treatments-xml/data/03/D2/47/03D247023910FFE4FF69F9D53240F85A.xml

<document id="E1CAE192784AB75A13B6962843E570CA" ID-CLB-Dataset="24604" ID-DOI="10.11646/zootaxa.4711.2.6" ID-GBIF-Dataset="57cfb289-89a3-48b0-901c-c76fb029d2c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3577059" ID-ZooBank="0A02D6A2-9DF4-415D-9A1C-51B313F59998" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576479411414" checkinUser="plazi" docAuthor="Li, Xu, Che, Xing-Jin &amp; Zhou, Wei" docDate="2019" docId="03D247023910FFE4FF69F9D53240F85A" docLanguage="en" docName="zootaxa.4711.2.6.pdf" docOrigin="Zootaxa 4711 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homatula nigra Li &amp; Che &amp; Zhou 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="342" masterDocId="FFEB3F7A391BFFE8FFFEFFA4311DFF85" masterDocTitle="Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species" masterLastPageNumber="348" masterPageNumber="330" pageNumber="341" updateTime="1698774435811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E07F8FBE6A02AE4DF1BB0255DAC1B24B">Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species</mods:title>
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<mods:namePart id="17E4D4286920E37D423BAE0898D33B84">Li, Xu</mods:namePart>
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<mods:namePart id="28BC9CF862CBD156FB0F6C97EC3ED746">Che, Xing-Jin</mods:namePart>
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<treatment id="03D247023910FFE4FF69F9D53240F85A" ID-DOI="http://doi.org/10.5281/zenodo.5587689" ID-GBIF-Taxon="160747846" ID-Zenodo-Dep="5587689" LSID="urn:lsid:plazi:treatment:03D247023910FFE4FF69F9D53240F85A" httpUri="http://treatment.plazi.org/id/03D247023910FFE4FF69F9D53240F85A" lastPageId="12" lastPageNumber="342" pageId="11" pageNumber="341">
<subSubSection id="C361A59F3910FFE3FF69F9D530A2F909" box="[151,447,1649,1676]" pageId="11" pageNumber="341" type="nomenclature">
<paragraph id="8BC4F6143910FFE3FF69F9D530A2F909" blockId="11.[151,447,1649,1712]" box="[151,447,1649,1676]" pageId="11" pageNumber="341">
<heading id="D08C41783910FFE3FF69F9D530A2F909" bold="true" box="[151,447,1649,1676]" fontSize="11" level="1" pageId="11" pageNumber="341" reason="1">
<emphasis id="B90F2A063910FFE3FF69F9D530A2F909" bold="true" box="[151,447,1649,1676]" pageId="11" pageNumber="341">
<taxonomicName id="4C7B8D973910FFE3FF69F9D53047F909" authority="Li &amp; Che &amp; Zhou, 2019" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[151,346,1649,1676]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="species" species="nigra" status="sp. nov.">
<emphasis id="B90F2A063910FFE3FF69F9D53047F909" bold="true" box="[151,346,1649,1676]" italics="true" pageId="11" pageNumber="341">Homatula nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C977D3910FFE3FE9FF9D630A2F909" box="[353,447,1650,1676]" pageId="11" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C361A59F3910FFE3FF69F9323041F935" box="[151,348,1686,1712]" pageId="11" pageNumber="341" type="description">
<paragraph id="8BC4F6143910FFE3FF69F9323041F935" blockId="11.[151,447,1649,1712]" box="[151,348,1686,1712]" pageId="11" pageNumber="341">
(
<figureCitation id="1340EA913910FFE3FF5EF93231EAF935" box="[160,247,1686,1712]" captionStart="FIGURE 2" captionStartId="5.[151,250,932,957]" captionTargetBox="[155,1432,320,847]" captionTargetId="figure@5.[151,1436,271,892]" captionTargetPageId="5" captionText="FIGURE 2. Relationship between anterior end of adipose crest and anal-fin base of Homatula. (A) H. anteridorsalis, SWFC 1610019, paratype, 108.1 mm SL; (B) H. cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; (C) H. nigra, SWFC 1901005, holotype, 99.8 mm SL; (D) H. pycnolepis, SWFC 0612085, 99.6 mm SL; (E) H. laxiclathra, SWFC 1810010, 97.0 mm SL); (F) H. nanpanjiangensis, SWFC 1903001, 97.1 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3577063" httpUri="https://zenodo.org/record/3577063/files/figure.png" pageId="11" pageNumber="341">Fig. 2C</figureCitation>
, 
<figureCitation id="1340EA913910FFE3FEFDF932304EF935" box="[259,339,1686,1712]" captionStart="FIGURE 10" captionStartId="13.[151,250,1151,1176]" captionTargetBox="[174,1413,174,1111]" captionTargetId="figure@13.[174,1413,174,1111]" captionTargetPageId="13" captionText="FIGURE 10. Homatula nigra, (A–B, D) SWFC 1901005, holotype, 99.8 mm SL; (C) SWFC 0411007, paratype, 87.4 mm SL, China: Yunnan: Baoshan City: Changning County: Gengga Town: Xiangshui Village." figureDoi="http://doi.org/10.5281/zenodo.3577081" httpUri="https://zenodo.org/record/3577081/files/figure.png" pageId="11" pageNumber="341">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3910FFE3FF69F97A301CF80D" pageId="11" pageNumber="341" type="materials_examined">
<paragraph id="8BC4F6143910FFE3FF69F97A354AF899" blockId="11.[151,1437,1757,2036]" pageId="11" pageNumber="341">
<materialsCitation id="3B13FC493910FFE3FF69F97A354AF899" ID-GBIF-Occurrence="2512877918" collectingDate="2019-01-08" country="China" county="Changning County" latitude="24.48535" location="Gengga Town" longLatPrecision="1" longitude="99.49442" municipality="Baoshan City" pageId="11" pageNumber="341" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90F2A063910FFE3FF69F97A3016F97D" bold="true" box="[151,267,1758,1784]" pageId="11" pageNumber="341">
<typeStatus id="54C048B63910FFE3FF69F97A301BF97D" box="[151,262,1758,1784]" pageId="11" pageNumber="341" type="holotype">Holotype</typeStatus>
.
</emphasis>
SWFC 1901005, 
<quantity id="4C835BF13910FFE3FE18F97A334DF97D" box="[486,592,1758,1784]" metricMagnitude="-2" metricUnit="m" metricValue="9.98" pageId="11" pageNumber="341" unit="mm" value="99.8">99.8 mm</quantity>
SL; 
<collectingCountry id="F36CB6843910FFE3FD6FF97A33C5F97D" box="[657,728,1758,1784]" name="China" pageId="11" pageNumber="341">China</collectingCountry>
: 
<collectingRegion id="49BF38F63910FFE3FD16F97A325FF97D" box="[744,834,1758,1784]" country="China" name="Yunnan" pageId="11" pageNumber="341">Yunnan</collectingRegion>
: 
<collectingMunicipality id="6BA06C6E3910FFE3FCACF97A32E9F97D" box="[850,1012,1758,1784]" pageId="11" pageNumber="341">Baoshan City</collectingMunicipality>
: 
<collectingCounty id="62A58E983910FFE3FBFDF97A35FFF97D" box="[1027,1250,1758,1784]" pageId="11" pageNumber="341">Changning County</collectingCounty>
: 
<location id="8EA4A0CF3910FFE3FB0CF97A3484F97D" LSID="urn:lsid:plazi:treatment:03D247023910FFE4FF69F9D53240F85A:8EA4A0CF3910FFE3FB0CF97A3484F97D" box="[1266,1433,1758,1784]" country="China" county="Changning County" latitude="24.48535" longLatPrecision="1" longitude="99.49442" municipality="Baoshan City" name="Gengga Town" pageId="11" pageNumber="341" stateProvince="Yunnan">Gengga Town</location>
: 
<location id="8EA4A0CF3910FFE3FF69F8A6307FF899" LSID="urn:lsid:plazi:treatment:03D247023910FFE4FF69F9D53240F85A:8EA4A0CF3910FFE3FF69F8A6307FF899" box="[151,354,1794,1820]" country="China" county="Changning County" latitude="24.48535" longLatPrecision="1" longitude="99.49442" municipality="Baoshan City" name="Xiangshui Village" pageId="11" pageNumber="341" stateProvince="Yunnan">Xiangshui Village</location>
(
<geoCoordinate id="EE4F90D33910FFE3FE8FF8A6330DF899" box="[369,528,1794,1820]" degrees="24" direction="north" minutes="29" orientation="latitude" pageId="11" pageNumber="341" precision="1" seconds="7.26" value="24.48535">24°29’7.26”N</geoCoordinate>
<geoCoordinate id="EE4F90D33910FFE3FDE9F8A633D8F899" box="[535,709,1794,1820]" degrees="99" direction="east" minutes="29" orientation="longitude" pageId="11" pageNumber="341" precision="1" seconds="39.94" value="99.49442">99°29’39.94”E</geoCoordinate>
); collected by XJ. 
<location id="8EA4A0CF3910FFE3FC6AF8A632D9F899" LSID="urn:lsid:plazi:treatment:03D247023910FFE4FF69F9D53240F85A:8EA4A0CF3910FFE3FC6AF8A632D9F899" box="[916,964,1794,1820]" country="China" county="Changning County" latitude="24.48535" longLatPrecision="1" longitude="99.49442" municipality="Baoshan City" name="Che" pageId="11" pageNumber="341" stateProvince="Yunnan">Che</location>
, 
<date id="FFC5D0D43910FFE3FC2EF8A63549F899" box="[976,1108,1794,1820]" pageId="11" pageNumber="341" value="2019-01-08">
<collectingDate id="EF81293C3910FFE3FC2EF8A63549F899" box="[976,1108,1794,1820]" pageId="11" pageNumber="341" value="2019-01-08">8 Jan. 2019</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC4F6143910FFE3FF39F882301CF80D" blockId="11.[151,1437,1757,2036]" pageId="11" pageNumber="341">
<materialsCitation id="3B13FC493910FFE3FF39F882345FF8C5" ID-GBIF-Occurrence="2512877913" box="[199,1346,1830,1856]" collectingDate="2019-01-08" country="China" county="Changning County" latitude="24.48535" location="Gengga Town" longLatPrecision="1" longitude="99.49442" municipality="Baoshan City" pageId="11" pageNumber="341" specimenCount="4" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B90F2A063910FFE3FF39F882305BF8C5" bold="true" box="[199,326,1830,1856]" pageId="11" pageNumber="341">
<typeStatus id="54C048B63910FFE3FF39F882305CF8C5" box="[199,321,1830,1856]" pageId="11" pageNumber="341" type="paratype">Paratypes</typeStatus>
.
</emphasis>
SWFC 1901001-1901004, 
<specimenCount id="9D7D3D9D3910FFE3FD76F88233D8F8C5" box="[648,709,1830,1856]" pageId="11" pageNumber="341" type="generic">4 ex.</specimenCount>
, 92.0– 
<quantity id="4C835BF13910FFE3FCE8F8823297F8C5" box="[790,906,1830,1856]" metricMagnitude="-1" metricUnit="m" metricValue="1.091" pageId="11" pageNumber="341" unit="mm" value="109.1">109.1 mm</quantity>
SL; other data same as the holotype
</materialsCitation>
. 
<materialsCitation id="3B13FC493910FFE3FAAEF88231E0F80D" ID-GBIF-Occurrence="2512877912" collectingDate="2004-11-23" collectorName="X. Li &amp; W. Zhou" country="China" county="Changning County" latitude="24.48535" location="Gengga Town" longLatPrecision="1" longitude="99.49442" municipality="Baoshan City" pageId="11" pageNumber="341" specimenCount="5" stateProvince="Yunnan" typeStatus="paratype">
SWFC 0411005-0411009, 
<specimenCount id="9D7D3D9D3910FFE3FE8FF8ED30B6F8E1" box="[369,427,1865,1892]" pageId="11" pageNumber="341" type="generic">5 ex.</specimenCount>
, 
<quantity id="4C835BF13910FFE3FE49F8ED3346F8E1" box="[439,603,1865,1892]" metricMagnitude="-2" metricUnit="m" metricValue="7.13" metricValueMax="8.74" metricValueMin="5.52" pageId="11" pageNumber="341" unit="mm" value="71.3" valueMax="87.4" valueMin="55.2">55.2–87.4 mm</quantity>
SL; collected by 
<collectorName id="268E93C23910FFE3FCDDF8EE3242F8E1" box="[803,863,1866,1892]" pageId="11" pageNumber="341">X. Li</collectorName>
&amp; 
<collectorName id="268E93C23910FFE3FC7AF8EE32F1F8E1" box="[900,1004,1866,1892]" pageId="11" pageNumber="341">W. Zhou</collectorName>
, 
<date id="FFC5D0D43910FFE3FC06F8EE358BF8E1" box="[1016,1174,1866,1892]" pageId="11" pageNumber="341" value="2004-11-23">
<collectingDate id="EF81293C3910FFE3FC06F8EE358BF8E1" box="[1016,1174,1866,1892]" pageId="11" pageNumber="341" value="2004-11-23">23 Nov. 2004</collectingDate>
</date>
; other data same as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3910FFE4FF39F83530DFFE0E" lastPageId="12" lastPageNumber="342" pageId="11" pageNumber="341" type="diagnosis">
<paragraph id="8BC4F6143910FFE4FF39F83530DFFE0E" blockId="11.[151,1437,1757,2036]" lastBlockId="12.[151,1437,153,2015]" lastPageId="12" lastPageNumber="342" pageId="11" pageNumber="341">
<emphasis id="B90F2A063910FFE3FF39F835305FF829" bold="true" box="[199,322,1937,1964]" pageId="11" pageNumber="341">Diagnosis.</emphasis>
<taxonomicName id="4C7B8D973910FFE3FEB9F83630E7F829" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[327,506,1938,1964]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063910FFE3FEB9F83630E7F829" box="[327,506,1938,1964]" italics="true" pageId="11" pageNumber="341">Homatula nigra</emphasis>
</taxonomicName>
differs from other species of the densely-scaled group in 
<taxonomicName id="4C7B8D973910FFE3FB8EF83635FDF829" authorityName="Nichols" authorityYear="1925" box="[1136,1248,1938,1964]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063910FFE3FB8EF83635FDF829" box="[1136,1248,1938,1964]" italics="true" pageId="11" pageNumber="341">Homatula</emphasis>
</taxonomicName>
by the following combination of characters: no marks on flank in 
<taxonomicName id="4C7B8D973910FFE3FD53F8133216F855" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[685,779,1975,2000]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063910FFE3FD53F8133216F855" box="[685,779,1975,2000]" italics="true" pageId="11" pageNumber="341">H. nigra</emphasis>
</taxonomicName>
(vs. vertical bars on flank along lateral line in all congeners of densely-scaled group in 
<taxonomicName id="4C7B8D973910FFE3FE38F87E332BF871" authorityName="Nichols" authorityYear="1925" box="[454,566,2010,2036]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063910FFE3FE38F87E332BF871" box="[454,566,2010,2036]" italics="true" pageId="11" pageNumber="341">Homatula</emphasis>
</taxonomicName>
); no free protrusions between pelvic fins (vs. pair of free protrusions between pelvic fins in 
<taxonomicName id="4C7B8D973917FFE4FEC8FF3E30CFFF36" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[310,466,153,179]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063917FFE4FEC8FF3E30CFFF36" box="[310,466,153,179]" italics="true" pageId="12" pageNumber="342">H. pycnolepis</emphasis>
</taxonomicName>
); gill opening smaller, its upper angle level with lower edge of eye (vs. gill opening larger, its upper angle aligned with middle point of eye in 
<taxonomicName id="4C7B8D973917FFE4FCC7FF1A32F2FF52" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[825,1007,189,215]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063917FFE4FCC7FF1A32F2FF52" box="[825,1007,189,215]" italics="true" pageId="12" pageNumber="342">H. acuticephala</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D973917FFE4FC00FF1A35A8FF52" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[1022,1205,189,215]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A063917FFE4FC00FF1A35A8FF52" box="[1022,1205,189,215]" italics="true" pageId="12" pageNumber="342">H. anguillioides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D973917FFE4FB3BFF1A346CFF52" authorityName="Endruweit, Min &amp; Yang" authorityYear="2018" box="[1221,1393,189,215]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="coccinocola">
<emphasis id="B90F2A063917FFE4FB3BFF1A346CFF52" box="[1221,1393,189,215]" italics="true" pageId="12" pageNumber="342">H. coccinocola</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D973917FFE4FA7FFF1A3017FF7E" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A063917FFE4FA7FFF1A3017FF7E" italics="true" pageId="12" pageNumber="342">H. erhaiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7B8D973917FFE4FEB8FF4630C2FF7E" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[326,479,225,251]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063917FFE4FEB8FF4630C2FF7E" box="[326,479,225,251]" italics="true" pageId="12" pageNumber="342">H. pycnolepis</emphasis>
</taxonomicName>
) (
<figureCitation id="1340EA913917FFE4FE09FF453326FF7E" box="[503,571,224,251]" captionStart="FIGURE 5" captionStartId="7.[151,250,559,584]" captionTargetBox="[166,1421,174,540]" captionTargetId="figure@7.[166,1421,174,540]" captionTargetPageId="7" captionText="FIGURE 5. Gill openings of Homatula. (A) Larger gill opening; upper angle aligned with middle point of eye (H. pycnolepis, SWFC 0612085, 99.6 mm SL); (B) smaller gill opening; upper angle level with lower edge of eye (H. cryptoclathrata, SWFC 1610006, holotype, 121.6 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.3577071" httpUri="https://zenodo.org/record/3577071/files/figure.png" pageId="12" pageNumber="342">Fig. 5</figureCitation>
); with pelvic axillary lobe (vs. without pelvic axillary lobe in 
<taxonomicName id="4C7B8D973917FFE4FB14FF463480FF7E" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[1258,1437,225,251]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063917FFE4FB14FF463480FF7E" box="[1258,1437,225,251]" italics="true" pageId="12" pageNumber="342">H. acuticephala</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7B8D973917FFE4FF39FEA23061FE9A" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[199,380,261,287]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A063917FFE4FF39FEA23061FE9A" box="[199,380,261,287]" italics="true" pageId="12" pageNumber="342">H. anguillioides</emphasis>
</taxonomicName>
); marked notch on lower jaw (vs. lower jaw spoon-shaped and lacking notch in 
<taxonomicName id="4C7B8D973917FFE4FAF9FEA23480FE9A" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[1287,1437,261,287]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A063917FFE4FAF9FEA23480FE9A" box="[1287,1437,261,287]" italics="true" pageId="12" pageNumber="342">H. erhaiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7B8D973917FFE4FF36FE8E3060FEC6" authorityName="Min, Yang &amp; Chen" authorityYear="2012" box="[200,381,297,323]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="wuliangensis">
<emphasis id="B90F2A063917FFE4FF36FE8E3060FEC6" box="[200,381,297,323]" italics="true" pageId="12" pageNumber="342">H. wuliangensis</emphasis>
</taxonomicName>
); dorsal-fin base shorter than longest branched dorsal-fin ray (vs. dorsal-fin base longer than longest branched dorsal-fin ray in 
<taxonomicName id="4C7B8D973917FFE4FDE7FEEA33D0FEE2" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[537,717,333,359]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063917FFE4FDE7FEEA33D0FEE2" box="[537,717,333,359]" italics="true" pageId="12" pageNumber="342">H. acuticephala</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D973917FFE4FD25FEEA3292FEE2" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[731,911,333,359]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A063917FFE4FD25FEEA3292FEE2" box="[731,911,333,359]" italics="true" pageId="12" pageNumber="342">H. anguillioides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D973917FFE4FC60FEEA3573FEE2" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[926,1134,333,359]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="cryptoclathrata">
<emphasis id="B90F2A063917FFE4FC60FEEA3573FEE2" box="[926,1134,333,359]" italics="true" pageId="12" pageNumber="342">H. cryptoclathrata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D973917FFE4FB83FEEA340FFEE2" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[1149,1298,333,359]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A063917FFE4FB83FEEA340FFEE2" box="[1149,1298,333,359]" italics="true" pageId="12" pageNumber="342">H. erhaiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7B8D973917FFE4FAAEFEEA3014FE0E" authorityName="Min, Yang &amp; Chen" authorityYear="2012" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="wuliangensis">
<emphasis id="B90F2A063917FFE4FAAEFEEA3014FE0E" italics="true" pageId="12" pageNumber="342">H. wuliangensis</emphasis>
</taxonomicName>
) (
<tableCitation id="C6F9C3AF3917FFE4FEDFFED5309DFE0E" box="[289,384,369,395]" captionStart="TABLE 1" captionStartId="2.[151,239,579,605]" captionTargetBox="[165,2015,624,1571]" captionText="TABLE 1. Comparisons of characteristic among the species of the densely-scaled group of Homatula" httpUri="http://table.plazi.org/id/DF04A69C3919FFEAFF69FDE735C8FDD8" pageId="12" pageNumber="342" tableUuid="DF04A69C3919FFEAFF69FDE735C8FDD8">Tables 1</tableCitation>
&amp; 
<tableCitation id="C6F9C3AF3917FFE4FE5AFED530A8FE0E" box="[420,437,369,395]" captionStart="TABLE 2" captionStartId="14.[151,239,470,496]" captionTargetBox="[165,1975,507,1700]" captionText="TABLE 2. Morphometric data of three new Homatula species and H. pycnolepis" httpUri="http://table.plazi.org/id/DF04A69C3915FFE6FF69FE7232E8FE75" pageId="12" pageNumber="342" tableUuid="DF04A69C3915FFE6FF69FE7232E8FE75">2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3917FFE4FF39FE30344CFA1A" pageId="12" pageNumber="342" type="description">
<paragraph id="8BC4F6143917FFE4FF39FE303519FE7D" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
<emphasis id="B90F2A063917FFE4FF39FE303044FE2A" bold="true" box="[199,345,404,431]" pageId="12" pageNumber="342">Description.</emphasis>
Based on 
<typeStatus id="54C048B63917FFE4FE2EFE31332FFE2A" box="[464,562,405,431]" pageId="12" pageNumber="342" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D7D3D9D3917FFE4FD95FE3133F3FE2A" box="[619,750,405,431]" pageId="12" pageNumber="342" type="generic" typeStatus="paratypes">9 paratypes</specimenCount>
, maximum standard length is 
<quantity id="4C835BF13917FFE4FBBDFE3135A4FE2A" box="[1091,1209,405,431]" metricMagnitude="-1" metricUnit="m" metricValue="1.091" pageId="12" pageNumber="342" unit="mm" value="109.1">109.1 mm</quantity>
. Morphometric data are shown in 
<tableCitation id="C6F9C3AF3917FFE4FECDFE1D3090FE56" box="[307,397,441,467]" captionStart="TABLE 2" captionStartId="14.[151,239,470,496]" captionTargetBox="[165,1975,507,1700]" captionText="TABLE 2. Morphometric data of three new Homatula species and H. pycnolepis" httpUri="http://table.plazi.org/id/DF04A69C3915FFE6FF69FE7232E8FE75" pageId="12" pageNumber="342" tableUuid="DF04A69C3915FFE6FF69FE7232E8FE75">Table 2</tableCitation>
. Dorsal-fin rays iii, 8½; anal-fin rays iii, 5½; pectoral-fin rays i, 9–10; pelvic-fin rays i, 7; caudal-fin branched rays 9+8; vertebrae 4+41 (
<specimenCount id="9D7D3D9D3917FFE4FD58FE79322FFE7D" box="[678,818,477,504]" pageId="12" pageNumber="342" type="generic">2 specimens</specimenCount>
)–42 (
<specimenCount id="9D7D3D9D3917FFE4FC88FE7932E5FE7D" box="[886,1016,477,504]" pageId="12" pageNumber="342" type="generic">1 specimen</specimenCount>
).
</paragraph>
<paragraph id="8BC4F6143917FFE4FF39FDA532D6FD02" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">Body elongate and cylindrical, anteriorly slightly depressed, posteriorly compressed laterally. Maximum body depth immediately in front of dorsal fin. Dorsal and ventral profiles almost straight. Except for scaleless head and thorax, body is entirely covered by small scales, with scattered scales on abdomen. Caudal peduncle compressed laterally. Lateral line complete, extending directly along midline of body.</paragraph>
<paragraph id="8BC4F6143917FFE4FF39FD353223FC92" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">Head short and depressed in frontal view, wider than high, roughly triangular in dorsal view. Snout blunt, slightly shorter than postorbital length of head. Eye small, close to dorsal profile of head, invisible from ventral view. Interorbital space wide and flat. Nostrils closely set, nearer to anterior margin of eye than to snout tip; anterior nare pierced in front side of a flap-like tube with pointed tip.</paragraph>
<paragraph id="8BC4F6143917FFE4FF39FC853557FC4E" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
Mouth inferior and arched. Lips thick, slightly furrowed, but not papillated; upper lip with small median incision, lower lip with marked median incision. Jaws covered by lips; upper jaw with well developed processus dentiformis corresponding with marked notch on lower jaw. Three pairs of barbels; two rostral pairs, inner barbel not reaching corner of mouth and outer barbel pair reaching vertical line of anterior nostril; one maxillary barbel pair long, reaching to vertical line between middle and posterior margin of eye (
<figureCitation id="1340EA913917FFE4FC1BFC153520FC4E" box="[997,1085,945,971]" captionStart="FIGURE 7" captionStartId="8.[151,250,1933,1958]" captionTargetBox="[217,1391,853,1887]" captionTargetId="figure@8.[151,1436,838,1899]" captionTargetPageId="8" captionText="FIGURE 7. Mouths of Homatula. (A) H. anteridorsalis, SWFC 1610006, holotype, 121.6 mm SL; (B) H. cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; (C) H. nigra, SWFC 1901005, holotype, 99.8 mm SL; (D) H. pycnolepis, SWFC 0612085, 99.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3577075" httpUri="https://zenodo.org/record/3577075/files/figure.png" pageId="12" pageNumber="342">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BC4F6143917FFE4FF39FC713431FAB6" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">Dorsal fin with 3 simple and 8–9 branched rays; distal margin convex; origin slightly forward of midway between snout tip and caudal-fin base. Pectoral fin with 1 simple and 9–10 branched rays, inserted slightly posterior to vertical through posteriormost point of operculum, tip of longest fin ray not extending beyond halfway to insertion of pelvic fin. Pelvic fin with 1 simple and 6–7 branched rays, inserted below first or second branched rays of dorsal fin, tip of longest fin ray not extending beyond half distance between pelvic-fin insertion and anal-fin origin. Pelvic axillary lobe with pointed tip. Anal fin with 3 simple and 5 branched rays, with convex distal edge; origin closer to pelvic-fin insertion than to caudal-fin base. Posterior margin of caudal fin oblique or micro-concave. Caudal peduncle uniformly deep; with adipose crests along dorsal and ventral midlines. Adipose crest along dorsal midline of caudal peduncle not extending through posterior end of anal-fin base. Anus located posteriorly, approximately 1/3 the distance between the pelvic-fin insertion and anal-fin origin, closer to the caudal-fin base.</paragraph>
<paragraph id="8BC4F6143917FFE4FF39FA98344CFA1A" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
<emphasis id="B90F2A063917FFE4FF39FA98304FFAD2" bold="true" box="[199,338,1340,1367]" pageId="12" pageNumber="342">Coloration:</emphasis>
Flank of fresh specimen black with no bars, dorsum of body dark brown, abdomen pale yellow. After fixation in 95% alcohol, flank is brown, abdomen grayish yellow. Dark brown vertical bar on caudal-fin base. Dorsal surface of pectoral, pelvic, and anal fins reddish orange; dorsal and caudal fins yellowish (
<figureCitation id="1340EA913917FFE4FB22FA213458FA1A" box="[1244,1349,1413,1439]" captionStart="FIGURE 10" captionStartId="13.[151,250,1151,1176]" captionTargetBox="[174,1413,174,1111]" captionTargetId="figure@13.[174,1413,174,1111]" captionTargetPageId="13" captionText="FIGURE 10. Homatula nigra, (A–B, D) SWFC 1901005, holotype, 99.8 mm SL; (C) SWFC 0411007, paratype, 87.4 mm SL, China: Yunnan: Baoshan City: Changning County: Gengga Town: Xiangshui Village." figureDoi="http://doi.org/10.5281/zenodo.3577081" httpUri="https://zenodo.org/record/3577081/files/figure.png" pageId="12" pageNumber="342">Fig. 10D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3917FFE4FF39FA0C33A0F9D6" pageId="12" pageNumber="342" type="distribution">
<paragraph id="8BC4F6143917FFE4FF39FA0C33DBFA62" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
<emphasis id="B90F2A063917FFE4FF39FA0C307EFA46" bold="true" box="[199,355,1448,1475]" pageId="12" pageNumber="342">Distribution:</emphasis>
Known only from Damengtong-he, a branch of the Mengboluo-he, the first tributary of the Nujiang drainage (the upper Salween River) (
<figureCitation id="1340EA913917FFE4FD8AFA6933A5FA62" box="[628,696,1485,1511]" captionStart="FIGURE 8" captionStartId="9.[151,250,1995,2020]" captionTargetBox="[159,1425,702,1967]" captionTargetId="figure@9.[151,1436,695,1973]" captionTargetPageId="9" captionText="FIGURE 8. Distribution map of the densely-scaled group of Homatula. A symbol may represent several adjacent localities." figureDoi="http://doi.org/10.5281/zenodo.3577077" httpUri="https://zenodo.org/record/3577077/files/figure.png" pageId="12" pageNumber="342">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BC4F6143917FFE4FF39FA5433A0F9D6" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
<emphasis id="B90F2A063917FFE4FF39FA5430D0F98E" bold="true" box="[199,461,1520,1547]" pageId="12" pageNumber="342">Habitat and Ecology:</emphasis>
<taxonomicName id="4C7B8D973917FFE4FE26FA55338DF98E" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[472,656,1521,1547]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063917FFE4FE26FA55338DF98E" box="[472,656,1521,1547]" italics="true" pageId="12" pageNumber="342">Homatula nigra</emphasis>
</taxonomicName>
is omnivorous, mainly feeding on algae attached to rocks, organic residues, and small aquatic insects. It inhabits flowing and clear stream environments, dwelling at the bottom and shuttling back and forth among rocks and gravel.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3917FFE4FF39F9F93216F91E" pageId="12" pageNumber="342" type="etymology">
<paragraph id="8BC4F6143917FFE4FF39F9F93216F91E" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
<emphasis id="B90F2A063917FFE4FF39F9F9304FF9F2" bold="true" box="[199,338,1629,1655]" pageId="12" pageNumber="342">Etymology:</emphasis>
The specific epithet, 
<taxonomicName id="4C7B8D973917FFE4FDBAF9FA339CF9F2" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[580,641,1630,1655]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063917FFE4FDBAF9FA339CF9F2" box="[580,641,1630,1655]" italics="true" pageId="12" pageNumber="342">nigra</emphasis>
</taxonomicName>
, an adjective, is from the Latin word 
<emphasis id="B90F2A063917FFE4FBD6F9FA357EF9F2" box="[1064,1123,1630,1655]" italics="true" pageId="12" pageNumber="342">niger</emphasis>
(black), alluding to the pattern of bars on the flank in front of the dorsal-fin origin.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3917FFE4FF39F9013240F85A" pageId="12" pageNumber="342" type="discussion">
<paragraph id="8BC4F6143917FFE4FF39F9013240F85A" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="342">
<emphasis id="B90F2A063917FFE4FF39F901302FF93A" bold="true" box="[199,306,1701,1727]" pageId="12" pageNumber="342">Remark:</emphasis>
The body color of 
<taxonomicName id="4C7B8D973917FFE4FDECF902336EF93A" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[530,627,1702,1727]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063917FFE4FDECF902336EF93A" box="[530,627,1702,1727]" italics="true" pageId="12" pageNumber="342">H. nigra</emphasis>
</taxonomicName>
is unique compared with the body color of other species of 
<taxonomicName id="4C7B8D973917FFE4FADBF9013488F93A" authorityName="Nichols" authorityYear="1925" box="[1317,1429,1701,1727]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063917FFE4FADBF9013488F93A" box="[1317,1429,1701,1727]" italics="true" pageId="12" pageNumber="342">Homatula</emphasis>
</taxonomicName>
. There are no bars on its flank and the whole body is black. However, when the specimens are fixed and preserved in alcohol, the discoloration of the specimen is significant and &quot;irregular cloud marks&quot; will appear on the flank. The flank of the fresh specimen collected in 
<date id="FFC5D0D43917FFE4FDA9F8B533DEF8AE" box="[599,707,1809,1835]" pageId="12" pageNumber="342" value="2019-01">Jan. 2019</date>
was black with no bars, the dorsum of the body was dark brown, and the abdomen was pale yellow (
<figureCitation id="1340EA913917FFE4FDDBF8913392F8CA" box="[549,655,1845,1871]" captionStart="FIGURE 10" captionStartId="13.[151,250,1151,1176]" captionTargetBox="[174,1413,174,1111]" captionTargetId="figure@13.[174,1413,174,1111]" captionTargetPageId="13" captionText="FIGURE 10. Homatula nigra, (A–B, D) SWFC 1901005, holotype, 99.8 mm SL; (C) SWFC 0411007, paratype, 87.4 mm SL, China: Yunnan: Baoshan City: Changning County: Gengga Town: Xiangshui Village." figureDoi="http://doi.org/10.5281/zenodo.3577081" httpUri="https://zenodo.org/record/3577081/files/figure.png" pageId="12" pageNumber="342">Fig. 10D</figureCitation>
). In the field, we noticed that when individuals of fresh specimens of 
<taxonomicName id="4C7B8D973917FFE4FF69F8FD301AF8F6" authorityName="Nichols" authorityYear="1925" box="[151,263,1881,1907]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063917FFE4FF69F8FD301AF8F6" box="[151,263,1881,1907]" italics="true" pageId="12" pageNumber="342">Homatula</emphasis>
</taxonomicName>
are pushed and squeezed in a narrow vessel, the color on the body will be lost soon after touching, even before fixing in alcohol; furthermore, the discoloration is uneven when the specimen is fixed with alcohol causing the fixed samples to show a false appearance of the &quot;irregular cloud marks.&quot; This phenomenon is also common in fresh specimens of the families 
<taxonomicName id="4C7B8D973917FFE4FE05F861336AF85A" authorityName="Swainson" authorityYear="1839" box="[507,631,1989,2015]" class="Actinopterygii" family="Balitoridae" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="family">Balitoridae</taxonomicName>
and 
<taxonomicName id="4C7B8D973917FFE4FD53F8613244F85A" box="[685,857,1989,2015]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="342" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>