953FFA5FCF93F1AB60CF20A" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864953FFA5FCF93F1AB60CF20A" box="[889,1064,1825,1849]" name="Hawaiian Islands" pageId="9" pageNumber="10">Hawaiian Islands</location>
, 1889,
<collectorName id="BA0A078B4953FFA5FBED3F1AB6F9F20A" box="[1133,1245,1825,1849]" pageId="9" pageNumber="10">O. Jenkins.</collectorName>
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</paragraph>
<paragraph id="1740625D4953FFA6FCCE3F79B2E2F777" blockId="9.[821,1432,1792,2077]" lastBlockId="10.[156,767,198,2076]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="258BBE4F4953FFA5FCCE3F79B6BFF26A" bold="true" box="[846,1179,1857,1882]" pageId="9" pageNumber="10">Non-type material examined.</emphasis>
<materialsCitation id="A79768004953FFA5FB343F7AB76CF249" ID-GBIF-Occurrence="3357498339" location="Hawaiian Archipelago" pageId="9" pageNumber="10" specimenCount="46">
<specimenCount id="01F9A9D44953FFA5FB343F7AB76FF26A" box="[1204,1355,1857,1881]" pageId="9" pageNumber="10" type="generic">46 specimens</specimenCount>
,
<quantity id="D007CFB84953FFA5FAE03F79B1AFF24A" metricMagnitude="-2" metricUnit="m" metricValue="2.09" metricValueMax="2.69" metricValueMin="1.49" pageId="9" pageNumber="10" unit="mm" value="20.9" valueMax="26.9" valueMin="14.9">14.9– 26.9 mm</quantity>
SL, all from the
<location id="122034864953FFA5FBDC3F59B76CF249" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864953FFA5FBDC3F59B76CF249" box="[1116,1352,1890,1914]" name="Hawaiian Archipelago" pageId="9" pageNumber="10">Hawaiian Archipelago</location>
</materialsCitation>
.
<materialsCitation id="A79768004953FFA5FAD93F59B1DAF288" ID-GBIF-Occurrence="3357498350" collectionCode="ANSP" county="Hawaiian Islands" location="Honolulu" municipality="Oahu" pageId="9" pageNumber="10" specimenCode="ANSP 28022" specimenCount="3" specimenCount-male="3">
<specimenCode id="4759CA264953FFA5FAD93F59B150F2A8" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9" pageNumber="10">ANSP 28022</specimenCode>
,
<specimenCount id="01F9A9D44953FFA5FCFF3FB8B1EBF2A9" box="[895,975,1922,1947]" pageId="9" pageNumber="10" type="male">3 males</specimenCount>
,
<quantity id="D007CFB84953FFA5FC5B3FB9B64DF2A9" box="[987,1129,1922,1946]" metricMagnitude="-2" metricUnit="m" metricValue="1.7549999999999997" metricValueMax="1.94" metricValueMin="1.5699999999999998" pageId="9" pageNumber="10" unit="mm" value="17.549999999999997" valueMax="19.4" valueMin="15.7">15.7–19.4mm</quantity>
SL,
<location id="122034864953FFA5FB193FB9B6D8F2A9" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864953FFA5FB193FB9B6D8F2A9" box="[1177,1276,1922,1946]" county="Hawaiian Islands" municipality="Oahu" name="Honolulu" pageId="9" pageNumber="10">Honolulu</location>
,
<collectingMunicipality id="F724F8274953FFA5FA883FB9B767F2A9" box="[1288,1347,1922,1946]" pageId="9" pageNumber="10">Oahu</collectingMunicipality>
,
<collectingCounty id="FE211AD14953FFA5FACE3FB9B187F288" pageId="9" pageNumber="10">Hawaiian Islands</collectingCounty>
, 1900, O
</materialsCitation>
.
<materialsCitation id="A79768004953FFA5FB873F98B7B0F2EF" ID-GBIF-Occurrence="3357498351" collectionCode="ANSP" county="Hawaiian Islands" location="Waikiki" municipality="Oahu" pageId="9" pageNumber="10" specimenCode="ANSP 95035" specimenCount="1">
Jenkins;
<specimenCode id="4759CA264953FFA5FBDE3F98B6C6F288" box="[1118,1250,1955,1979]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9" pageNumber="10">ANSP 95035</specimenCode>
, male and female,
<quantity id="D007CFB84953FFA5FCB53FFFB1E6F2E8" box="[821,962,1988,2012]" metricMagnitude="-2" metricUnit="m" metricValue="1.785" metricValueMax="2.08" metricValueMin="1.49" pageId="9" pageNumber="10" unit="mm" value="17.85" valueMax="20.8" valueMin="14.9">14.9–20.8 mm</quantity>
SL,
<location id="122034864953FFA5FC6E3FF8B61BF2E8" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864953FFA5FC6E3FF8B61BF2E8" box="[1006,1087,1987,2011]" county="Hawaiian Islands" municipality="Oahu" name="Waikiki" pageId="9" pageNumber="10">Waikiki</location>
,
<collectingMunicipality id="F724F8274953FFA5FBC83FF8B6A7F2E8" box="[1096,1155,1987,2011]" pageId="9" pageNumber="10">Oahu</collectingMunicipality>
,
<collectingCounty id="FE211AD14953FFA5FB0C3FF8B71FF2E8" box="[1164,1339,1987,2011]" pageId="9" pageNumber="10">Hawaiian Islands</collectingCounty>
, 1992, C
</materialsCitation>
.
<materialsCitation id="A79768004953FFA5FCB53FDFB613FD2F" ID-GBIF-Occurrence="3357498340" collectionCode="V" county="Oahu" location="Waikiki Reef" municipality="Honolulu" pageId="9" pageNumber="10" specimenCode="BPBM 3465" specimenCount="1">
Edmondson;
<specimenCode id="4759CA264953FFA5FC3F3FDEB618F2CF" box="[959,1084,2020,2044]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="10" type="Museum">BPBM 3465</specimenCode>
, male,
<quantity id="D007CFB84953FFA5FB073FDFB6F9F2CF" box="[1159,1245,2020,2044]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="9" pageNumber="10" unit="mm" value="25.8">25.8 mm</quantity>
SL,
<location id="122034864953FFA5FA8E3FDFB7B0F2CF" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864953FFA5FA8E3FDFB7B0F2CF" box="[1294,1428,2020,2044]" county="Oahu" municipality="Honolulu" name="Waikiki Reef" pageId="9" pageNumber="10">Waikiki Reef</location>
,
<collectingMunicipality id="F724F8274953FFA5FCB5303EB1BCFD2E" box="[821,920,2053,2077]" pageId="9" pageNumber="10">Honolulu</collectingMunicipality>
,
<collectingCounty id="FE211AD14953FFA5FC23303EB1FAFD2E" box="[931,990,2053,2077]" pageId="9" pageNumber="10">Oahu</collectingCounty>
, 1923,
<collectionCode id="71EEFA984953FFA5FBA6303EB613FD2F" box="[1062,1079,2053,2076]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="10" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="A79768004953FFA6FBC1303EB346F5CD" ID-GBIF-Occurrence="3357498309" collectingDate="1975-09-13" collectionCode="BPBM" collectorName="Pietschmann" elevation="2" lastPageId="10" lastPageNumber="11" location="Kaneohe Bay" municipality="Oahu" pageId="9" pageNumber="10" specimenCode="BPBM 19658" specimenCount="1">
<collectorName id="BA0A078B4953FFA5FBC1303EB6E3FD2E" box="[1089,1223,2053,2077]" pageId="9" pageNumber="10">Pietschmann</collectorName>
;
<specimenCode id="4759CA264953FFA5FB53303EB77FFD2F" box="[1235,1371,2052,2076]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="10" type="Museum">BPBM 19658</specimenCode>
, male and female, 19.0–
<quantity id="D007CFB84950FFA6FED238FDB38EF5ED" box="[338,426,198,222]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="10" pageNumber="11" unit="mm" value="20.9">20.9 mm</quantity>
SL,
<location id="122034864950FFA6FE5A38FDB046F5EC" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864950FFA6FE5A38FDB046F5EC" box="[474,610,198,223]" municipality="Oahu" name="Kaneohe Bay" pageId="10" pageNumber="11">Kaneohe Bay</location>
,
<collectingMunicipality id="F724F8274950FFA6FDED38FDB08CF5ED" box="[621,680,198,222]" pageId="10" pageNumber="11">Oahu</collectingMunicipality>
,
<quantity id="D007CFB84950FFA6FD3438FDB0FEF5ED" box="[692,730,198,222]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2.0">
<elevation id="9CD2856E4950FFA6FD3438FDB0FEF5ED" box="[692,730,198,222]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2.0">2 m</elevation>
</quantity>
,
<date id="6341449D4950FFA6FD6738FDB361F5CC" pageId="10" pageNumber="11" value="1975-09-13">
<collectingDate id="7305BD754950FFA6FD6738FDB361F5CC" pageId="10" pageNumber="11" value="1975-09-13">13 September 1975</collectingDate>
</date>
, B
</materialsCitation>
. Carlson
<emphasis id="258BBE4F4950FFA6FE4838D3B3D9F5CC" box="[456,509,231,255]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
;
<materialsCitation id="A79768004950FFA6FD8B38DCB0ADF40C" ID-GBIF-Occurrence="3357498346" collectingDate="1991-09-16" collectionCode="BPBM" country="United States Pacific Island Wildlife Refuges" elevation="8" location="Sand Island" pageId="10" pageNumber="11" specimenCode="BPBM 34846" specimenCount="18" specimenCount-female="5" specimenCount-male="13">
<specimenCode id="4759CA264950FFA6FD8B38DCB0B2F5CC" box="[523,662,231,255]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="10" pageNumber="11" type="Museum">BPBM 34846</specimenCode>
,
<specimenCount id="01F9A9D44950FFA6FD2338DCB0DBF5CC" box="[675,767,231,255]" pageId="10" pageNumber="11" type="male">13 males</specimenCount>
and
<specimenCount id="01F9A9D44950FFA6FF4A393CB309F42C" box="[202,301,263,287]" pageId="10" pageNumber="11" type="female">5 females</specimenCount>
,
<quantity id="D007CFB84950FFA6FEB9393CB3E3F42C" box="[313,455,263,287]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" metricValueMax="2.3899999999999997" metricValueMin="1.95" pageId="10" pageNumber="11" unit="mm" value="21.7" valueMax="23.9" valueMin="19.5">19.5–23.9 mm</quantity>
SL, southeast side of
<location id="122034864950FFA6FD2A393CB2EEF473" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864950FFA6FD2A393CB2EEF473" country="United States Pacific Island Wildlife Refuges" name="Sand Island" pageId="10" pageNumber="11">Sand Island</location>
,
<collectingCountry id="6FE822CD4950FFA6FF583913B343F473" box="[216,359,296,320]" name="United States Pacific Island Wildlife Refuges" pageId="10" pageNumber="11">Midway Atoll</collectingCountry>
,
<quantity id="D007CFB84950FFA6FEF43913B3BDF40C" box="[372,409,296,320]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="m" value="8.0">
<elevation id="9CD2856E4950FFA6FEF43913B3BDF40C" box="[372,409,296,320]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="m" value="8.0">8 m</elevation>
</quantity>
,
<date id="6341449D4950FFA6FE273913B055F473" box="[423,625,296,320]" pageId="10" pageNumber="11" value="1991-09-16">
<collectingDate id="7305BD754950FFA6FE273913B055F473" box="[423,625,296,320]" pageId="10" pageNumber="11" value="1991-09-16">16 September 1991</collectingDate>
</date>
, J
</materialsCitation>
. Randall
<emphasis id="258BBE4F4950FFA6FD6D3912B291F453" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
;
<materialsCitation id="A79768004950FFA6FF433972B3A1F4F1" ID-GBIF-Occurrence="3357498337" collectingDate="1961-04-21" collectionCode="FMNH" collectorName="L. Woods et al." country="United States Pacific Island Wildlife Refuges" county="Oahu" location="Kaneohe Bay" municipality="Kahuku" pageId="10" pageNumber="11" specimenCode="FMNH 47658, FMNH 63605" specimenCount="14" specimenCount-female="5" specimenCount-male="9">
<specimenCode id="4759CA264950FFA6FF433972B372F453" box="[195,342,328,352]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11" type="Museum">FMNH 47658</specimenCode>
, male,
<quantity id="D007CFB84950FFA6FE233972B3DDF453" box="[419,505,329,353]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="10" pageNumber="11" unit="mm" value="22.1">22.1 mm</quantity>
SL,
<location id="122034864950FFA6FDAB3973B090F452" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864950FFA6FDAB3973B090F452" box="[555,692,328,353]" country="United States Pacific Island Wildlife Refuges" county="Oahu" municipality="Kahuku" name="Kaneohe Bay" pageId="10" pageNumber="11">Kaneohe Bay</location>
,
<collectorName id="BA0A078B4950FFA6FD403973B0DFF453" box="[704,763,328,352]" pageId="10" pageNumber="11">Oahu</collectorName>
,
<collectorName id="BA0A078B4950FFA6FF1C3952B32EF4B2" box="[156,266,361,385]" pageId="10" pageNumber="11">April–May</collectorName>
1935,
<collectorName id="BA0A078B4950FFA6FED23952B020F4B2" box="[338,516,361,385]" pageId="10" pageNumber="11">Shedd Aquarium</collectorName>
;
<specimenCode id="4759CA264950FFA6FD913951B087F4B2" box="[529,675,361,385]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11" type="Museum">FMNH 63605</specimenCode>
,
<specimenCount id="01F9A9D44950FFA6FD2F3952B0DBF4B2" box="[687,767,361,385]" pageId="10" pageNumber="11" type="male">9 males</specimenCount>
and
<specimenCount id="01F9A9D44950FFA6FF4C39B2B315F492" box="[204,305,393,417]" pageId="10" pageNumber="11" type="female">5 females</specimenCount>
, 17.0–
<quantity id="D007CFB84950FFA6FEF539B2B3E8F492" box="[373,460,393,417]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="10" pageNumber="11" unit="mm" value="26.9">26.9 mm</quantity>
SL,
<collectingMunicipality id="F724F8274950FFA6FD8039B2B076F492" box="[512,594,393,417]" pageId="10" pageNumber="11">Kahuku</collectingMunicipality>
,
<collectingCounty id="FE211AD14950FFA6FDE039B2B0BFF492" box="[608,667,393,417]" pageId="10" pageNumber="11">Oahu</collectingCounty>
,
<date id="6341449D4950FFA6FD2839B1B2EAF4F1" pageId="10" pageNumber="11" value="1961-04-21">
<collectingDate id="7305BD754950FFA6FD2839B1B2EAF4F1" pageId="10" pageNumber="11" value="1961-04-21">21 April 1961</collectingDate>
</date>
, L. Woods
<emphasis id="258BBE4F4950FFA6FECB3990B3A4F4F1" box="[331,384,426,450]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
;
</materialsCitation>
<materialsCitation id="A79768004950FFA6FE0E3991B31FF730" ID-GBIF-Occurrence="3357498326" accessionNumber="ROM 28022" collectionCode="ROM, US" country="United States Pacific Island Wildlife Refuges" county="Fish" location="Honolulu" municipality="Oahu" pageId="10" pageNumber="11" specimenCount="3" specimenCount-male="3">
<collectionCode id="71EEFA984950FFA6FE0E3991B3E3F4F1" box="[398,455,426,450]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="10" pageNumber="11" type="Museum">ROM</collectionCode>
28022,
<specimenCount id="01F9A9D44950FFA6FD9D3991B049F4F1" box="[541,621,426,450]" pageId="10" pageNumber="11" type="male">3 males</specimenCount>
(one in poor condition, its SL not included), 16.0–
<quantity id="D007CFB84950FFA6FD9639F1B048F4D1" box="[534,620,458,482]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="10" pageNumber="11" unit="mm" value="18.5">18.5 mm</quantity>
SL,
<location id="122034864950FFA6FD1839F0B0DFF4D0" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864950FFA6FD1839F0B0DFF4D0" box="[664,763,459,483]" country="United States Pacific Island Wildlife Refuges" county="Fish" municipality="Oahu" name="Honolulu" pageId="10" pageNumber="11">Honolulu</location>
,
<collectingMunicipality id="F724F8274950FFA6FF1C39D0B2F3F730" box="[156,215,491,515]" pageId="10" pageNumber="11">Oahu</collectingMunicipality>
,
<collectionCode id="71EEFA984950FFA6FF6439D0B327F730" box="[228,259,491,515]" country="South Africa" name="University of Stellenbosch" pageId="10" pageNumber="11">US</collectionCode>
<collectingCounty id="FE211AD14950FFA6FE8D39D0B31FF730" box="[269,315,491,515]" pageId="10" pageNumber="11">Fish</collectingCounty>
</materialsCitation>
.
<materialsCitation id="A79768004950FFA6FEC739D0B0B4F717" ID-GBIF-Occurrence="3357498311" accessionNumber="ROM 95035" collectionCode="ROM" collectorName="Commission" country="United States Pacific Island Wildlife Refuges" county="Oahu" location="Waikiki" municipality="Honolulu" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2">
<collectorName id="BA0A078B4950FFA6FEC739D0B3EFF730" box="[327,459,491,515]" pageId="10" pageNumber="11">Commission</collectorName>
;
<collectionCode id="71EEFA984950FFA6FE5939D0B036F730" box="[473,530,491,515]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="10" pageNumber="11" type="Museum">ROM</collectionCode>
95035,
<specimenCount id="01F9A9D44950FFA6FDE739D0B09DF730" box="[615,697,491,515]" pageId="10" pageNumber="11" type="male">2 males</specimenCount>
,
<quantity id="D007CFB84950FFA6FD4739D0B2D6F710" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.04" metricValueMin="1.56" pageId="10" pageNumber="11" unit="mm" value="18.0" valueMax="20.4" valueMin="15.6">15.6– 20.4 mm</quantity>
SL,
<location id="122034864950FFA6FEA63A37B353F717" LSID="urn:lsid:plazi:treatment:9F56D34B4953FFA6FC1A3D02B60AFD2F:122034864950FFA6FEA63A37B353F717" box="[294,375,524,548]" country="United States Pacific Island Wildlife Refuges" county="Oahu" municipality="Honolulu" name="Waikiki" pageId="10" pageNumber="11">Waikiki</location>
,
<collectingMunicipality id="F724F8274950FFA6FE043A37B3C3F717" box="[388,487,524,548]" pageId="10" pageNumber="11">Honolulu</collectingMunicipality>
,
<collectingCounty id="FE211AD14950FFA6FE753A37B014F717" box="[501,560,524,548]" pageId="10" pageNumber="11">Oahu</collectingCounty>
, 1922, C
</materialsCitation>
. Edmondson.
</paragraph>
</subSubSection>
<subSubSection id="5FE531D64950FFA6FF353A77B0DCF0C0" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="1740625D4950FFA6FF353A77B0DCF0C0" blockId="10.[156,767,198,2076]" pageId="10" pageNumber="11">
<emphasis id="258BBE4F4950FFA6FF353A77B301F757" bold="true" box="[181,293,588,612]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
A small-sized species of
<taxonomicName id="D0FF19DE4950FFA6FDE63A76B0DAF757" authorityName="Ruppell" authorityYear="1835" box="[614,766,589,612]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="258BBE4F4950FFA6FDE63A76B0DAF757" box="[614,766,589,612]" italics="true" pageId="10" pageNumber="11">Enneapterygius</emphasis>
</taxonomicName>
with the following combination of characters: 12–15 (mode 14) second dorsal-fin spines; 9–11 (9) third dorsal-fin soft rays; 14–17 (15) pectoral-fin rays; 19–21 (19) anal-fin soft rays; 35–37 (36) scale rows in longitudinal series; 17–20 (19) pored lateral-line scales; 16–20 (18) notched lateral-line scales; 1 or 2 scale rows between last pored lateral-line scale and first notched lateral-line scale; mandibular pore formula 4–6+1+4–5 (4+1+4); short dorsal-fin spines and rays [
<emphasis id="258BBE4F4950FFA6FF253B49B2E2F6BA" box="[165,198,882,905]" italics="true" pageId="10" pageNumber="11">e.g.</emphasis>
, lengths of second spine 8.0–11.3% of SL (mean 9.2%) and third spine 6.5–9.1% of SL (7.8%) in first dorsal fin; first spine 12.5–14.2% of SL (13.4%), second spine 12.9–15.3% of SL (14.2%), and third spine 13.6–15.8% of SL (14.8%) in second dorsal fin; first soft ray 14.2–17.3% of SL (15.9%), second soft ray 13.6–16.7% of SL (15.6%), and third soft ray 13.4–16.2% of SL (14.8%) in third dorsal fin]; long second dorsal-fin base [27.4–38.1% of SL (mean 31.8%)]; long anal-fin base [39.5–47.5% of SL (mean 43.1%)]; nasal tentacle bilobed (
<figureCitation id="8FC47ED84950FFA6FEAE3CADB3B8F19D" box="[302,412,1174,1198]" captionStart="Fig" captionStartId="8.[821,852,893,915]" captionTargetBox="[843,1407,206,852]" captionTargetId="figure-404@8.[841,1411,205,854]" captionTargetPageId="8" captionText="Fig. 8. Tentacle on anterior nostril. A–B, holotype (KAUM–I. 21960, 35.1mm SL) of Enneapterygius phoenicosoma n. sp.; C–D, non-type (ANSP 28022, 18.6mm SL) of E. atriceps. Upper (A, C) and lower (B, D) represent lateral and posterior views respectively. Scale bars 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4583162" httpUri="https://zenodo.org/record/4583162/files/figure.png" pageId="10" pageNumber="11">Fig. 8C, D</figureCitation>
; rarely unbranched; 2 of
<specimenCount id="01F9A9D44950FFA6FD233CADB2F6F1FD" pageId="10" pageNumber="11" type="generic">47 specimens</specimenCount>
with unbranched tentacle); lower two-thirds of head, including lower half of eye, snout, lips, cheek, and opercle, and pectoral-fin base dense black in nuptial males; upper head and body reddish-brown in females and males, reddish in nuptial males, with 4–6 H-shaped and/or upside-down Y-shaped oblique brown bars on lateral surface of body; caudal-fin base with vertical brownish band, its width subequal to pupil diameter, narrow vertical white band behind brownish band; narrow, indistinct, white line along posterior margin of black area on pectoral-fin base; no fins black.
</paragraph>
</subSubSection>
<subSubSection id="5FE531D64950FFA6FF353DC7B3FBF306" pageId="10" pageNumber="11" type="distribution">
<paragraph id="1740625D4950FFA6FF353DC7B3FBF306" blockId="10.[156,767,198,2076]" pageId="10" pageNumber="11">
<emphasis id="258BBE4F4950FFA6FF353DC7B365F327" bold="true" box="[181,321,1532,1556]" pageId="10" pageNumber="11">Distribution.</emphasis>
Known only from the Hawaiian Archipelago (
<bibRefCitation id="736E1FAC4950FFA6FF6B3E26B340F306" author="Fricke, R." box="[235,356,1565,1589]" pageId="10" pageNumber="11" refId="ref9457" refString="Fricke, R. 1997. Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein, ix + 607 pp." type="book" year="1997">Fricke 1997</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
<subSubSection id="5FE531D64950FFA6FF353E06B60AFD2F" pageId="10" pageNumber="11" type="discussion">
<paragraph id="1740625D4950FFA6FF353E06B0B0F3E4" blockId="10.[156,767,198,2076]" pageId="10" pageNumber="11">
<emphasis id="258BBE4F4950FFA6FF353E06B33DF366" bold="true" box="[181,281,1597,1621]" pageId="10" pageNumber="11">Remarks.</emphasis>
Counts and measurements of the above-listed
<typeStatus id="C844DCFF4950FFA6FF3B3E65B330F345" box="[187,276,1630,1654]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
and non-types of
<taxonomicName id="D0FF19DE4950FFA6FE513E65B01DF346" authorityName=": Fricke" authorityYear="1997" box="[465,569,1630,1653]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FE513E65B01DF346" box="[465,569,1630,1653]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
are given in
<tableCitation id="5A7D57E64950FFA6FD403E65B282F3A5" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Tables 1</tableCitation>
–3. The cephalic sensory pore system of the species is illustrated in
<figureCitation id="8FC47ED84950FFA6FE9C3EA4B37EF385" box="[284,346,1695,1719]" captionStart="Fig" captionStartId="8.[156,187,1519,1541]" captionTargetBox="[177,739,206,1479]" captionTargetId="figure-452@8.[176,746,205,1481]" captionTargetPageId="8" captionText="Fig. 7. Cephalic sensory pore system of Enneapterygius atriceps. Dorsal (A), lateral (B), and ventral (C) views of head (ANSP 95035, male, 20.8mm SL, Waikiki, Oahu, Hawaiian Islands). Nostrils and tentacles not illustrated; scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4583166" httpUri="https://zenodo.org/record/4583166/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
. Color photographs of male and female specimens were published by
<bibRefCitation id="736E1FAC4950FFA6FE473E84B0B4F3E4" author="Randall, J. E." box="[455,656,1727,1751]" pageId="10" pageNumber="11" refId="ref9974" refString="Randall, J. E. 2007. Reef and Shore Fishes of the Hawaiian Islands. Sea Grant College Program, University of Hawai' i, Honolulu, xiv + 546 pp." type="book" year="2007">Randall (2007: 373)</bibRefCitation>
.
</paragraph>
<paragraph id="1740625D4950FFA6FF353EE4B182F4D1" blockId="10.[156,767,198,2076]" lastBlockId="10.[820,1432,198,2076]" pageId="10" pageNumber="11">
<emphasis id="258BBE4F4950FFA6FF353EE4B36EF3C4" bold="true" box="[181,330,1759,1783]" pageId="10" pageNumber="11">Comparisons.</emphasis>
<taxonomicName id="D0FF19DE4950FFA6FEE33EDBB0B2F3CB" authority="Motomura &amp; Ota &amp; Meguro &amp; Tashiro, 2015" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[355,662,1760,1784]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma" status="sp. nov.">
<emphasis id="258BBE4F4950FFA6FEE33EDBB0B2F3CB" box="[355,662,1760,1784]" italics="true" pageId="10" pageNumber="11">Enneapterygius phoenicosoma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EB803344950FFA6FD243EDBB0C6F3C4" box="[676,738,1760,1783]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
is similar to the Hawaiian endemic species
<taxonomicName id="D0FF19DE4950FFA6FDC13F3AB08CF22B" authorityName=": Fricke" authorityYear="1997" box="[577,680,1793,1816]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FDC13F3AB08CF22B" box="[577,680,1793,1816]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
in sharing the following features: 16–20 pored lateral-line scales; a single scale row between the last pored lateral-line scale and the first notched lateral-line scale; the lower two-thirds of the head, including the lower half of the eye, snout, lips, cheek, and opercle, and the pectoral-fin base dense black in nuptial males; the upper head and body reddish in nuptial males but reddish-brown in females and non-nuptial males, with H-shaped or upside-down Y-shaped oblique brown bars on the lateral surface of the body; the caudal-fin base with a vertical brownish band, its width subequal to the pupil diameter, and a vertical narrow white band behind the brownish band; a narrow, indistinct, white line along the posterior margin of the pectoral-fin base; and no black fins. With the exception of
<taxonomicName id="D0FF19DE4950FFA6FBA33952B6E8F4B2" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[1059,1228,361,385]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FBA33952B6E8F4B2" box="[1059,1228,361,385]" italics="true" pageId="10" pageNumber="11">E. phoenicosoma</emphasis>
</taxonomicName>
and
<taxonomicName id="D0FF19DE4950FFA6FA853952B74AF4B3" authorityName=": Fricke" authorityYear="1997" box="[1285,1390,361,384]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FA853952B74AF4B3" box="[1285,1390,361,384]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
, no nominal species with combination of the above-mentioned characters have been reported in the genus (
<bibRefCitation id="736E1FAC4950FFA6FA953991B7B0F4F1" author="Fricke, R." box="[1301,1428,426,450]" pageId="10" pageNumber="11" refId="ref9457" refString="Fricke, R. 1997. Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein, ix + 607 pp." type="book" year="1997">Fricke 1997</bibRefCitation>
; this study).
</paragraph>
<paragraph id="1740625D4950FFA6FCCE39D0B188F61B" blockId="10.[820,1432,198,2076]" pageId="10" pageNumber="11">
The new species is clearly distinguished from
<taxonomicName id="D0FF19DE4950FFA6FAAF39D7B7B3F730" authorityName=": Fricke" authorityYear="1997" box="[1327,1431,492,515]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FAAF39D7B7B3F730" box="[1327,1431,492,515]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
by having a simple nasal tentacle (
<emphasis id="258BBE4F4950FFA6FB223A37B692F710" box="[1186,1206,524,547]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
bilobed in the latter;
<figureCitation id="8FC47ED84950FFA6FCB43A17B156F777" box="[820,882,556,580]" captionStart="Fig" captionStartId="8.[821,852,893,915]" captionTargetBox="[843,1407,206,852]" captionTargetId="figure-404@8.[841,1411,205,854]" captionTargetPageId="8" captionText="Fig. 8. Tentacle on anterior nostril. A–B, holotype (KAUM–I. 21960, 35.1mm SL) of Enneapterygius phoenicosoma n. sp.; C–D, non-type (ANSP 28022, 18.6mm SL) of E. atriceps. Upper (A, C) and lower (B, D) represent lateral and posterior views respectively. Scale bars 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4583162" httpUri="https://zenodo.org/record/4583162/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
) and 16–18 (mode 17) anal-fin soft rays [
<emphasis id="258BBE4F4950FFA6FABB3A16B76BF777" box="[1339,1359,557,580]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
19–21 (19);
<tableCitation id="5A7D57E64950FFA6FCF53A77B1CAF756" box="[885,1006,588,613]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[156,204,507,530]" captionStartId-1="6.[156,204,1241,1264]" captionTargetId-1="graphics-294@6.[156,2041,1516,1517]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." captionText-1="Table 2. Frequency distribution of selected fin ray counts of Enneapterygius phoenicosoma n. sp. and E. atriceps." httpUri-0="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" httpUri-1="http://table.plazi.org/id/438032D5495CFFAAFF1C3CE2B6BFF1C3" pageId="10" pageNumber="11" tableUuid-0="438032D5495FFFA9FF1C39C0B7F0F721" tableUuid-1="438032D5495CFFAAFF1C3CE2B6BFF1C3">Tables 1–2</tableCitation>
].
<taxonomicName id="D0FF19DE4950FFA6FB893A76B71AF757" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[1033,1342,588,612]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FB893A76B71AF757" box="[1033,1342,588,612]" italics="true" pageId="10" pageNumber="11">Enneapterygius phoenicosoma</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="D0FF19DE4950FFA6FC383A55B605F7B6" authorityName=": Fricke" authorityYear="1997" box="[952,1057,622,645]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FC383A55B605F7B6" box="[952,1057,622,645]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
in having 11–13 (mode 12) second dorsal-fin spines [
<emphasis id="258BBE4F4950FFA6FC763AB5B62EF796" box="[1014,1034,654,677]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
12–15 (14);
<tableCitation id="5A7D57E64950FFA6FB1A3AB6B729F795" box="[1178,1293,653,678]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[156,204,507,530]" captionStartId-1="6.[156,204,1241,1264]" captionTargetId-1="graphics-294@6.[156,2041,1516,1517]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." captionText-1="Table 2. Frequency distribution of selected fin ray counts of Enneapterygius phoenicosoma n. sp. and E. atriceps." httpUri-0="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" httpUri-1="http://table.plazi.org/id/438032D5495CFFAAFF1C3CE2B6BFF1C3" pageId="10" pageNumber="11" tableUuid-0="438032D5495FFFA9FF1C39C0B7F0F721" tableUuid-1="438032D5495CFFAAFF1C3CE2B6BFF1C3">Tables 1–2</tableCitation>
], 15–17 (16) pectoral-fin rays [
<emphasis id="258BBE4F4950FFA6FC753A94B62DF7F5" box="[1013,1033,687,710]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
14–17 (15);
<tableCitation id="5A7D57E64950FFA6FB193A95B729F7F5" box="[1177,1293,686,710]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[156,204,507,530]" captionStartId-1="6.[156,204,1241,1264]" captionTargetId-1="graphics-294@6.[156,2041,1516,1517]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." captionText-1="Table 2. Frequency distribution of selected fin ray counts of Enneapterygius phoenicosoma n. sp. and E. atriceps." httpUri-0="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" httpUri-1="http://table.plazi.org/id/438032D5495CFFAAFF1C3CE2B6BFF1C3" pageId="10" pageNumber="11" tableUuid-0="438032D5495FFFA9FF1C39C0B7F0F721" tableUuid-1="438032D5495CFFAAFF1C3CE2B6BFF1C3">Tables 1–2</tableCitation>
], 32–35 (33) scale rows in the longitudinal series [
<emphasis id="258BBE4F4950FFA6FB3C3AF4B6F4F7D5" box="[1212,1232,719,742]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
35–37 (36);
<tableCitation id="5A7D57E64950FFA6FAD83AF5B164F634" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Tables 1</tableCitation>
, 3], and 16–19 (17) pored lateral-line scales [
<emphasis id="258BBE4F4950FFA6FA8A3ACBB73AF634" box="[1290,1310,752,775]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
17–20 (19);
<tableCitation id="5A7D57E64950FFA6FCB43B2BB1A2F61B" box="[820,902,784,808]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Tables 1</tableCitation>
, 3].
</paragraph>
<paragraph id="1740625D4950FFA6FCCE3B0AB6CBF023" blockId="10.[820,1432,198,2076]" pageId="10" pageNumber="11">
In morphometrics,
<taxonomicName id="D0FF19DE4950FFA6FB943B0AB69FF67B" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[1044,1211,816,840]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FB943B0AB69FF67B" box="[1044,1211,816,840]" italics="true" pageId="10" pageNumber="11">E. phoenicosoma</emphasis>
</taxonomicName>
tends to have slightly longer spines and rays of the dorsal fins than
<taxonomicName id="D0FF19DE4950FFA6FB7F3B6AB741F65B" authorityName=": Fricke" authorityYear="1997" box="[1279,1381,849,872]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FB7F3B6AB741F65B" box="[1279,1381,849,872]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
:
<emphasis id="258BBE4F4950FFA6FAF13B6AB7B6F65B" box="[1393,1426,849,872]" italics="true" pageId="10" pageNumber="11">e.g.</emphasis>
, lengths of the second spine (mean 10.1% of SL
<emphasis id="258BBE4F4950FFA6FAAC3B49B764F6BA" box="[1324,1344,882,905]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
9.2% in
<taxonomicName id="D0FF19DE4950FFA6FCB43BA9B1B8F69A" authorityName=": Fricke" authorityYear="1997" box="[820,924,914,937]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FCB43BA9B1B8F69A" box="[820,924,914,937]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
;
<tableCitation id="5A7D57E64950FFA6FC2A3BA9B1D0F699" box="[938,1012,914,938]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
,
<figureCitation id="8FC47ED84950FFA6FB823BA9B66AF699" box="[1026,1102,914,938]" captionStart="Fig" captionStartId="9.[157,188,1228,1250]" captionTargetBox="[188,1397,207,1187]" captionTargetId="figure-380@9.[187,1399,205,1189]" captionTargetPageId="9" captionText="Fig. 9. Comparisons of lengths of (A) second spine of first dorsal fin, (B) first spine of second dorsal fin, (C) second spine of second dorsal fin, (D) third spine of second dorsal fin, (E) second dorsal-fin base, and (F) anal-fin base (% standard length) with standard length in Enneapterygius phoenicosoma n. sp. (stars) and E. atriceps (circles)." figureDoi="http://doi.org/10.5281/zenodo.4583170" httpUri="https://zenodo.org/record/4583170/files/figure.png" pageId="10" pageNumber="11">Fig. 9A</figureCitation>
) and third spine (8.5%
<emphasis id="258BBE4F4950FFA6FAC53BA9B77DF69A" box="[1349,1369,914,937]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
7.8%;
<tableCitation id="5A7D57E64950FFA6FCB43B89B1A5F6F9" box="[820,897,946,970]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
) in the first dorsal fin; lengths of the first spine (15.2%
<emphasis id="258BBE4F4950FFA6FC033BE8B1B3F6D9" box="[899,919,979,1002]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
13.4%;
<tableCitation id="5A7D57E64950FFA6FC6C3BE8B613F6D8" box="[1004,1079,979,1003]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
,
<figureCitation id="8FC47ED84950FFA6FBC63BE8B6B6F6D8" box="[1094,1170,979,1003]" captionStart="Fig" captionStartId="9.[157,188,1228,1250]" captionTargetBox="[188,1397,207,1187]" captionTargetId="figure-380@9.[187,1399,205,1189]" captionTargetPageId="9" captionText="Fig. 9. Comparisons of lengths of (A) second spine of first dorsal fin, (B) first spine of second dorsal fin, (C) second spine of second dorsal fin, (D) third spine of second dorsal fin, (E) second dorsal-fin base, and (F) anal-fin base (% standard length) with standard length in Enneapterygius phoenicosoma n. sp. (stars) and E. atriceps (circles)." figureDoi="http://doi.org/10.5281/zenodo.4583170" httpUri="https://zenodo.org/record/4583170/files/figure.png" pageId="10" pageNumber="11">Fig. 9B</figureCitation>
), second spine (15.9%
<emphasis id="258BBE4F4950FFA6FA043BE8B7BCF6D9" box="[1412,1432,979,1002]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
14.2%;
<tableCitation id="5A7D57E64950FFA6FC013BC8B1E8F138" box="[897,972,1011,1035]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
,
<figureCitation id="8FC47ED84950FFA6FC5B3BC8B60DF138" box="[987,1065,1011,1035]" captionStart="Fig" captionStartId="9.[157,188,1228,1250]" captionTargetBox="[188,1397,207,1187]" captionTargetId="figure-380@9.[187,1399,205,1189]" captionTargetPageId="9" captionText="Fig. 9. Comparisons of lengths of (A) second spine of first dorsal fin, (B) first spine of second dorsal fin, (C) second spine of second dorsal fin, (D) third spine of second dorsal fin, (E) second dorsal-fin base, and (F) anal-fin base (% standard length) with standard length in Enneapterygius phoenicosoma n. sp. (stars) and E. atriceps (circles)." figureDoi="http://doi.org/10.5281/zenodo.4583170" httpUri="https://zenodo.org/record/4583170/files/figure.png" pageId="10" pageNumber="11">Fig. 9C</figureCitation>
), and third spine (15.8%
<emphasis id="258BBE4F4950FFA6FAB73BCFB76FF138" box="[1335,1355,1012,1035]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
14.8%;
<tableCitation id="5A7D57E64950FFA6FCB43C2FB1A4F11F" box="[820,896,1044,1068]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
,
<figureCitation id="8FC47ED84950FFA6FC0F3C2FB1FAF11F" box="[911,990,1044,1068]" captionStart="Fig" captionStartId="9.[157,188,1228,1250]" captionTargetBox="[188,1397,207,1187]" captionTargetId="figure-380@9.[187,1399,205,1189]" captionTargetPageId="9" captionText="Fig. 9. Comparisons of lengths of (A) second spine of first dorsal fin, (B) first spine of second dorsal fin, (C) second spine of second dorsal fin, (D) third spine of second dorsal fin, (E) second dorsal-fin base, and (F) anal-fin base (% standard length) with standard length in Enneapterygius phoenicosoma n. sp. (stars) and E. atriceps (circles)." figureDoi="http://doi.org/10.5281/zenodo.4583170" httpUri="https://zenodo.org/record/4583170/files/figure.png" pageId="10" pageNumber="11">Fig. 9D</figureCitation>
) in the second dorsal fin; and the lengths of the first soft ray (17.3%
<emphasis id="258BBE4F4950FFA6FBD73C0EB64FF17F" box="[1111,1131,1077,1100]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
15.9%;
<tableCitation id="5A7D57E64950FFA6FB423C0FB729F17F" box="[1218,1293,1076,1100]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
), second ray (16.9%
<emphasis id="258BBE4F4950FFA6FC043C6DB1BCF15E" box="[900,920,1110,1133]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
15.6%;
<tableCitation id="5A7D57E64950FFA6FC6E3C6EB61DF15E" box="[1006,1081,1109,1133]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
), and third ray (15.9%
<emphasis id="258BBE4F4950FFA6FAB73C6DB76FF15E" box="[1335,1355,1110,1133]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
14.8%;
<tableCitation id="5A7D57E64950FFA6FCB43C4EB15FF1BE" box="[820,891,1141,1165]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
) in the third dorsal fin. In contrast, the lengths of the second dorsal-fin base and anal-fin base in
<taxonomicName id="D0FF19DE4950FFA6FB703CACB7B3F19D" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[1264,1431,1174,1198]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FB703CACB7B3F19D" box="[1264,1431,1174,1198]" italics="true" pageId="10" pageNumber="11">E. phoenicosoma</emphasis>
</taxonomicName>
tend to be slightly shorter than those of
<taxonomicName id="D0FF19DE4950FFA6FB633C8CB769F1FD" authorityName=": Fricke" authorityYear="1997" box="[1251,1357,1207,1230]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FB633C8CB769F1FD" box="[1251,1357,1207,1230]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
(mean 29.2% of SL and 40.5%, respectively,
<emphasis id="258BBE4F4950FFA6FB283CE3B698F1DC" box="[1192,1212,1240,1263]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
31.8% and 43.1%, respectively, in
<taxonomicName id="D0FF19DE4950FFA6FC3C3CC3B605F03C" authorityName=": Fricke" authorityYear="1997" box="[956,1057,1272,1295]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FC3C3CC3B605F03C" box="[956,1057,1272,1295]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
;
<tableCitation id="5A7D57E64950FFA6FBAD3CCCB651F023" box="[1069,1141,1271,1296]" captionStart="Table 1" captionStartId="5.[156,204,507,530]" captionTargetPageId="5" captionText="Table 1. Counts and measurements, expressed as percentages of standard length, of Enneapterygius phoenicosoma n. sp. and E.atriceps. Means in parentheses." httpUri="http://table.plazi.org/id/438032D5495FFFA9FF1C39C0B7F0F721" pageId="10" pageNumber="11" tableUuid="438032D5495FFFA9FF1C39C0B7F0F721">Table 1</tableCitation>
,
<figureCitation id="8FC47ED84950FFA6FB003CCCB6C7F023" box="[1152,1251,1271,1296]" captionStart="Fig" captionStartId="9.[157,188,1228,1250]" captionTargetBox="[188,1397,207,1187]" captionTargetId="figure-380@9.[187,1399,205,1189]" captionTargetPageId="9" captionText="Fig. 9. Comparisons of lengths of (A) second spine of first dorsal fin, (B) first spine of second dorsal fin, (C) second spine of second dorsal fin, (D) third spine of second dorsal fin, (E) second dorsal-fin base, and (F) anal-fin base (% standard length) with standard length in Enneapterygius phoenicosoma n. sp. (stars) and E. atriceps (circles)." figureDoi="http://doi.org/10.5281/zenodo.4583170" httpUri="https://zenodo.org/record/4583170/files/figure.png" pageId="10" pageNumber="11">Fig. 9E, F</figureCitation>
).
</paragraph>
<paragraph id="1740625D4950FFA6FCCE3D22B670F366" blockId="10.[820,1432,198,2076]" pageId="10" pageNumber="11">
<taxonomicName id="D0FF19DE4950FFA6FCCE3D22B659F003" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[846,1149,1304,1328]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FCCE3D22B659F003" box="[846,1149,1304,1328]" italics="true" pageId="10" pageNumber="11">Enneapterygius phoenicosoma</emphasis>
</taxonomicName>
grows larger than
<taxonomicName id="D0FF19DE4950FFA6FACC3D22B178F063" authorityName=": Fricke" authorityYear="1997" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FACC3D22B178F063" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
(largest recorded size,
<quantity id="D007CFB84950FFA6FBCE3D02B680F063" box="[1102,1188,1337,1361]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="10" pageNumber="11" unit="mm" value="37.2">37.2 mm</quantity>
SL
<emphasis id="258BBE4F4950FFA6FB4F3D02B6C7F063" box="[1231,1251,1337,1360]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
<quantity id="D007CFB84950FFA6FB6B3D03B766F063" box="[1259,1346,1336,1360]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="10" pageNumber="11" unit="mm" value="26.9">26.9 mm</quantity>
SL; this study).
<bibRefCitation id="736E1FAC4950FFA6FC023D62B628F042" author="Fricke, R." box="[898,1036,1369,1393]" pageId="10" pageNumber="11" refId="ref9457" refString="Fricke, R. 1997. Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein, ix + 607 pp." type="book" year="1997">Fricke (1997)</bibRefCitation>
examined
<specimenCount id="01F9A9D44950FFA6FB003D62B72EF042" box="[1152,1290,1369,1393]" pageId="10" pageNumber="11" type="generic">87 specimens</specimenCount>
of
<taxonomicName id="D0FF19DE4950FFA6FAAF3D61B7B3F042" authorityName=": Fricke" authorityYear="1997" box="[1327,1431,1370,1393]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FAAF3D61B7B3F042" box="[1327,1431,1370,1393]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
ranging from
<quantity id="D007CFB84950FFA6FC443D41B661F0A2" box="[964,1093,1402,1426]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.6" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="18.5" valueMax="26.0" valueMin="11.0">11 to 26 mm</quantity>
SL. On the basis of examination of otoliths of
<taxonomicName id="D0FF19DE4950FFA6FC413DA0B7B3F081" authority=", Longenecker and Langston (2005)" authorityName=", Longenecker and Langston" authorityYear="2005" box="[961,1431,1434,1459]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FC413DA0B60DF081" box="[961,1065,1435,1458]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
,
<bibRefCitation id="736E1FAC4950FFA6FBB73DA1B7B3F081" author="Longenecker, K. &amp; Langston, R." box="[1079,1431,1434,1459]" pageId="10" pageNumber="11" pagination="243 - 251" refId="ref9726" refString="Longenecker, K. and Langston, R. 2005. Life history of the Hawaiian blackhead triplefin, Enneapterygius atriceps (Blennioidei, Tripterygiidae). Environmental Biology of Fishes 73: 243 - 251." type="journal article" year="2005">Longenecker and Langston (2005)</bibRefCitation>
</taxonomicName>
determined the age of the oldest individual of
<taxonomicName id="D0FF19DE4950FFA6FA943D80B758F0E1" authorityName=", Longenecker and Langston" authorityYear="2005" box="[1300,1404,1467,1490]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="258BBE4F4950FFA6FA943D80B758F0E1" box="[1300,1404,1467,1490]" italics="true" pageId="10" pageNumber="11">E. atriceps</emphasis>
</taxonomicName>
as 117 days, with sexual maturity occurring at about 55 days and
<quantity id="D007CFB84950FFA6FCE23DC7B182F320" box="[866,934,1532,1556]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="mm" value="15.0">15 mm</quantity>
SL. No biological information on
<taxonomicName id="D0FF19DE4950FFA6FA873DC7B170F307" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FA873DC7B170F307" italics="true" pageId="10" pageNumber="11">E. phoenicosoma</emphasis>
</taxonomicName>
is known. These two species are allopatrically distributed in the Pacific Ocean (
<figureCitation id="8FC47ED84950FFA6FB8B3E06B663F366" box="[1035,1095,1597,1621]" captionStart="Fig" captionStartId="4.[821,852,646,668]" captionTargetBox="[823,1428,206,605]" captionTargetId="figure-452@4.[822,1430,205,607]" captionTargetPageId="4" captionText="Fig. 6. Distributional records of Enneapterygius phoenicosoma n. sp. and E. atriceps. Stars, localities of specimens of E. phoenicosoma; closed circles, localities of specimens of E. atriceps examined in this study; open circles, localities of specimens of E. atriceps examined by Fricke (1997)." figureDoi="http://doi.org/10.5281/zenodo.4583154" httpUri="https://zenodo.org/record/4583154/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="1740625D4950FFA6FCCE3E65B60AFD2F" blockId="10.[820,1432,198,2076]" pageId="10" pageNumber="11">
<taxonomicName id="D0FF19DE4950FFA6FCCE3E65B641F346" baseAuthorityName="Jordan &amp; Snyder" baseAuthorityYear="1902" box="[846,1125,1629,1653]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="etheostoma">
<emphasis id="258BBE4F4950FFA6FCCE3E65B641F346" box="[846,1125,1629,1653]" italics="true" pageId="10" pageNumber="11">Enneapterygius etheostoma</emphasis>
</taxonomicName>
(
<collectingCountry id="6FE822CD4950FFA6FBFB3E66B6E5F346" box="[1147,1217,1629,1653]" name="Jordan" pageId="10" pageNumber="11">Jordan</collectingCountry>
and Snyder, 1902) co-occurs with
<taxonomicName id="D0FF19DE4950FFA6FC583E44B6A6F3A5" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[984,1154,1662,1686]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FC583E44B6A6F3A5" box="[984,1154,1662,1686]" italics="true" pageId="10" pageNumber="11">E. phoenicosoma</emphasis>
</taxonomicName>
through at the entire distributional range of the latter in Japanese waters. Coloration of females and pale males in the two species is very similar to each other either in life or when fresh. Besides the coloration of nuptial males, however,
<taxonomicName id="D0FF19DE4950FFA6FB243F3AB76AF22B" authorityName="Motomura &amp; Ota &amp; Meguro &amp; Tashiro" authorityYear="2015" box="[1188,1358,1792,1816]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phoenicosoma">
<emphasis id="258BBE4F4950FFA6FB243F3AB76AF22B" box="[1188,1358,1792,1816]" italics="true" pageId="10" pageNumber="11">E. phoenicosoma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="D0FF19DE4950FFA6FCEF3F1AB1DFF20B" baseAuthorityName="Jordan &amp; Snyder" baseAuthorityYear="1902" box="[879,1019,1824,1848]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="etheostoma">
<emphasis id="258BBE4F4950FFA6FCEF3F1AB1DFF20B" box="[879,1019,1824,1848]" italics="true" pageId="10" pageNumber="11">E. etheostoma</emphasis>
</taxonomicName>
in the following features: nasal tentacle simple (
<emphasis id="258BBE4F4950FFA6FC0A3F79B1BAF26A" box="[906,926,1858,1881]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
bilobed in the latter), 11–13 (mode 12) second dorsal-fin spines [
<emphasis id="258BBE4F4950FFA6FC743F59B62CF24A" box="[1012,1032,1890,1913]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
12–16 (14)], 16–18 (17) anal-fin soft rays [
<emphasis id="258BBE4F4950FFA6FCED3FB8B1A5F2A9" box="[877,897,1923,1946]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
17–20 (19)], 2½–4 (3½) scale rows between base of first spine of second dorsal fin and pored lateral-line scale [
<emphasis id="258BBE4F4950FFA6FCBD3FFFB175F2E8" box="[829,849,1988,2011]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
4–7½ (6)], and 2–3 (2½) scale rows above last pored lateral-line scale [
<emphasis id="258BBE4F4950FFA6FC6C3FDFB624F2C8" box="[1004,1024,2020,2043]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
3½–5½ (4)] (
<bibRefCitation id="736E1FAC4950FFA6FB113FDFB72AF2CF" author="Fricke, R." box="[1169,1294,2020,2044]" pageId="10" pageNumber="11" refId="ref9457" refString="Fricke, R. 1997. Tripterygiid Fishes of the Western and Central Pacific (Teleostei). Koeltz Scientific Books, Koenigstein, ix + 607 pp." type="book" year="1997">Fricke 1997</bibRefCitation>
; Meguro and Motomura unpub. data).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue Copy permalink